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165-520: The Oligocene is often considered an important time of transition, a link between the archaic world of the tropical Eocene and the more modern ecosystems of the Miocene. Major changes during the Oligocene included a global expansion of grasslands , and a regression of tropical broad leaf forests to the equatorial belt . The start of the Oligocene is marked by a notable extinction event called

330-599: A Greek periplus written by an unknown author around the 1st century, contains a detailed description of the Red Sea's ports and sea routes. The Periplus also describes how Hippalus first discovered the direct route from the Red Sea to India . The Red Sea was favored for Roman trade with India starting with the reign of Augustus , when the Roman Empire gained control over the Mediterranean, Egypt , and

495-409: A cell wall . Newly dead animals may be covered by an exoskeleton . Fragmentation processes, which break through these protective layers, accelerate the rate of microbial decomposition. Animals fragment detritus as they hunt for food, as does passage through the gut. Freeze-thaw cycles and cycles of wetting and drying also fragment dead material. The chemical alteration of the dead organic matter

660-495: A food chain . Real systems are much more complex than this—organisms will generally feed on more than one form of food, and may feed at more than one trophic level. Carnivores may capture some prey that is part of a plant-based trophic system and others that are part of a detritus-based trophic system (a bird that feeds both on herbivorous grasshoppers and earthworms, which consume detritus). Real systems, with all these complexities, form food webs rather than food chains which present

825-619: A habitat . Ecosystem ecology is the "study of the interactions between organisms and their environment as an integrated system". The size of ecosystems can range up to ten orders of magnitude , from the surface layers of rocks to the surface of the planet. The Hubbard Brook Ecosystem Study started in 1963 to study the White Mountains in New Hampshire . It was the first successful attempt to study an entire watershed as an ecosystem. The study used stream chemistry as

990-665: A Phanerozoic low in diversity by the late Oligocene, and the temperate forests and jungles of the Eocene were replaced by forest and scrubland. The closing of the Tethys Seaway destroyed its tropical biota. The Oi1 event of the Eocene-Oligocene transition covered the continent of Antarctica with ice sheets, leaving Nothofagus and mosses and ferns clinging to life around the periphery of Antarctica in tundra conditions. Angiosperms continued their expansion throughout

1155-609: A central role over a wide range, for example, in the slow development of soil from bare rock and the faster recovery of a community from disturbance . Disturbance also plays an important role in ecological processes. F. Stuart Chapin and coauthors define disturbance as "a relatively discrete event in time that removes plant biomass". This can range from herbivore outbreaks, treefalls, fires, hurricanes, floods, glacial advances , to volcanic eruptions . Such disturbances can cause large changes in plant, animal and microbe populations, as well as soil organic matter content. Disturbance

1320-507: A critical role in global nutrient cycling and ecosystem function. Phosphorus enters ecosystems through weathering . As ecosystems age this supply diminishes, making phosphorus-limitation more common in older landscapes (especially in the tropics). Calcium and sulfur are also produced by weathering, but acid deposition is an important source of sulfur in many ecosystems. Although magnesium and manganese are produced by weathering, exchanges between soil organic matter and living cells account for

1485-491: A critical threshold value. Brachiopod oxygen isotope ratios from New Zealand suggest that a proto-Subtropical Convergence developed during the Early Oligocene, with northern New Zealand being subtropical and southern and eastern New Zealand being cooled by cold, subantarctic water. Oligocene climate following the Eocene-Oligocene event is poorly known. There were several pulses of glaciation in middle Oligocene, about

1650-517: A faster recovery. More severe and more frequent disturbance result in longer recovery times. From one year to another, ecosystems experience variation in their biotic and abiotic environments. A drought , a colder than usual winter, and a pest outbreak all are short-term variability in environmental conditions. Animal populations vary from year to year, building up during resource-rich periods and crashing as they overshoot their food supply. Longer-term changes also shape ecosystem processes. For example,

1815-427: A function-based typology has been proposed to leverage the strengths of these different approaches into a unified system. Human activities are important in almost all ecosystems. Although humans exist and operate within ecosystems, their cumulative effects are large enough to influence external factors like climate. Ecosystems provide a variety of goods and services upon which people depend. Ecosystem goods include

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1980-698: A general level, for example, tropical forests , temperate grasslands , and arctic tundra . There can be any degree of subcategories among ecosystem types that comprise a biome, e.g., needle-leafed boreal forests or wet tropical forests. Although ecosystems are most commonly categorized by their structure and geography, there are also other ways to categorize and classify ecosystems such as by their level of human impact (see anthropogenic biome ), or by their integration with social processes or technological processes or their novelty (e.g. novel ecosystem ). Each of these taxonomies of ecosystems tends to emphasize different structural or functional properties. None of these

2145-577: A major mountain chain in the Oligocene, as subduction became more direct into the coastline. Climate during the Oligocene reflected a general cooling trend following the Early Eocene Climatic Optimum . This transformed the Earth's climate from a greenhouse to an icehouse climate. The Eocene-Oligocene transition was a major cooling event and reorganization of the biosphere, being part of a broader trend of global cooling lasting from

2310-672: A means of monitoring ecosystem properties, and developed a detailed biogeochemical model of the ecosystem. Long-term research at the site led to the discovery of acid rain in North America in 1972. Researchers documented the depletion of soil cations (especially calcium) over the next several decades. Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Studies can be carried out at

2475-528: A more important role in moving nutrients around. This can be especially important as the soil thaws in the spring, creating a pulse of nutrients that become available. Decomposition rates are low under very wet or very dry conditions. Decomposition rates are highest in wet, moist conditions with adequate levels of oxygen. Wet soils tend to become deficient in oxygen (this is especially true in wetlands ), which slows microbial growth. In dry soils, decomposition slows as well, but bacteria continue to grow (albeit at

2640-406: A number of common, non random properties in the topology of their network. The carbon and nutrients in dead organic matter are broken down by a group of processes known as decomposition. This releases nutrients that can then be re-used for plant and microbial production and returns carbon dioxide to the atmosphere (or water) where it can be used for photosynthesis. In the absence of decomposition,

2805-682: A popular destination for diving after the expeditions of Hans Hass in the 1950s, and later by Jacques-Yves Cousteau . Popular tourist resorts include El Gouna , Hurghada , Safaga , Marsa Alam , on the west shore of the Red Sea, and Sharm-el-Sheikh , Dahab , and Taba on the Egyptian side of Sinaï , as well as Aqaba in Jordan and Eilat in Israel in an area known as the Red Sea Riviera . The popular tourist beach of Sharm el-Sheikh

2970-421: A process known as denitrification . Mycorrhizal fungi which are symbiotic with plant roots, use carbohydrates supplied by the plants and in return transfer phosphorus and nitrogen compounds back to the plant roots. This is an important pathway of organic nitrogen transfer from dead organic matter to plants. This mechanism may contribute to more than 70 Tg of annually assimilated plant nitrogen, thereby playing

3135-458: A semiarid prairie home and included entelodonts like Archaeotherium , camelids (such as Poebrotherium ), running rhinoceratoids , three-toed equids (such as Mesohippus ), nimravids , protoceratids , and early canids like Hesperocyon . Merycoidodonts, an endemic American group, were very diverse during this time. Australia and South America became geographically isolated and developed their own distinctive endemic fauna. These included

3300-696: A significant portion of ecosystem fluxes. Potassium is primarily cycled between living cells and soil organic matter. Biodiversity plays an important role in ecosystem functioning. Ecosystem processes are driven by the species in an ecosystem, the nature of the individual species, and the relative abundance of organisms among these species. Ecosystem processes are the net effect of the actions of individual organisms as they interact with their environment. Ecological theory suggests that in order to coexist, species must have some level of limiting similarity —they must be different from one another in some fundamental way, otherwise, one species would competitively exclude

3465-408: A slower rate) even after soils become too dry to support plant growth. Ecosystems are dynamic entities. They are subject to periodic disturbances and are always in the process of recovering from past disturbances. When a perturbation occurs, an ecosystem responds by moving away from its initial state. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance,

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3630-464: A small effect on ecosystem function. Ecologically distinct species, on the other hand, have a much larger effect. Similarly, dominant species have a large effect on ecosystem function, while rare species tend to have a small effect. Keystone species tend to have an effect on ecosystem function that is disproportionate to their abundance in an ecosystem. An ecosystem engineer is any organism that creates, significantly modifies, maintains or destroys

3795-468: A variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Biomes are general classes or categories of ecosystems. However, there is no clear distinction between biomes and ecosystems. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of

3960-443: A variety of goods and services upon which people depend, and may be part of. Ecosystem goods include the "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like the maintenance of hydrological cycles , cleaning air and water,

4125-522: A variety of scales, ranging from whole-ecosystem studies to studying microcosms or mesocosms (simplified representations of ecosystems). American ecologist Stephen R. Carpenter has argued that microcosm experiments can be "irrelevant and diversionary" if they are not carried out in conjunction with field studies done at the ecosystem scale. In such cases, microcosm experiments may fail to accurately predict ecosystem-level dynamics. Biomes are general classes or categories of ecosystems. However, there

4290-849: Is a direct translation of the Ancient Greek Erythra Thalassa ( Ἐρυθρὰ Θάλασσα ). The sea itself was once referred to as the Erythraean Sea by Europeans. As well as Mare Rubrum in Latin (alternatively Sinus Arabicus , literally "Arabian Gulf"), the Romans called it Pontus Herculis (Sea of Hercules). Other designations include the Arabic : البحر الأحمر , romanized :  Al-Baḥr Al-Aḥmar (alternatively بحر القلزم Baḥr Al-Qulzum , literally "the Sea of Clysma "),

4455-877: Is a sea inlet of the Indian Ocean , lying between Africa and Asia. Its connection to the ocean is in the south, through the Bab-el-Mandeb strait and the Gulf of Aden . To its north lie the Sinai Peninsula , the Gulf of Aqaba , and the Gulf of Suez —leading to the Suez Canal . It is underlain by the Red Sea Rift , which is part of the Great Rift Valley . The Red Sea has a surface area of roughly 438,000 km (169,000 sq mi),

4620-431: Is a system that environments and their organisms form through their interaction. The biotic and abiotic components are linked together through nutrient cycles and energy flows. Ecosystems are controlled by external and internal factors . External factors such as climate , parent material which forms the soil and topography , control the overall structure of an ecosystem but are not themselves influenced by

4785-617: Is about 2,250 km (1,400 mi) long, and 355 km (221 mi) wide at its widest point. It has an average depth of 490 m (1,610 ft), and in the central Suakin Trough it reaches its maximum depth of 3,040 m (9,970 ft). Approximately 40% of the Red Sea is quite shallow at less than 100 m (330 ft) deep, and about 25% is less than 50 m (160 ft) deep. The extensive shallow shelves are noted for their marine life and corals . More than 1,000 invertebrate species and 200 types of soft and hard coral live in

4950-406: Is an international synthesis by over 1000 of the world's leading biological scientists that analyzes the state of the Earth's ecosystems and provides summaries and guidelines for decision-makers. The report identified four major categories of ecosystem services: provisioning, regulating, cultural and supporting services. It concludes that human activity is having a significant and escalating impact on

5115-480: Is considered that it will become an ocean in time (as proposed in the model of John Tuzo Wilson ). In 1949, a deep water survey reported anomalously hot brines in the central portion of the Red Sea. Later work in the 1960s confirmed the presence of hot, 60 °C (140 °F), saline brines and associated metalliferous muds. The hot solutions were emanating from an active subseafloor rift . Lake Asal in Djibouti

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5280-442: Is consumed by animals while still alive and enters the plant-based trophic system. After plants and animals die, the organic matter contained in them enters the detritus-based trophic system. Ecosystem respiration is the sum of respiration by all living organisms (plants, animals, and decomposers) in the ecosystem. Net ecosystem production is the difference between gross primary production (GPP) and ecosystem respiration. In

5445-639: Is discernible in pollen counts from the Tibetan Plateau, which also show that the South Asian Monsoon had already developed by the late Oligocene. Around 25.8 Ma, the South Asian Monsoon underwent an episode of major intensification brought on by the uplift of the Tibetan Plateau. A deep 400,000-year glaciated Oligocene-Miocene boundary event is recorded at McMurdo Sound and King George Island . The early Eocene climate

5610-447: Is eligible as an experimental site to study the evolution of the deep hot brines of the Red Sea. By observing the strontium isotope composition of the Red Sea brines, it is possible to deduce how these salt waters found at the bottom of the Red Sea could have evolved in a similar way to Lake Asal, which ideally represents their compositional extreme. The high salinity of the waters was not hospitable to living organisms. Sometime during

5775-569: Is followed by succession, a "directional change in ecosystem structure and functioning resulting from biotically driven changes in resource supply." The frequency and severity of disturbance determine the way it affects ecosystem function. A major disturbance like a volcanic eruption or glacial advance and retreat leave behind soils that lack plants, animals or organic matter. Ecosystems that experience such disturbances undergo primary succession . A less severe disturbance like forest fires, hurricanes or cultivation result in secondary succession and

5940-554: Is governed by three sets of factors—the physical environment (temperature, moisture, and soil properties), the quantity and quality of the dead material available to decomposers, and the nature of the microbial community itself. Temperature controls the rate of microbial respiration; the higher the temperature, the faster the microbial decomposition occurs. Temperature also affects soil moisture, which affects decomposition. Freeze-thaw cycles also affect decomposition—freezing temperatures kill soil microorganisms, which allows leaching to play

6105-405: Is no clear distinction between biomes and ecosystems. Biomes are always defined at a very general level. Ecosystems can be described at levels that range from very general (in which case the names are sometimes the same as those of biomes) to very specific, such as "wet coastal needle-leafed forests". Biomes vary due to global variations in climate . Biomes are often defined by their structure: at

6270-506: Is placed at the last appearance of the foraminiferan genus Hantkenina in a quarry at Massignano , Italy . However, this GSSP has been criticized as excluding the uppermost part of the type Eocene Priabonian Stage and because it is slightly earlier than important climate shifts that form natural markers for the boundary, such as the global oxygen isotope shift marking the expansion of Antarctic glaciation (the Oi1 event). The upper boundary of

6435-531: Is primarily achieved through bacterial and fungal action. Fungal hyphae produce enzymes that can break through the tough outer structures surrounding dead plant material. They also produce enzymes that break down lignin , which allows them access to both cell contents and the nitrogen in the lignin. Fungi can transfer carbon and nitrogen through their hyphal networks and thus, unlike bacteria, are not dependent solely on locally available resources. Decomposition rates vary among ecosystems. The rate of decomposition

6600-485: Is termed its resistance . The capacity of a system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Resilience thinking also includes humanity as an integral part of the biosphere where we are dependent on ecosystem services for our survival and must build and maintain their natural capacities to withstand shocks and disturbances. Time plays

6765-554: Is that some Asiatic languages used color words to refer to the cardinal directions . Herodotus on one occasion uses Red Sea and Southern Sea interchangeably. The name in Hebrew Yam Suph ( Hebrew : ים סוף , lit.   'Sea of Reeds') is of biblical origin. The name in Coptic : ⲫⲓⲟⲙ `ⲛϩⲁϩ Phiom Enhah ("Sea of Hah") is connected to Ancient Egyptian root ḥ-ḥ which refers to water and sea (for example

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6930-422: Is the "best" classification. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of the definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Different approaches to ecological classifications have been developed in terrestrial, freshwater and marine disciplines, and

7095-416: Is the driving force in the Red Sea to transport material as suspension or as bedload. Wind-induced currents play an important role in the Red Sea in resuspending bottom sediments and transferring materials from sites of dumping to sites of burial in quiescent environment of deposition. Wind-generated current measurement is therefore important in order to determine the sediment dispersal pattern and its role in

7260-473: The 2023-2024 Israel-Hamas war . One ship was hijacked and taken back to Yemen. The Red Sea is between arid land, desert and semi-desert . Reef systems are better developed along the Red Sea mainly because of its greater depths and an efficient water circulation pattern. The Red Sea water mass-exchanges its water with the Arabian Sea , Indian Ocean via the Gulf of Aden . These physical factors reduce

7425-581: The Bartonian to the Rupelian. The transition is marked by the Oi1 event, an oxygen isotope excursion occurring approximately 33.55 million years ago, during which oxygen isotope ratios decreased by 1.3 ‰ . About 0.3–0.4 ‰ of this is estimated to be due to major expansion of Antarctic ice sheets. The remaining 0.9 to 1.0 ‰ was due to about 5 to 6 °C (9 to 10 °F) of global cooling . The transition likely took place in three closely spaced steps over

7590-499: The Coptic ⲫⲓⲟⲙ ̀ⲛϣⲁⲣⲓ Phiom ̀nšari , Syriac ܝܡܐ ܣܘܡܩܐ Yammāʾ summāqā, Somali Badda cas and Tigrinya Qeyyiḥ bāḥrī (ቀይሕ ባሕሪ). The name of the sea may signify the seasonal blooms of the red-coloured Trichodesmium erythraeum near the water's surface. A theory favored by some modern scholars is that the name red is referring to the direction south, just as the Black Sea 's name may refer to north. The basis of this theory

7755-570: The Grande Coupure ; it featured the replacement of European fauna with Asian fauna , except for the endemic rodent and marsupial families. By contrast, the Oligocene–Miocene boundary is not set at an easily identified worldwide event but rather at regional boundaries between the warmer late Oligocene and the relatively cooler Miocene. The lower boundary of the Oligocene (its Global Boundary Stratotype Section and Point or GSSP)

7920-774: The Mediterranean Sea . To the point that a number of them now form a significant component of the Mediterranean ecosystem. The resulting change in biodiversity, without precedent in human memory, is currently accelerating, in particular for fishes according to surveys engaged by the Mediterranean Science Commission . The rich diversity of the Red Sea is in part due to the 2,000 km (1,240 mi) of coral reef extending along its coastline; these fringing reefs are 5000–7000 years old and are largely formed of stony acropora and porites corals. The reefs form platforms and sometimes lagoons along

8085-655: The Red Sea slave trade from Africa to the Middle East. During the Middle Ages , the Red Sea was an important part of the spice trade route. In 1183, Raynald of Châtillon launched a raid down the Red Sea to attack the Muslim pilgrim convoys to Mecca. The possibility that Raynald's fleet might sack the holy cities of Mecca and Medina caused fury throughout the Muslim world. However, it appears that Reynald's target

8250-810: The Second World War , the Americans and Soviets exerted their influence whilst the volume of oil tanker traffic intensified. However, the Six-Day War culminated in the closure of the Suez Canal from 1967 to 1975. Today, in spite of patrols by the major maritime fleets in the waters of the Red Sea, the Suez Canal has never recovered its supremacy over the Cape route, which is believed to be less vulnerable to piracy. Iranian -backed Yemini Houthis have attacked Western ships, including warships , during

8415-672: The Septuagint translation of the Book of Exodus from Hebrew to Koine Greek in approximately the third century BC. In that version, the Yam Suph ( Hebrew : ים סוף , lit.   'Sea of Reeds') is translated as Erythra Thalassa (Red Sea). The Red Sea is one of four seas named in English after common color terms – the others being the Black Sea , the White Sea and

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8580-788: The Tertiary period, the Bab el Mandeb closed and the Red Sea evaporated to an empty hot dry salt-floored sink. Effects causing this would have been: A number of volcanic islands rise from the center of the sea. Most are dormant. However, in 2007, Jabal al-Tair island in the Bab el Mandeb strait erupted violently. Two new islands were formed in 2011 and 2013 in the Zubair Archipelago , a small chain of islands owned by Yemen. The first island, Sholan Island, emerged in an eruption in December 2011,

8745-632: The Yellow Sea . The direct rendition of the Greek Erythra thalassa in Latin as Mare Erythraeum refers to the north-western part of the Indian Ocean , and also to a region on Mars . The earliest known exploration of the Red Sea was conducted by ancient Egyptians , as they attempted to establish commercial routes to Punt . One such expedition took place around 2500 BC, and another around 1500 BC (by Hatshepsut ). Both involved long voyages down

8910-426: The archaeocete cetaceans began to decrease in diversity due to their lack of echolocation, which was very useful as the water became colder and cloudier. Other factors to their decline could include climate changes and competition with today's modern cetaceans and the requiem sharks , which also appeared in this epoch. Early desmostylians , like Behemotops , are known from the Oligocene. Pinnipeds appeared near

9075-664: The creodonts that had dominated the Paleocene in the Old World. Rodents and rabbits underwent tremendous diversification due to the increase in suitable habitats for ground-dwelling seed eaters, as habitats for squirrel-like nut- and fruit-eaters diminished. The primates, once present in Eurasia, were reduced in range to Africa and South America. Many groups, such as equids , entelodonts , rhinos , merycoidodonts , and camelids , became more able to run during this time, adapting to

9240-968: The global economy , with cargo vessels traveling between the Indian Ocean and the Mediterranean Sea every year, thus shortening the path between Asia and Europe almost by half (as compared to traveling around Africa via the Atlantic Ocean). 12% of global trade passes through the Red Sea. This includes 30% of global container traffic. The sea is known for its recreational diving sites, such as Ras Mohammed , SS Thistlegorm (shipwreck), Elphinstone Reef , The Brothers , Daedalus Reef , St. John's Reef , Rocky Island in Egypt and less known sites in Sudan such as Sanganeb , Abington , Angarosh and Shaab Rumi . The Red Sea became

9405-555: The resource inputs are generally controlled by external processes like climate and parent material, the availability of these resources within the ecosystem is controlled by internal factors like decomposition, root competition or shading. Other factors like disturbance, succession or the types of species present are also internal factors. Primary production is the production of organic matter from inorganic carbon sources. This mainly occurs through photosynthesis . The energy incorporated through this process supports life on earth, while

9570-528: The sabkhas through a network of channels. However, south of Jeddah in the Shoiaba area, the water from the lagoon may cover the adjoining sabkhas as far as 3 km (2 mi), whereas north of Jeddah in the Al-Kharrar area the sabkhas are covered by a thin sheet of water as far as 2 km (1.2 mi). The prevailing north and northeast winds influence the movement of water in the coastal inlets to

9735-435: The "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . They also include less tangible items like tourism and recreation, and genes from wild plants and animals that can be used to improve domestic species. Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like

9900-438: The 44 recorded species of shark, and by many species (over 175) of nudibranch , many of which are endemic to the Red Sea. Other coastal Red Sea habitats include sea grass beds, salt pans , mangrove and salt marsh . Furthermore the deep Red Sea brine pools have been extensively studied with regard to their microbial life, characterized by its diversity and adaptation to extreme environments. The high marine biodiversity of

10065-604: The 6th century BC, Darius the Great , who was a prominent ruler of the Achaemenid Empire in Persia, undertook significant efforts to improve and extend navigation in the Red Sea. He sent reconnaissance missions to explore the Red Sea area and to identify its various navigational hazards, such as rocks and currents. This effort was significant, as it contributed to safer and more efficient navigation routes. In addition to

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10230-536: The Achaemenid Empire's economic and political power in the region. In the late 4th century BC, Alexander the Great sent Greek naval expeditions down the Red Sea to the Indian Ocean . Greek navigators continued to explore and compile data on the Red Sea. Agatharchides collected information about the sea in the 2nd century BC. The Periplus of the Erythraean Sea (" Periplus of the Red Sea "),

10395-600: The Antarctic Circumpolar Current. There is some evidence that it occurred much earlier, during the early Eocene. The other major oceanic gateway opening during this time was the Tasman, or Tasmanian, depending on the paper, gateway between Australia and Antarctica. The time frame for this opening is less disputed than the Drake Passage and is largely considered to have occurred around 34 Ma. As

10560-717: The Arctic may have influenced this oceanic cooling, though the extent of this influence is still a matter of some significant dispute. The opening and closing of ocean gateways: the opening of the Drake Passage ; the opening of the Tasmanian Gateway and the closing of the Tethys seaway; along with the final formation of the Greenland – Iceland – Faroes Ridge; played vital parts in reshaping oceanic currents during

10725-534: The Dong Ho Formation of Oligocene age shows that the Oligocene flora of what is now Vietnam was very similar to its present flora. Kelps make their first appearance in the fossil record during the earliest Oligocene. Most extant mammal families had appeared by the end of the Oligocene. These included primitive three-toed horses, rhinoceroses, camels, deer, and peccaries. Carnivores such as dogs , nimravids , bears, weasels, and raccoons began to replace

10890-399: The Drake Passage may also have been in place by the end of the early Oligocene. This may have been interrupted by a temporary constriction of the Drake Passage from sometime in the middle to late Oligocene (29 to 22 mya) to the middle Miocene (15 mya). The reorganization of the oceanic tectonic plates of the northeastern Pacific, which had begun in the Paleocene, culminated with the arrival of

11055-522: The Drake Passage opened and enabled the formation of the Antarctic Circumpolar Current (ACC), which would have kept the cold waters of Antarctica circulating around that continent and strengthened the formation of Antarctic Bottom Water (ABW). With the cold water concentrated around Antarctica, sea surface temperatures and, consequently, continental temperatures would have dropped. The onset of Antarctic glaciation occurred during

11220-916: The East African large igneous province , which also initiated rifting along the Red Sea and Gulf of Aden . The Alps were rapidly rising in Europe as the African Plate continued to push north into the Eurasian Plate , isolating the remnants of the Tethys Sea . Sea levels were lower in the Oligocene than in the early Eocene, exposing large coastal plains in Europe and the Gulf Coast and Atlantic Coast of North America. The Obik Sea , which had separated Europe from Asia, retreated early in

11385-489: The Eocene-Oligocene transition are not yet fully understood. The timing is wrong for this to be caused either by known impact events or by the volcanic activity on the Ethiopean Plateau. Two other possible drivers of climate change, not mutually exclusive, have been proposed. The first is thermal isolation of the continent of Antarctica by development of the Antarctic Circumpolar Current . Deep sea cores from south of New Zealand suggest that cold deep-sea currents were present by

11550-442: The Grande Coupure. The lowering of sea levels closed the Turgai Strait across the Obik Sea, which had previously separated Asia from Europe. This allowed Asian mammals, such as rhinoceroses and ruminants , to enter Europe and drive endemic species to extinction. Lesser faunal turnovers occurred simultaneously with the Oi2 event and towards the end of the Oligocene. There was significant diversification of mammals in Eurasia, including

11715-446: The Jurassic, but the exact timing of the establishment of ocean channels between the two continents remains uncertain. However, one estimate is that a deep channel was in place between the two continents by the end of the early Oligocene. The timing of the formation of the Drake Passage between South America and Antarctica is also uncertain, with estimates ranging from 49 to 17 mya (early Eocene to Miocene), but oceanic circulation through

11880-744: The Murray and Mendocino Fracture Zones at the North American subduction zone in the Oligocene. This initiated strike-slip movement along the San Andreas Fault and extensional tectonics in the Basin and Range province , ended volcanism south of the Cascades, and produced clockwise rotation of many western North American terranes. The Rocky Mountains were at their peak. A new volcanic arc was established in western North America, far inland from

12045-504: The New World and Old World monkeys. The South American continent was home to animals such as pyrotheres and astrapotheres , as well as litopterns and notoungulates . Sebecosuchians , terror birds , and carnivorous metatheres , like the borhyaenids remained the dominant predators. Africa was also relatively isolated and retained its endemic fauna. These included mastodonts, hyraxes, arsinoitheres, and other archaic forms. Egypt in

12210-733: The Norwegian-Greenland Sea indicate a drop in winter temperatures at high latitudes of about 5 °C (9.0 °F) just prior to the Oi1 event. Borehole dating from the Southeast Faroes drift indicates that deep-ocean circulation from the Arctic Ocean to the North Atlantic Ocean began in the early Oligocene. The best terrestrial record of Oligocene climate comes from North America, where temperatures dropped by 7 to 11 °C (13 to 20 °F) in

12375-606: The Norwegian-Greenland Sea indicated that glaciers had appeared in Greenland by the start of the Oligocene. Continental ice sheets in Antarctica reached sea level during the transition. Glacially rafted debris of early Oligocene age in the Weddell Sea and Kerguelen Plateau , in combination with Oi1 isotope shift, provides unambiguous evidence of a continental ice sheet on Antarctica by the early Oligocene. The causes of

12540-427: The Oligocene exists mostly in isotopic proxies. Patterns of extinction and patterns of species migration can also be studied to gain insight into ocean conditions. For a while, it was thought that the glaciation of Antarctica may have significantly contributed to the cooling of the ocean, however, recent evidence tends to deny this. Isotopic evidence suggests that during the early Oligocene, the main source of deep water

12705-636: The Oligocene is defined by its GSSP at Carrosio , Italy , which coincides with the first appearance of the foraminiferan Paragloborotalia kugleri and with the base of magnetic polarity chronozone C6Cn.2n. Oligocene faunal stages from youngest to oldest are: During the Oligocene Epoch, the continents continued to drift toward their present positions. Antarctica became more isolated as deep ocean channels were established between Antarctica and Australia and South America . Australia had been very slowly rifting away from West Antarctica since

12870-516: The Oligocene was an environment of lush forested deltas. Nevertheless, the Early Oligocene saw a major reduction in the diversity of many Afro-Arabian mammal clades, including hyaenodonts, primates, and hystricognath and anomaluroid rodents. During the Oligocene, the Tethyan marine biodiversity hotspot collapsed as the Tethys Ocean contracted. The seas around Southeast Asia and Australia became

13035-526: The Oligocene, creating a persistent land connection between the continents. The Paratethys Sea stretched from what is now the Balkan Peninsula across Central Asia to the Tian Shan region of what is now Xinjiang . There appears to have been a land bridge in the early Oligocene between North America and Europe, since the faunas of the two regions are very similar. However, towards the end of

13200-492: The Oligocene, there was a brief marine incursion in Europe. The rise of the Himalayas during the Oligocene remains poorly understood. One recent hypothesis is that a separate microcontinent collided with south Asia in the early Eocene, and India itself did not collide with south Asia until the end of the Oligocene. The Tibetan Plateau may have reached nearly its present elevation by the late Oligocene. The Andes first became

13365-527: The Oligocene. The Oligocene saw the emergence of parrotfishes, as the centre of marine biodiversity shifted from the Central Tethys eastward into the Indo-Pacific. The fossil record of marine mammals is a little spotty during this time, and not as well known as the Eocene or Miocene, but some fossils have been found. The baleen whales and toothed whales had just appeared, and their ancestors,

13530-600: The Oligocene. As the continents shifted to a more modern configuration, so too did ocean circulation. The Drake Passage is located between South America and Antarctica . Once the Tasmanian Gateway between Australia and Antarctica opened, all that kept Antarctica from being completely isolated by the Southern Ocean was its connection to South America. As the South American continent moved north,

13695-423: The Red Sea and its coasts is extremely low, averaging 60 mm (2.36 in) per year. The rain is mostly short showers, often with thunderstorms and occasionally with dust storms . The scarcity of rainfall and no major source of fresh water to the Red Sea result in excess evaporation as high as 2,050 mm (81 in) per year and high salinity with minimal seasonal variation. A recent underwater expedition to

13860-492: The Red Sea inside the Bab al-Mandab , as the first fleet from Europe in modern times to have sailed these waters. Later in 1524 the city was delivered to Governor Heitor da Silveira as an agreement for protection from the Ottomans . In 1798, France ordered General Napoleon to invade Egypt and take control of the Red Sea. Although he failed in his mission, the engineer Jean-Baptiste Lepère , who took part in it, revitalised

14025-408: The Red Sea is dominated by persistent north-west winds , with speeds ranging between 7 km/h (4.3 mph) and 12 km/h (7.5 mph). The rest of the Red Sea and the Gulf of Aden are subjected to regular and seasonally reversible winds. The wind regime is characterized by seasonal and regional variations in speed and direction with average speed generally increasing northward. Wind

14190-465: The Red Sea offshore from Sudan and Eritrea found surface water temperatures 28 °C (82 °F) in winter and up to 34 °C (93 °F) in the summer, but despite that extreme heat, the coral was healthy with much fish life with very little sign of coral bleaching , with only 9% infected by Thalassomonas loyana , the 'white plague' agent. Favia favus coral there harbours a virus, BA3, which kills T. loyana . Scientists are investigating

14355-582: The Red Sea. The biblical Book of Exodus tells the account of the Israelites ' crossing of the sea , which the Hebrew text calls Yam Suph ( Hebrew : יַם סוּף ). Yam Suph was traditionally identified as the Red Sea. Rabbi Saadia Gaon (882‒942), in his Judeo-Arabic translation of the Pentateuch , identifies the crossing place of the Red Sea as Baḥar al-Qulzum , meaning the Gulf of Suez . In

14520-464: The Red Sea. The net value of the latter predominates, resulting in an overall drift to the north end of the Red Sea. Generally, the velocity of the tidal current is 50–60 cm/s (20–24 in/s) with a maximum of 1 m/s (3.3 ft/s) at the mouth of the al-Kharrar Lagoon. However, the range of the north-northeast current along the Saudi coast is 8–29 cm/s (3–11 in/s). The north part of

14685-481: The absence of disturbance, net ecosystem production is equivalent to the net carbon accumulation in the ecosystem. Energy can also be released from an ecosystem through disturbances such as wildfire or transferred to other ecosystems (e.g., from a forest to a stream to a lake) by erosion . In aquatic systems , the proportion of plant biomass that gets consumed by herbivores is much higher than in terrestrial systems. In trophic systems, photosynthetic organisms are

14850-418: The adjacent sabkhas, especially during storms. Winter mean sea level is 0.5 m (1.6 ft) higher than in summer. Tidal velocities passing through constrictions caused by reefs, sand bars and low islands commonly exceed 1–2 m/s (3–7 ft/s). Coral reefs in the Red Sea are near Egypt, Eritrea, Israel, Saudi Arabia, and Sudan. Detailed information regarding current data is lacking, partially because

15015-402: The amount of energy available to the ecosystem. Parent material determines the nature of the soil in an ecosystem, and influences the supply of mineral nutrients. Topography also controls ecosystem processes by affecting things like microclimate , soil development and the movement of water through a system. For example, ecosystems can be quite different if situated in a small depression on

15180-504: The amount of light available, the amount of leaf area a plant has to capture light (shading by other plants is a major limitation of photosynthesis), the rate at which carbon dioxide can be supplied to the chloroplasts to support photosynthesis, the availability of water, and the availability of suitable temperatures for carrying out photosynthesis. Energy and carbon enter ecosystems through photosynthesis, are incorporated into living tissue, transferred to other organisms that feed on

15345-542: The area is recognized by the Egyptian government, which set up the Ras Mohammed National Park in 1983. The rules and regulations governing this area protect local marine life, which has become a major draw for diving enthusiasts who should be aware that although most Red Sea species are innocuous, a few are hazardous to humans. There is extensive demand for desalinated water to meet the needs of

15510-550: The atmosphere, contributing to global cooling. The gradual separation of the clump of continental crust and the deepening of the tectonic ridge in the North Atlantic that would become Greenland, Iceland, and the Faroe Islands helped to increase the deep water flow in that area. More information about the evolution of North Atlantic Deep Water will be given a few sections down. Evidence for ocean-wide cooling during

15675-625: The beginnings of sediment drift deposition in the North Atlantic, such as the Feni and Southeast Faroe drifts. The chilling of the South Ocean deep water began in earnest once the Tasmanian Gateway and the Drake Passage opened fully. Regardless of the time at which the opening of the Drake Passage occurred, the effect on the cooling of the Southern Ocean would have been the same. Ecosystem An ecosystem (or ecological system )

15840-433: The biodiversity of the world ecosystems, reducing both their resilience and biocapacity . The report refers to natural systems as humanity's "life-support system", providing essential ecosystem services. The assessment measures 24 ecosystem services and concludes that only four have shown improvement over the last 50 years, 15 are in serious decline, and five are in a precarious condition. Red Sea The Red Sea

16005-408: The carbon makes up much of the organic matter in living and dead biomass, soil carbon and fossil fuels . It also drives the carbon cycle , which influences global climate via the greenhouse effect . Through the process of photosynthesis, plants capture energy from light and use it to combine carbon dioxide and water to produce carbohydrates and oxygen . The photosynthesis carried out by all

16170-536: The coast and occasional other features such as cylinders (such as the Blue Hole (Red Sea) at Dahab ). The Red Sea also hosts many offshore reefs, including several true atolls. Many of the unusual offshore reef formations defy classic (i.e., Darwinian) coral reef classification schemes, and are generally attributed to the high levels of tectonic activity that characterize the area. Both offshore and coastal reefs are visited by pelagic species of fish, including many of

16335-1447: The coast of Africa ] and Aqaba [A line running from Ràs al Fasma Southwesterly to Requin Island ( 27°57′N 34°36′E  /  27.950°N 34.600°E  / 27.950; 34.600 ) through Tiran Island to the Southwest point thereof and thence Westward on a parallel (27°54'N) to the coast of the Sinai Peninsula ]. On the South. A line joining Husn Murad ( 12°40′N 43°30′E  /  12.667°N 43.500°E  / 12.667; 43.500 ) and Ras Siyyan ( 12°29′N 43°20′E  /  12.483°N 43.333°E  / 12.483; 43.333 ). Exclusive economic zones in Red Sea: Note: Hala'ib Triangle disputed between Sudan and Egypt and calculated for both. Red Sea has names in many languages (Modern Arabic : البحر الأحمر , romanized :  al-Baḥr al-ʾAḥmar , Medieval Arabic: بحر القلزم , romanized:  Baḥr al-Qulzum ; Biblical Hebrew : יַם-סוּף , romanized:  Yam Sūp̄ or Hebrew : הַיָּם הָאָדוֹם , romanized :  hayYām hāʾĀḏōm ; Coptic : ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ Phiom Enhah or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ Phiom ǹšari ; Tigrinya : ቀይሕ ባሕሪ Qeyih Bahri ; Somali : Badda Cas ; Afar : "Qasa Bad". Red Sea

16500-747: The coast, reaching from central Mexico through the Mogollon-Datil volcanic field to the San Juan volcanic field , then through Utah and Nevada to the ancestral Northern Cascades. Huge ash deposits from these volcanoes created the White River and Arikaree Groups of the High Plains, with their excellent fossil beds. Between 31 and 26 mya, the Ethiopia-Yemen Continental Flood Basalts were emplaced by

16665-432: The combustion of fossil fuels, ammonia gas which evaporates from agricultural fields which have had fertilizers applied to them, and dust. Anthropogenic nitrogen inputs account for about 80% of all nitrogen fluxes in ecosystems. When plant tissues are shed or are eaten, the nitrogen in those tissues becomes available to animals and microbes. Microbial decomposition releases nitrogen compounds from dead organic matter in

16830-418: The concept to draw attention to the importance of transfers of materials between organisms and their environment. He later refined the term, describing it as "The whole system, ... including not only the organism-complex, but also the whole complex of physical factors forming what we call the environment". Tansley regarded ecosystems not simply as natural units, but as "mental isolates". Tansley later defined

16995-468: The currents are weak and both spatially and temporally variable. The variation of temporal and spatial currents is as low as 0.5 m (1.6 ft) and are governed all by wind. During the summer, northwesterly winds drive surface water south for about four months at a velocity of 15–20 cm/s (6–8 in/s), whereas in winter the flow is reversed resulting in the inflow of water from the Gulf of Aden into

17160-403: The dead organic matter would accumulate in an ecosystem, and nutrients and atmospheric carbon dioxide would be depleted. Decomposition processes can be separated into three categories— leaching , fragmentation and chemical alteration of dead material. As water moves through dead organic matter, it dissolves and carries with it the water-soluble components. These are then taken up by organisms in

17325-447: The definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Biotic factors of the ecosystem are living things; such as plants, animals, and bacteria, while abiotic are non-living components; such as water, soil and atmosphere. Plants allow energy to enter the system through photosynthesis , building up plant tissue. Animals play an important role in

17490-442: The difficulty may be that there were strong regional variations in the response to climate shifts. Evidence of a relatively warm Oligocene suggests an enigmatic climate state, neither hothouse nor icehouse. The late Oligocene (26.5 to 24 mya) likely saw a warming trend in spite of low pCO2 levels, though this appears to vary by region. However, Antarctica remained heavily glaciated during this warming period. The late Oligocene warming

17655-414: The earliest Oligocene. This change is seen from Alaska to the Gulf Coast. Upper Eocene paleosols reflect annual precipitation of over a meter of rain, but early Oligocene precipitation was less than half this. In central North America, the cooling was by 8.2 ± 3.1 °C over a period of 400,000 years, though there is little indication of significant increase in aridity during this interval. Ice-rafted debris in

17820-494: The early Oligocene, and the effect of the Drake Passage opening on this glaciation has been the subject of much research. However, some controversy still exists as to the exact timing of the passage opening, whether it occurred at the start of the Oligocene or nearer the end. Even so, many theories agree that at the Eocene/Oligocene (E/O) boundary, a yet shallow flow existed between South America and Antarctica, permitting

17985-490: The early Oligocene. However, the timing of this event remains controversial. The other possibility, for which there is considerable evidence, is a drop in atmospheric carbon dioxide levels ( pCO2 ) during the transition. The pCO2 is estimated to have dropped just before the transition, to 760 ppm at the peak of ice sheet growth, then rebounded slightly before resuming a more gradual fall. Climate modeling suggests that glaciation of Antarctica took place only when pCO2 dropped below

18150-655: The ecosystem or to gradual disruption of biotic processes and degradation of abiotic conditions of the ecosystem. Once the original ecosystem has lost its defining features, it is considered "collapsed ". Ecosystem restoration can contribute to achieving the Sustainable Development Goals . An ecosystem (or ecological system) consists of all the organisms and the abiotic pools (or physical environment) with which they interact. The biotic and abiotic components are linked together through nutrient cycles and energy flows. "Ecosystem processes" are

18315-528: The ecosystem. Internal factors are controlled, for example, by decomposition , root competition, shading, disturbance, succession, and the types of species present. While the resource inputs are generally controlled by external processes, the availability of these resources within the ecosystem is controlled by internal factors. Therefore, internal factors not only control ecosystem processes but are also controlled by them. Ecosystems are dynamic entities—they are subject to periodic disturbances and are always in

18480-423: The effect of high salinity caused by evaporation in the north and relatively hot water in the south. The climate of the Red Sea is the result of two monsoon seasons: a northeasterly monsoon and a southwesterly monsoon. Monsoon winds occur because of differential heating between the land and the sea. Very high surface temperatures and high salinities make this one of the warmest and saltiest bodies of seawater in

18645-430: The end of the epoch from an otter -like ancestor. The Oligocene sees the beginnings of modern ocean circulation, with tectonic shifts causing the opening and closing of ocean gateways. Cooling of the oceans had already commenced by the Eocene/Oligocene boundary, and they continued to cool as the Oligocene progressed. The formation of permanent Antarctic ice sheets during the early Oligocene and possible glacial activity in

18810-652: The erosion and accretion of the coastal rock exposure and the submerged coral beds. The Red Sea was formed by the Arabian Peninsula being split from the Horn of Africa by movement of the Red Sea Rift . This split started in the Eocene and accelerated during the Oligocene . The sea is still widening (in 2005, following a three-week period of tectonic activity it had grown by 8 m [26 ft]), and it

18975-421: The flow of energy through a lake was the primary driver of the ecosystem. Hutchinson's students, brothers Howard T. Odum and Eugene P. Odum , further developed a "systems approach" to the study of ecosystems. This allowed them to study the flow of energy and material through ecological systems. Ecosystems are controlled by both external and internal factors. External factors, also called state factors, control

19140-458: The forests of eastern North America still show legacies of cultivation which ceased in 1850 when large areas were reverted to forests. Another example is the methane production in eastern Siberian lakes that is controlled by organic matter which accumulated during the Pleistocene . Ecosystems continually exchange energy and carbon with the wider environment . Mineral nutrients, on

19305-804: The gateway widened, the Antarctic Circumpolar Current strengthened. The Tethys Seaway was not a gateway, but rather a sea in its own right. Its closing during the Oligocene had significant impact on both ocean circulation and climate. The collisions of the African plate with the European plate and of the Indian subcontinent with the Asian plate, cut off the Tethys Seaway that had provided a low-latitude ocean circulation. The closure of Tethys built some new mountains (the Zagros range) and drew down more carbon dioxide from

19470-527: The giant indricotheres , that grew up to 6 meters (20 ft) at the shoulder and weighed up to 20 tons. Paraceratherium was one of the largest land mammals ever to walk the Earth. However, the indricotheres were an exception to a general tendency for Oligocene mammals to be much smaller than their Eocene counterparts. The earliest deer, giraffes, pigs, and cattle appeared in the mid-Oligocene in Eurasia. The first felid , Proailurus , originated in Asia during

19635-691: The landscape, versus one present on an adjacent steep hillside. Other external factors that play an important role in ecosystem functioning include time and potential biota , the organisms that are present in a region and could potentially occupy a particular site. Ecosystems in similar environments that are located in different parts of the world can end up doing things very differently simply because they have different pools of species present. The introduction of non-native species can cause substantial shifts in ecosystem function. Unlike external factors, internal factors in ecosystems not only control ecosystem processes but are also controlled by them. While

19800-452: The late Oligocene and spread to Europe. There was only limited migration between Asia and North America. The cooling of central North America at the Eocene-Oligocene transition resulted in a large turnover of gastropods, amphibians, and reptiles. Mammals were much less affected. Crocodilians and pond turtles replaced by dry land tortoises. Molluscs shifted to more drought-tolerant forms. The White River Fauna of central North America inhabited

19965-423: The living and dead plant matter, and eventually released through respiration. The carbon and energy incorporated into plant tissues (net primary production) is either consumed by animals while the plant is alive, or it remains uneaten when the plant tissue dies and becomes detritus . In terrestrial ecosystems , the vast majority of the net primary production ends up being broken down by decomposers . The remainder

20130-408: The maintenance of hydrological cycles, cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. While material from the ecosystem had traditionally been recognized as being the basis for things of economic value, ecosystem services tend to be taken for granted. The Millennium Ecosystem Assessment

20295-405: The maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. Many ecosystems become degraded through human impacts, such as soil loss , air and water pollution , habitat fragmentation , water diversion , fire suppression , and introduced species and invasive species . These threats can lead to abrupt transformation of

20460-536: The major constituents of the Red Sea sediments are as follows: The Red Sea is a rich and diverse ecosystem. For example more than 1200 fish species have been recorded in the Red Sea, of which 10% are endemic, found nowhere else. Since the opening of the Suez Canal in November 1869, over a thousand marine species from the Red Sea—from plankton, seaweeds, invertebrates to fishes—have migrated northward and settled in

20625-535: The maritime explorations, during the reign of Darius the Great, a canal was constructed linking the Nile River to the northern end of the Red Sea at Suez. This canal is sometimes referred to as the ancient Suez Canal. It played a pivotal role in improving trade and communication between the Nile Valley and the Red Sea, and beyond to the Indian Ocean. This canal was a predecessor to the modern Suez Canal , which

20790-433: The movement of matter and energy through the system, by feeding on plants and on one another. They also influence the quantity of plant and microbial biomass present. By breaking down dead organic matter , decomposers release carbon back to the atmosphere and facilitate nutrient cycling by converting nutrients stored in dead biomass back to a form that can be readily used by plants and microbes. Ecosystems provide

20955-572: The names of the Ogdoad gods Heh and Hauhet ). Historically, it was also known to western geographers as Mare Mecca (Sea of Mecca), and Sinus Arabicus (Gulf of Arabia). Some ancient geographers called the Red Sea the Arabian Gulf or Gulf of Arabia. The association of the Red Sea with the biblical account of the Israelites crossing the Red Sea is ancient, and was made explicit in

21120-506: The new dominant hotspot of marine biodiversity. At sea, 97% of marine snail species, 89% of clams, and 50% of echinoderms of the Gulf Coast did not survive past the earliest Oligocene. New species evolved, but the overall diversity diminished. Cold-water mollusks migrated around the Pacific Rim from Alaska and Siberia. The marine animals of Oligocene oceans resembled today's fauna, such as the bivalves . Calcareous cirratulids appeared in

21285-519: The northern Red Sea. The route had been used by previous states but grew in the volume of traffic under the Romans. From Indian ports goods from China were introduced to the Roman world. Contact between Rome and China depended on the Red Sea, but the route was broken by the Aksumite Empire around the 3rd century AD. From antiquity until the 20th-century , the Red Sea was also a trade route of

21450-574: The opening of the Drake Passage is around 30 Ma. One of the possible issues with this timing was the continental debris cluttering up the seaway between the two plates in question. This debris, along with what is known as the Shackleton Fracture Zone , has been shown in a recent study to be fairly young, only about 8 million years old. The study concludes that the Drake Passage would be free to allow significant deep water flow by around 31 Ma. This would have facilitated an earlier onset of

21615-467: The opening of the Drake Passage is during the early Miocene. Despite the shallow flow between South America and Antarctica, there was not enough of a deep water opening to allow for significant flow to create a true Antarctic Circumpolar Current. If the opening occurred as late as hypothesized, then the Antarctic Circumpolar Current could not have had much of an effect on early Oligocene cooling, as it would not have existed. The earliest hypothesized time for

21780-537: The other hand, are mostly cycled back and forth between plants, animals, microbes and the soil. Most nitrogen enters ecosystems through biological nitrogen fixation , is deposited through precipitation, dust, gases or is applied as fertilizer . Most terrestrial ecosystems are nitrogen-limited in the short term making nitrogen cycling an important control on ecosystem production. Over the long term, phosphorus availability can also be critical. Macronutrients which are required by all plants in large quantities include

21945-509: The other. Despite this, the cumulative effect of additional species in an ecosystem is not linear: additional species may enhance nitrogen retention, for example. However, beyond some level of species richness, additional species may have little additive effect unless they differ substantially from species already present. This is the case for example for exotic species . The addition (or loss) of species that are ecologically similar to those already present in an ecosystem tends to only have

22110-403: The overall structure of an ecosystem and the way things work within it, but are not themselves influenced by the ecosystem. On broad geographic scales, climate is the factor that "most strongly determines ecosystem processes and structure". Climate determines the biome in which the ecosystem is embedded. Rainfall patterns and seasonal temperatures influence photosynthesis and thereby determine

22275-514: The period from 33.8 to 33.5 mya. By the end of the transition, sea levels had dropped by 105 meters (344 ft), and ice sheets were 25% greater in extent than in the modern world. The effects of the transition can be seen in the geological record at many locations around the world. Ice volumes rose as temperature and sea levels dropped. Playa lakes of the Tibetan Plateau disappeared at the transition, pointing to cooling and aridification of central Asia. Pollen and spore counts in marine sediments of

22440-640: The plains that were spreading as the Eocene rainforests receded. Brontotheres died out in the Earliest Oligocene, and creodonts died out outside Africa and the Middle East at the end of the period. Multituberculates , an ancient lineage of primitive mammals that originated back in the Jurassic , also became extinct in the Oligocene, aside from the gondwanatheres . The Eocene-Oligocene transition in Europe and Asia has been characterized as

22605-519: The plan for a canal which had been envisaged during the reign of the Pharaohs . Several canals were built in ancient times from the Nile to the Red Sea along or near the line of the present Sweet Water Canal , but none lasted for long. The Suez Canal was opened in November 1869. During the first half of the 20th-century, the Red Sea slave trade attracted substantional international condemnation. After

22770-405: The plants in an ecosystem is called the gross primary production (GPP). About half of the gross GPP is respired by plants in order to provide the energy that supports their growth and maintenance. The remainder, that portion of GPP that is not used up by respiration, is known as the net primary production (NPP). Total photosynthesis is limited by a range of environmental factors. These include

22935-428: The population and the industries along the Red Sea. There are at least 18 desalination plants along the Red Sea coast of Saudi Arabia which discharge warm brine and treatment chemicals ( chlorine and anti-scalants ) that bleach and kill corals and cause diseases in the fish. This is only localized, but it may intensify with time and profoundly impact the fishing industry. The Red Sea serves an important role in

23100-585: The primary nutrients (which are most limiting as they are used in largest amounts): Nitrogen, phosphorus, potassium. Secondary major nutrients (less often limiting) include: Calcium, magnesium, sulfur. Micronutrients required by all plants in small quantities include boron, chloride, copper, iron, manganese, molybdenum, zinc. Finally, there are also beneficial nutrients which may be required by certain plants or by plants under specific environmental conditions: aluminum, cobalt, iodine, nickel, selenium, silicon, sodium, vanadium. Until modern times, nitrogen fixation

23265-413: The primary producers. The organisms that consume their tissues are called primary consumers or secondary producers — herbivores . Organisms which feed on microbes ( bacteria and fungi ) are termed microbivores . Animals that feed on primary consumers— carnivores —are secondary consumers. Each of these constitutes a trophic level. The sequence of consumption—from plant to herbivore, to carnivore—forms

23430-419: The process of recovering from some past disturbance. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance, is termed its resistance . The capacity of a system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Ecosystems can be studied through

23595-516: The sea. The Red Sea is the world's northernmost tropical sea, and has been designated a Global 200 ecoregion. The International Hydrographic Organization defines the limits of the Red Sea as follows: On the North. The Southern limits of the Gulfs of Suez [A line running from Ràs Muhammed (27°43'N) to the South point of Shadwan Island (34°02'E) and thence Westward on a parallel (27°27'N) to

23760-526: The second island, Jadid, emerged in September 2013. Approximately 40% of the Red Sea is quite shallow at less than 100 m (330 ft) deep, with about 25% less than 50 m (160 ft) deep. Undiscovered oil reserves in the region have been estimated at 5,041 MMBO. Undiscovered gas reserves in the region have been estimated at 112,349 BCFG. Undiscovered natural gas reserves have been estimated at 3,077 MMBNGL. Most of these plays are controlled by

23925-611: The soil, react with mineral soil, or are transported beyond the confines of the ecosystem (and are considered lost to it). Newly shed leaves and newly dead animals have high concentrations of water-soluble components and include sugars , amino acids and mineral nutrients. Leaching is more important in wet environments and less important in dry ones. Fragmentation processes break organic material into smaller pieces, exposing new surfaces for colonization by microbes. Freshly shed leaf litter may be inaccessible due to an outer layer of cuticle or bark , and cell contents are protected by

24090-511: The soil, where plants, fungi, and bacteria compete for it. Some soil bacteria use organic nitrogen-containing compounds as a source of carbon, and release ammonium ions into the soil. This process is known as nitrogen mineralization . Others convert ammonium to nitrite and nitrate ions, a process known as nitrification . Nitric oxide and nitrous oxide are also produced during nitrification. Under nitrogen-rich and oxygen-poor conditions, nitrates and nitrites are converted to nitrogen gas ,

24255-437: The south near the Gulf of Aden, but it fluctuates between 0.20 m (0.66 ft) and 0.30 m (0.98 ft) away from the nodal point. The central Red Sea (Jeddah area) is therefore almost tideless, and as such the annual water level changes are more significant. Because of the small tidal range the water during high tide inundates the coastal sabkhas as a thin sheet of water up to a few hundred metres rather than flooding

24420-479: The southern part and 41 ‰ in the northern part around the Gulf of Suez , with an average of 40 ‰. (Average salinity for the world's seawater is ~35 ‰ on the Practical Salinity Scale, or PSU; that translates to 3.5% of actual dissolved salts). In general, tide ranges between 0.6 m (2.0 ft) in the north, near the mouth of the Gulf of Suez and 0.9 m (3.0 ft) in

24585-483: The spatial extent of ecosystems using the term " ecotope ". G. Evelyn Hutchinson , a limnologist who was a contemporary of Tansley's, combined Charles Elton 's ideas about trophic ecology with those of Russian geochemist Vladimir Vernadsky . As a result, he suggested that mineral nutrient availability in a lake limited algal production . This would, in turn, limit the abundance of animals that feed on algae. Raymond Lindeman took these ideas further to suggest that

24750-479: The start of an Antarctic Circumpolar Current. Stemming from the issue of when the opening of the Drake Passage took place, is the dispute over how great of an influence the opening of the Drake Passage had on the global climate. While early researchers concluded that the advent of the ACC was highly important, perhaps even the trigger, for Antarctic glaciation and subsequent global cooling, other studies have suggested that

24915-641: The structure of the basin. Normal faults are common as the Red Sea occupies an active diverging margin. These targets are commonly found below the Salt deposits of the Middle Miocene . Modern development is focused on the following fields. The Durwara 2 Field was discovered in 1963, while the Suakin 1 Field and the Bashayer 1A Field were discovered in 1976, on the Egyptian side of the Red Sea. The Barqan Field

25080-580: The time of the Oi2 oxygen isotope shift. This led to the largest drop of sea level in past 100 million years, by about 75 meters (246 ft). This is reflected in a mid-Oligocene incision of continental shelves and unconformities in marine rocks around the world. Some evidence suggests that the climate remained warm at high latitudes even as ice sheets experienced cyclical growth and retreat in response to orbital forcing and other climate drivers. Other evidence indicates significant cooling at high latitudes. Part of

25245-421: The transfers of energy and materials from one pool to another. Ecosystem processes are known to "take place at a wide range of scales". Therefore, the correct scale of study depends on the question asked. The term "ecosystem" was first used in 1935 in a publication by British ecologist Arthur Tansley . The term was coined by Arthur Roy Clapham , who came up with the word at Tansley's request. Tansley devised

25410-442: The unique properties of these coral and their commensal algae to see if they can be used to salvage bleached coral elsewhere. The Red Sea is one of the saltiest bodies of water in the world, owing to high evaporation and low precipitation; no significant rivers or streams drain into the sea, and its southern connection to the Gulf of Aden , an arm of the Indian Ocean, is narrow. Its salinity ranges from between ~36  ‰ in

25575-416: The world as tropical and sub- tropical forests were replaced by temperate deciduous forests . Open plains and deserts became more common and grasses expanded from their water-bank habitat in the Eocene moving out into open tracts. The decline in pCO2 favored C4 photosynthesis , which is found only in angiosperms and is particularly characteristic of grasses. However, even at the end of the period, grass

25740-481: The world. The average surface water temperature of the Red Sea during the summer is about 26 °C (79 °F) in the north and 30 °C (86 °F) in the south, with only about 2 °C (3.6 °F) variation during the winter months. The overall average water temperature is 22 °C (72 °F). Temperature and visibility remain good to around 200 m (660 ft). The sea is known for its strong winds and unpredictable local currents. The rainfall over

25905-410: The δO signature is too strong for glaciation to be the main trigger for cooling. Through study of Pacific Ocean sediments, other researchers have shown that the transition from warm Eocene ocean temperatures to cool Oligocene ocean temperatures took only 300,000 years, which strongly implies that feedbacks and factors other than the ACC were integral to the rapid cooling. The latest hypothesized time for

26070-514: Was closed to all swimming in December 2010 due to several serious shark attacks , including a fatality. As of December 2010, scientists are investigating the attacks and have identified, but not verified, several possible causes including over-fishing which causes large sharks to hunt closer to shore, tourist boat operators who chum offshore for shark-photo opportunities, and reports of ships throwing dead livestock overboard. The sea's narrowness, significant depth, and sharp drop-offs, all combine to form

26235-481: Was constructed in the 19th century and continues to be one of the world's most important waterways. The construction of the canal during Darius's reign is evidenced by ancient records, including inscriptions. Darius commemorated the completion of the canal by creating stelae (stone monuments) with inscriptions in several languages, describing the construction and its benefits. The canal not only facilitated trade but also solidified Darius's control over Egypt and enhanced

26400-736: Was discovered in 1969, and the Midyan Field in 1992, both within the Midyan Basin on the Saudi Arabian side of the Red Sea. The 20-m thick Middle Miocene Maqna Formation is an oil source rock in the basin. Oil seeps occur near the Farasan Islands , the Dahlak Archipelago , along the coast of Eritrea, and in the southeastern Red Sea along the coasts of Saudi Arabia and Yemen. In terms of mineral resources

26565-539: Was not quite common enough for modern savannas . In North America, much of the dense forest was replaced by patchy scrubland with riparian forests. Subtropical species dominated with cashews and lychee trees present, and temperate woody plants such as roses , beeches , and pines were common. The legumes spread, while sedges and ferns continued their ascent. In Europe, floral assemblages became increasingly affected by strengthening seasonality as it related to wildfire activity. The Ha Long megafossil flora from

26730-771: Was the North Pacific and the Southern Ocean . As the Greenland-Iceland-Faroe Ridge sank and thereby connected the Norwegian–Greenland sea with the Atlantic Ocean, the deep water of the North Atlantic began to come into play as well. Computer models suggest that once this occurred, a more modern in appearance thermo-haline circulation started. Evidence for the early Oligocene onset of chilled North Atlantic deep water lies in

26895-399: Was the lightly armed Muslim pilgrim convoys, rather than the well guarded cities of Mecca and Medina, and the belief in the Muslim world that Reynald was seeking to sack the holy cities, due to the proximity of those cities to the areas that Raynald raided. In 1513, trying to secure that channel to Portugal, Afonso de Albuquerque laid siege to Aden but was forced to retreat. They cruised

27060-679: Was the major source of nitrogen for ecosystems. Nitrogen-fixing bacteria either live symbiotically with plants or live freely in the soil. The energetic cost is high for plants that support nitrogen-fixing symbionts—as much as 25% of gross primary production when measured in controlled conditions. Many members of the legume plant family support nitrogen-fixing symbionts. Some cyanobacteria are also capable of nitrogen fixation. These are phototrophs , which carry out photosynthesis. Like other nitrogen-fixing bacteria, they can either be free-living or have symbiotic relationships with plants. Other sources of nitrogen include acid deposition produced through

27225-596: Was very warm, with crocodilians and temperate plants thriving north of the Arctic Circle . The cooling trend that began in the middle Eocene continued into the Oligocene, bringing both poles well below freezing for the first time in the Phanerozoic . The cooling climate, together with the opening of some land bridges and the closing of others, led to a profound reorganization of the biosphere and loss of taxonomic diversity. Land animals and marine organisms reached

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