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95-458: 8, see text Pelicans (genus Pelecanus ) are a genus of large water birds that make up the family Pelecanidae . They are characterized by a long beak and a large throat pouch used for catching prey and draining water from the scooped-up contents before swallowing. They have predominantly pale plumage, except for the brown and Peruvian pelicans . The bills, pouches, and bare facial skin of all pelicans become brightly coloured before

190-587: A clade containing otherwise seabirds , and also symplesiomorphic with sulids but synapomorphic with cormorants in some other respects. Thus, the Sulidae seem to be the oldest and most distinct lineage of those three, which are united in a suborder Sulae . Therein, the Sulidae are typically placed simply as a family ; sometimes, a superfamily Suloidea is recognized, wherein some of the primitive prehistoric forms (e.g. Empheresula , Eostega , and Masillastega ) are placed as basal lineages distinct from

285-592: A complex communal courtship involving a group of males chasing a single female in the air, on land, or in the water while pointing, gaping, and thrusting their bills at each other. They can finish the process in a day. The tree-nesting species have a simpler process in which perched males advertise for females. The location of the breeding colony is constrained by the availability of an ample supply of fish to eat, although pelicans can use thermals to soar and commute for hundreds of kilometres daily to fetch food. The Australian pelican has two reproductive strategies depending on

380-407: A genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of

475-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

570-588: A long history of cultural significance in mythology , and in Christian and heraldic iconography . The name comes from the Ancient Greek word pelekan (πελεκάν), which is itself derived from the word pelekys (πέλεκυς) meaning "axe". In classical times, the word was applied to both the pelican and the woodpecker. The genus Pelecanus was first formally described by Carl Linnaeus in his landmark 1758 10th edition of Systema Naturae . He described

665-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

760-659: A lower height than the brown pelican. The Australian and American white pelicans may feed by low plunge-dives landing feet-first and then scooping up the prey with the beak, but they—as well as the remaining pelican species—primarily feed while swimming on the water. Aquatic prey is most commonly taken at or near the water surface. Although principally a fish eater, the Australian pelican is also an eclectic and opportunistic scavenger and carnivore that forages in landfill sites, as well as taking carrion and "anything from insects and small crustaceans to ducks and small dogs". Food

855-516: A phenomenon known as ground effect to reduce drag and increase lift . As the air flows between the wings and the water surface, it is compressed to a higher density and exerts a stronger upward force against the bird above. Hence, substantial energy is saved while flying. Adult pelicans rely on visual displays and behaviour to communicate, particularly using their wings and bills. Agonistic behaviour consists of thrusting and snapping at opponents with their bills, or lifting and waving their wings in

950-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

1045-533: A response to periods of stress has been recorded. The sulids are distributed mainly in tropical and subtropical waters, but they, particularly gannets, are found in temperate regions, too. These birds are not truly pelagic seabirds like the related Procellariiformes , and usually stay rather close to the coasts, but the abundant colonies of sulids that exist on many Pacific islands suggest that they are not infrequently blown away from their home range by storms , and can wander for long distances in search of

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1140-514: A safe place to land if need be. All species feed entirely at sea, mostly on mid-sized fish and similarly sized marine invertebrates (e.g. cephalopods ). Many species feed communally, and some species follow fishing boats to scavenge discarded bycatch and chum . The typical hunting behavior is a dive from midair, taking the bird a 1–2 m under water. If prey manages to escape the diving birds at first, they may give chase using their legs and wings for underwater swimming. As noted above,

1235-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

1330-414: A threatening manner. Adult pelicans grunt when at the colony, but are generally silent elsewhere or outside breeding season. Conversely, colonies are noisy, as chicks vocalise extensively. Pelicans are gregarious and nest colonially. Pairs are monogamous for a single season, but the pair bond extends only to the nesting area; mates are independent away from the nest. The ground-nesting (white) species have

1425-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

1520-409: A valid description. Fossil finds from North America have been meagre compared with Europe, which has a richer fossil record. Several Pelecanus species have been described from fossil material, including: Pelicans are very large birds with very long bills characterised by a downcurved hook at the end of the upper mandible, and the attachment of a huge gular pouch to the lower. The slender rami of

1615-544: A varied taxonomic history. Tropicbirds , darters , cormorants, gannets, boobies, and frigatebirds, all traditional members of the order, have since been reclassified: tropicbirds into their own order, Phaethontiformes , and the remainder into the Suliformes. In their place, herons, ibises, spoonbills, the hamerkop, and the shoebill have now been transferred into the Pelecaniformes. Molecular evidence suggests that

1710-435: A week, they are able to put their heads into their parents' pouches and feed themselves. Sometimes before, but especially after being fed the pelican chick may seem to "throw a tantrum" by loudly vocalizing and dragging itself around in a circle by one wing and leg, striking its head on the ground or anything nearby and the tantrums sometimes end in what looks like a seizure that results in the chick falling briefly unconscious;

1805-569: A wingspan around 140 to 175 cm (4.59 to 5.74 ft). They have long, narrow, and pointed wings, and a quite long, graduated, and rather lozenge -shaped tail whose outer feathers are shorter than the central ones. Their flight muscles are rather small to allow for the small cross-section required for plunge-diving, as an adaptive trade-off relative to some sacrifice in flight performance. Consequently, they are very streamlined , reducing drag , so their bodies are " torpedo -shaped" and somewhat flat. They have stout legs and webbed feet , with

1900-464: A wingspan of as little as 1.83 m (6.0 ft). The largest is believed to be the Dalmatian, at up to 15 kg (33 lb) and 1.83 m (6.0 ft) in length, with a maximum wingspan of 3 m (9.8 ft). The Australian pelican's bill may grow up to 0.5 m (1.6 ft) long in large males, the longest of any bird. Pelicans have mainly light-coloured plumage, the exceptions being

1995-658: Is considered to be near threatened in the IUCN Red List of Threatened Species. Numbers declined substantially during the 20th century, one crucial factor being the eradication of the important Sittaung valley breeding colony in Burma through deforestation and the loss of feeding sites. The chief threats it faces are from habitat loss and human disturbance, but populations have mostly stabilised following increased protection in India and Cambodia. Genus The composition of

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2090-460: Is darker than that of adults. Newly hatched chicks are naked and pink, darkening to grey or black after 4 to 14 days, then developing a covering of white or grey down . Anatomical dissections of two brown pelicans in 1939 showed that pelicans have a network of air sacs under their skin situated across the ventral surface including the throat, breast, and undersides of the wings, as well as having air sacs in their bones. The air sacs are connected to

2185-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

2280-555: Is frequent. Sulids are related to a number of other aquatic birds, which all lack external nostrils and a brood patch , but have all four toes webbed and a gular sac . The closest living relatives of the Sulidae are the Phalacrocoracidae (cormorants and shags) and the Anhingidae (darters). The latter are somewhat intermediate between sulids and cormorants, but (like many cormorants) they are freshwater birds in

2375-539: Is highly variable. Pelicans live for 15 to 25 years in the wild, although one reached an age of 54 years in captivity. The diet of pelicans usually consists of fish, but occasionally amphibians , turtles, crustaceans , insects , birds, and mammals are also eaten. The size of the preferred prey fish varies depending on pelican species and location. For example, in Africa, the pink-backed pelican generally takes fish ranging in size from fry up to 400 g (0.9 lb) and

2470-587: Is known from the Australian, brown, and Peruvian pelicans. Non-native great white pelicans have been observed swallowing city pigeons in St. James's Park in London, England. Globally, pelican populations are adversely affected by these main factors: declining supplies of fish through overfishing or water pollution, destruction of habitat, direct effects of human activity such as disturbance at nesting colonies, hunting and culling, entanglement in fishing lines and hooks, and

2565-629: Is not stored in a pelican's throat pouch, contrary to popular folklore. Pelicans may also eat birds. In southern Africa, eggs and chicks of the Cape cormorant are an important food source for great white pelicans. Several other bird species have been recorded in the diet of this pelican in South Africa, including Cape gannet chicks on Malgas Island as well as crowned cormorants , kelp gulls , greater crested terns , and African penguins on Dassen Island and elsewhere. The Australian pelican, which

2660-719: Is particularly willing to take a wide range of prey items, has been recorded feeding on young Australian white ibis , and young and adult grey teals and silver gulls . Brown pelicans have been reported preying on young common murres in California and the eggs and nestlings of cattle egrets and nestling great egrets in Baja California , Mexico. Peruvian pelicans in Chile have been recorded feeding on nestlings of imperial shags , juvenile Peruvian diving petrels , and grey gulls . Cannibalism of chicks of their own species

2755-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

2850-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

2945-420: Is the case today). Pelicans swim well with their strong legs and their webbed feet. They rub the backs of their heads on their preen glands to pick up an oily secretion, which they transfer to their plumage to waterproof it. Holding their wings only loosely against their bodies, pelicans float with relatively little of their bodies below the water surface. They dissipate excess heat by gular flutter – rippling

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3040-421: Is typically two eggs . The eggs are unmarked (but may become stained by debris in the nest), whitish, pale blue, green, or pink, and have a coating that resembles lime . Egg weight ranges from 3.3 to 8.0% of the female's weight. Incubation lasts 42 to 55 days, depending on the species. Both sexes incubate ; like their relatives, they do not have brood patches , but their feet become vascularized and hot, and

3135-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

3230-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

3325-473: The Luberon in southeastern France, and is remarkably similar to modern forms. Its beak is almost complete and is morphologically identical to that of present-day pelicans, showing that this advanced feeding apparatus was already in existence at the time. An Early Miocene fossil has been named Miopelecanus gracilis on the basis of certain features originally considered unique, but later thought to lie within

3420-547: The Old World and spread into the Americas; this is reflected in the relationships within the genus as the eight species divide into Old World and New World lineages. This hypothesis is supported by fossil evidence from the oldest pelican taxa. Pelicans will frequent inland waterways but are most known for residing along maritime and coastal zones, where they feed principally on fish in their large throat pouches, diving into

3515-605: The Oligocene -Miocene boundary about 23 Mya. Microsula , which lived during that time, seems to have been a primitive booby that still had many symplesiomorphies with gannets. Like the other Phalacrocoraciformes, the sulids originated probably in the general region of the Atlantic or western Tethys Sea – probably the latter rather than the former, given that their earliest fossils are abundant in Europe , but absent from

3610-669: The Pelecaniformes in its obsolete paraphyletic circumscription, but pelicans , the namesake family of the Pelecaniformes, are actually more closely related to herons , ibises and spoonbills , the hamerkop , and the shoebill than to the sulids and allies. In recognition of this, the Sulae have been proposed for separation in a new order Phalacrocoraciformes , which also includes the frigatebirds (Fregatidae), as well as one or more prehistoric lineages that are entirely extinct today. The IOC World Bird List uses Suliformes as

3705-456: The behavioral traits of gannets and boobies differ considerably, but the Sulidae as a whole are characterized by several behavioral synapomorphies : Before taking off, they point their bills upwards (gannets) or forward (boobies). After landing again, they point downwards with their bills. In response to a threat, they do not attack, but shake their heads and point their bills towards the intruders. All sulids breed in colonies . Males examine

3800-451: The genus Sula . However, Sula (true boobies) and Morus (gannets) can be readily distinguished by morphological , behavioral , and DNA sequence characters. Abbott's booby ( Papasula ) is given its own genus, as it stands apart from both in these respects. It appears to be a distinct and ancient lineage, maybe closer to the gannets than to the true boobies. Sulids measure about 60 to 85 cm (24 to 33 in) in length and have

3895-422: The lores . Unlike their relatives (the darters and cormorants ), sulids have a well-developed preen gland whose waxy secretions they spread on their feathers for waterproofing and pest control. They moult their tail feathers irregularly and the flight feathers of their wings in stages, so that starting at the first moult, they always have some old feathers, some new ones, and some partly grown ones. Moult as

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3990-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

4085-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

4180-480: The shoebill and hamerkop storks (although these two birds are not actually true 'storks'), and are placed in the order Pelecaniformes . Ibises , spoonbills , herons , and bitterns have been classified in the same order. Fossil evidence of pelicans dates back at least 36 million years to the remains of a tibiotarsus recovered from late Eocene strata of Egypt that bears striking similarity to modern species of pelican. They are thought to have evolved in

4275-697: The American state of Utah 's Great Salt Lake , for example, some 600 miles (965 km) from the nearest coastline (the Pacific West Coast). They have also been seen hundreds of miles inland in North America, having flown northwards along the Mississippi River and other large waterways. Long thought to be related to frigatebirds , cormorants , tropicbirds , and gannets and boobies , pelicans instead are most closely related to

4370-463: The Australian white pelican was their next-closest relative. The great white pelican also belonged to this lineage, but was the first to diverge from the common ancestor of the other four species. This finding suggests that pelicans evolved in the Old World and spread into the Americas, and that preference for tree- or ground-nesting is more related to size than genetics. The fossil record shows that

4465-413: The Dalmatian, pink-backed, and spot-billed pelicans are the only ones to prefer solitary feeding. When fishing in groups, all pelican species have been known to work together to catch their prey, and Dalmatian pelicans may even cooperate with great cormorants . Large fish are caught with the bill-tip, then tossed up in the air to be caught and slid into the gullet head-first. A gull will sometimes stand on

4560-621: The abdomen, where surface protuberances may be caused by viscera changing size and position) to enable the overlying feathers to form more effective heat insulation and also to enable feathers to be held in position for good aerodynamics. Modern pelicans are found on all continents except Antarctica. They primarily inhabit warm regions, although breeding ranges extend to latitudes of 45° South (Australian pelicans in Tasmania ) and 60° North (American white pelicans in western Canada). Birds of inland and coastal waters, they are absent from polar regions,

4655-429: The airways of the respiratory system, and the pelican can keep its air sacs inflated by closing its glottis , but how air sacs are inflated is not clear. The air sacs serve to keep the pelican remarkably buoyant in the water and may also cushion the impact of the pelican's body on the water surface when they dive from flight into water to catch fish. Superficial air sacs may also help to round body contours (especially over

4750-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

4845-453: The birds place the eggs under the webs. Eggs lost during the first half of incubation are replaced. At hatching, parents move the eggs and then the hatchlings to the tops of their webs. The young hatch naked, but soon develop white down . They beg by touching the parent's bill and take regurgitated food straight from its gape . At first, at least one parent is always in attendance of the altricial young; after two weeks, both parents leave

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4940-457: The breeding season. The eight living pelican species have a patchy, seasonally-dependent yet global distribution, ranging latitudinally from the tropics to the temperate zone . Pelicans are absent from interior Amazonian South America, from polar regions and the open ocean; at least one species is known to migrate to the inland desert of Australia 's Red Centre , after heavy rains create temporary lakes. White pelicans are also observed at

5035-581: The brown and Peruvian pelicans. The bills, pouches, and bare facial skin of all species become brighter before breeding season commences. The throat pouch of the Californian subspecies of the brown pelican turns bright red, and fades to yellow after the eggs are laid, while the throat pouch of the Peruvian pelican turns blue. The American white pelican grows a prominent knob on its bill that is shed once females have laid eggs. The plumage of immature pelicans

5130-522: The colony area in flight and then pick a nest site, which they defend by fighting and territorial displays . Males then advertise to females by a special display and call. Their display behavior is characteristic, though not as diverse as the numerous variations found among the cormorants ; it typically includes the male shaking his head. Females search the colony in flight and on foot for a mate. Once they select males, pairs maintain their bonds by preening each other and by frequent copulation. The clutch

5225-517: The deep ocean, oceanic islands (except the Galapagos), and inland South America, as well as from the eastern coast of South America from the mouth of the Amazon River southwards. Subfossil bones have been recovered from as far south as New Zealand's South Island , although their scarcity and isolated occurrence suggests that these remains may have merely been vagrants from Australia (much as

5320-460: The distinguishing characteristics as a straight bill hooked at the tip, linear nostrils, a bare face, and fully webbed feet. This early definition included frigatebirds, cormorants, and sulids , as well as pelicans. The family Pelecanidae was introduced (as Pelicanea) by the French polymath Constantine Samuel Rafinesque in 1815. Pelicans give their name to the Pelecaniformes, an order which has

5415-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

5510-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

5605-457: The global population of the brown pelican at 300,000. Numbers of brown pelican plummeted in the 1950s and 1960s, largely as a consequence of environmental DDT pollution, and the species was listed as endangered in the US in 1970. With restrictions on DDT use in the US from 1972, its population has recovered, and it was delisted in 2009. The Peruvian pelican is listed as near threatened because, although

5700-519: The great white pelican prefers somewhat larger fish, up to 600 g (1.3 lb), but in Europe, the latter species has been recorded taking fish up to 1,850 g (4.1 lb). In deep water, white pelicans often fish alone. Nearer the shore, several encircle schools of small fish or form a line to drive them into the shallows, beating their wings on the water surface and then scooping up the prey. Although all pelican species may feed in groups or alone,

5795-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

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5890-536: The late Miocene around 6 million years ago, after which time the boobies steadily diverged. The gannets split more recently, only around 2.5 million years ago. The fossil record of sulids is quite extensive due to the many Miocene / Pliocene forms that have been recovered, but the lineage of sulids extends back to the Eocene , and all things (such as the Early Eocene frigatebird Limnofregata ) considered,

5985-537: The living Sulidae. However, the proposed family Pseudosulidae (or Enkurosulidae ) is almost certainly invalid. Abbott's booby ( Papasula abbotti ) Northern gannet ( Morus bassanus ) Cape gannet ( Morus capensis ) Australasian gannet ( Morus serrator ) Red-footed booby ( Sula sula ) Brown booby ( Sula leucogaster ) Masked booby ( Sula dactylatra ) Nazca booby ( Sula granti ) Blue-footed booby ( Sula nebouxii ) Peruvian booby ( Sula variegata ) The Sulae were traditionally included in

6080-586: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Sulidae Morus Papasula Sula Enkurosulidae Kashin, 1977 Pseudosulidae Harrison, 1975 The bird family Sulidae comprises the gannets and boobies . Collectively called sulids , they are medium-large coastal seabirds that plunge-dive for fish and similar prey . The 10 species in this family are often considered congeneric in older sources, placing all in

6175-574: The local degree of environmental predictability. Colonies of tens or hundreds, rarely thousands, of birds breed regularly on small coastal and subcoastal islands where food is seasonally or permanently available. In arid inland Australia, especially in the endorheic Lake Eyre basin , pelicans breed opportunistically in very large numbers of up to 50,000 pairs, when irregular major floods, which may be many years apart, fill ephemeral salt lakes and provide large amounts of food for several months before drying out again. In all species, copulation takes place at

6270-401: The lower bill and the flexible tongue muscles form the pouch into a basket for catching fish, and sometimes rainwater, though to not hinder the swallowing of large fish, the tongue itself is tiny. They have a long neck and short stout legs with large, fully webbed feet. Although they are among the heaviest of flying birds, they are relatively light for their apparent bulk because of air pockets in

6365-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

6460-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

6555-427: The nest site; it begins shortly after pairing and continues for 3–10 days before egg-laying. The male brings the nesting material, in ground-nesting species (which may not build a nest) sometimes in the pouch, and in tree-nesting species crosswise in the bill. The female then heaps the material up to form a simple structure. The eggs are oval, white, and coarsely textured. All species normally lay at least two eggs;

6650-470: The nest unguarded at times while they go fishing. The times for the chicks to fledge and become independent of their parents depend greatly on the food supply. Rarely does more than one chick survive to maturity, except in the Peruvian booby ( Sula variegata ), which has the biggest clutch (two to four eggs), and less often in the blue-footed booby ( S. nebouxii ). Siblicide by the stronger of two chicks

6745-498: The others by placement in the subgenus Leptopelecanus but in fact species with both sorts of appearance and nesting behavior are found in either. DNA sequencing of both mitochondrial and nuclear genes yielded quite different relationships; the three New World pelicans formed one lineage, with the American white pelican sister to the two brown pelicans, and the five Old World species the other. The Dalmatian, pink-backed, and spot-billed were all closely related to one another, while

6840-407: The outside directly. The eyes are angled forward, and provide a wider field of binocular vision than in most other birds. Their plumage is either all-white (or light brownish or greyish) with dark wingtips and (usually) tail, or at least some dark brown or black above with white underparts; gannets have a yellowish hue to their heads. The face usually has some sort of black markings, typically on

6935-625: The pelican lineage has existed for at least 36 million years; the oldest known pelican fossil was assigned to Eopelecanus aegyptiacus and was found in late Eocene (middle to late part of the early Priabonian stage/age) deposits of the Birket Qarun Formation within the Wadi Al-Hitan World Heritage Site in Egypt. A more complete fossil pelican of early Oligocene age is known from deposits at

7030-429: The pelican's head, peck it to distraction, and grab a fish from the open bill. Pelicans in their turn sometimes snatch prey from other waterbirds. The brown pelican usually plunge-dives head-first for its prey, from a height as great as 10–20 m (33–66 ft), especially for anchovies and menhaden . The only other pelican to feed using a similar technique is the Peruvian pelican, but its dives are typically from

7125-585: The population is estimated by BirdLife International to exceed 500,000 mature individuals, and is possibly increasing, it has been much higher in the past. It declined dramatically during the 1998 El Niño event and could experience similar declines in the future. Conservation needs include regular monitoring throughout the range to determine population trends, particularly after El Niño years, restricting human access to important breeding colonies, and assessing interactions with fisheries. The spot-billed pelican has an estimated population between 13,000 and 18,000 and

7220-629: The presence of pollutants such as DDT and endrin . Most species' populations are more or less stable, although three are classified by the IUCN as being at risk. All species breed readily in zoos, which is potentially useful for conservation management. The combined population of brown and Peruvian pelicans is estimated at 650,000 birds, with around 250,000 in the United States and Caribbean, and 400,000 in Peru. The National Audubon Society estimates

7315-503: The proposed order name. Within the family itself, three living genera— Sula (boobies, six species), Papasula (Abbott's booby), and Morus (gannets, three species)—are recognized. A 2011 study of multiple genes found Abbott's booby to be basal to all other gannets and boobies, and likely to have diverged from them around 22 million years ago, and the ancestors of the gannets and remaining boobies split around 17 million years ago. The most recent common ancestor of all boobies lived in

7410-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

7505-475: The range of interspecific variation in Pelecanus . The Late Eocene Protopelicanus may be a pelecaniform or suliform – or a similar aquatic bird such as a pseudotooth ( Pelagornithidae ), but is not generally considered a pelecanid . The supposed Miocene pelican Liptornis from Patagonia is a nomen dubium (of doubtful validity), being based on fragments providing insufficient evidence to support

7600-731: The reason is not clearly known, but a common belief is that it is to draw attention to itself and away from any siblings who are waiting to be fed. Parents of ground-nesting species sometimes drag older young around roughly by the head before feeding them. From about 25 days old, the young of these species gather in "pods" or " crèches " of up to 100 birds in which parents recognise and feed only their own offspring. By 6–8 weeks they wander around, occasionally swimming, and may practise communal feeding. Young of all species fledge 10–12 weeks after hatching. They may remain with their parents afterwards, but are now seldom or never fed. They are mature at three or four years old. Overall breeding success

7695-417: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or

7790-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in

7885-1487: The shoebill and the hamerkop form a sister group to the pelicans, though some doubt exists as to the exact relationships among the three lineages. The oldest known record of Pelicans is a right tibiotarsus very similar to those of modern species from the Birket Qarun Formation in the Wadi El Hitan in Egypt, dating to the late Eocene ( Priabonian ), referred to the genus Eopelecanus . P. rufescens P. philippensis P. crispus P. conspicillatus P. onocrotalus P. occidentalis P. thagus P. erythrorhynchos Suliformes Herons (Ardeidae) Ibises and spoonbills (Threskiornithidae) Hamerkop ( Scopus umbretta ) Shoebill ( Balaeniceps rex ) Pelicans ( Pelecanus ) Cladogram based on Hackett et al. (2008). The eight living pelican species were traditionally divided into two groups, one containing four ground-nesters with mainly white adult plumage ( Australian , Dalmatian , great white , and American white pelicans ), and one containing four grey- or brown-plumaged species which nest preferentially either in trees ( pink-backed , spot-billed and brown pelicans ), or on sea rocks ( Peruvian pelican ). The largely marine brown and Peruvian pelicans, formerly considered conspecific , are sometimes separated from

7980-484: The skeleton and beneath the skin, enabling them to float high in the water. The tail is short and square. The wings are long and broad, suitably shaped for soaring and gliding flight, and have the unusually large number of 30 to 35 secondary flight feathers . Males are generally larger than females and have longer bills. The smallest species is the brown pelican, small individuals of which can be no more than 2.75 kg (6.1 lb) and 1.06 m (3.5 ft) long, with

8075-564: The skin of the throat and pouch with the bill open to promote evaporative cooling . They roost and loaf communally on beaches, sandbanks, and in shallow water. A fibrous layer deep in the breast muscles can hold the wings rigidly horizontal for gliding and soaring. Thus, they use thermals for soaring to heights of 3000 m (10,000 ft) or more, combined both with gliding and with flapping flight in V formation , to commute distances up to 150 km (93 mi) to feeding areas. Pelicans also fly low (or "skim") over stretches of water, using

8170-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

8265-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

8360-503: The sulids seem to have diverged from the lineage leading to cormorants and darters around 50 million years ago (Mya), perhaps a bit earlier. The initial evolutionary radiation formed a number of genera that are now completely extinct, such as the freshwater Masillastega (which, as noted above, might not have been a modern-type sulid) or the bizarre Rhamphastosula (which had a bill shaped like an aracari 's). The modern genera evolved (like many other living genera of birds) around

8455-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

8550-510: The usual clutch size is one to three, rarely up to six. Both sexes incubate with the eggs on top of or below the feet; they may display when changing shifts. Incubation takes 30–36 days; hatching success for undisturbed pairs can be as high as 95%, but because of sibling competition or siblicide , in the wild, usually all but one nestling dies within the first few weeks (later in the pink-backed and spot-billed species). Both parents feed their young. Small chicks are fed by regurgitation ; after about

8645-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

8740-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

8835-738: The water and catching them at/near the water's surface. They can adapt to varying degrees of water salinity , from freshwater and brackish to—most commonly—seawater. They are gregarious birds, travelling in flocks, hunting cooperatively , and breeding colonially . Four white-plumaged species tend to nest on the ground, and four brown or grey-plumaged species nest mainly in trees. The relationship between pelicans and people has often been contentious. The birds have been persecuted because of their perceived competition with commercial and recreational fishing. Their populations have fallen through habitat destruction , disturbance, and environmental pollution, and three species are of conservation concern. They also have

8930-407: The web connecting all four toes. In some species, the webs are brightly colored and used in courtship displays . The bill is usually conspicuously colored, long, deep at the base, and pointed, with saw-like edges. The upper mandible curves down slightly at the tip and can be moved upward to accept large prey. To keep water out during plunges, the nostrils enter into the bill rather than opening to

9025-538: The well-studied contemporary American deposits. Prehistoric sulids (or suloids) only known from fossils are: For prehistoric species of the extant genera , see the genus articles. The Early Oligocene Prophalacrocorax ronzoni of Ronzon, France , was variously placed in the seaduck genus Mergus , in Sula , and after a distinct genus was established for it, in the Phalacrocoracidae. While it

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