126-647: The Proto-Tethys or Theic Ocean was an ancient ocean that existed from the latest Ediacaran to the Carboniferous (550–330 Ma ). The name "Proto-Tethys" has been used inconsistently for several concepts for a supposed predecessor of the Paleo-Tethys Ocean , a palaeocean that separated the margins of Gondwana , often referred to as peri-Gondwana, from various continents and Gondwana-derived continental fragments from Precambrian times and onwards. According to von Raumer & Stampfli 2008 , after
252-410: A halocline . If a zone undergoes a strong, vertical chemistry gradient with depth, it contains a chemocline . Temperature and salinity control ocean water density. Colder and saltier water is denser, and this density plays a crucial role in regulating the global water circulation within the ocean. The halocline often coincides with the thermocline, and the combination produces a pronounced pycnocline ,
378-493: A mid-ocean ridge , which creates a long mountain range beneath the ocean. Together they form the global mid-oceanic ridge system that features the longest mountain range in the world. The longest continuous mountain range is 65,000 km (40,000 mi). This underwater mountain range is several times longer than the longest continental mountain range – the Andes . Oceanographers state that less than 20% of
504-503: A biochemical pump that collects carbon from the organ interior (or from the soil ) and not from the atmosphere. Cyanobacteria possess carboxysomes , which increase the concentration of CO 2 around RuBisCO to increase the rate of photosynthesis. An enzyme, carbonic anhydrase , located within the carboxysome, releases CO 2 from dissolved hydrocarbonate ions (HCO 3 ). Before the CO 2 can diffuse out, RuBisCO concentrated within
630-496: A boundary between less dense surface water and dense deep water. Photosynthesis Photosynthesis ( / ˌ f oʊ t ə ˈ s ɪ n θ ə s ɪ s / FOH -tə- SINTH -ə-sis ) is a system of biological processes by which photosynthetic organisms , such as most plants, algae , and cyanobacteria , convert light energy , typically from sunlight, into the chemical energy necessary to fuel their metabolism . Photosynthesis usually refers to oxygenic photosynthesis ,
756-422: A depth of a few hundred meters or less. Human activity often has negative impacts on marine life within the continental shelf. Ocean temperatures depend on the amount of solar radiation reaching the ocean surface. In the tropics, surface temperatures can rise to over 30 °C (86 °F). Near the poles where sea ice forms, the temperature in equilibrium is about −2 °C (28 °F). In all parts of
882-409: A different leaf anatomy from C 3 plants, and fix the CO 2 at night, when their stomata are open. CAM plants store the CO 2 mostly in the form of malic acid via carboxylation of phosphoenolpyruvate to oxaloacetate , which is then reduced to malate. Decarboxylation of malate during the day releases CO 2 inside the leaves, thus allowing carbon fixation to 3-phosphoglycerate by RuBisCO. CAM
1008-595: A gentle breeze on a pond causes ripples to form. A stronger gust blowing over the ocean causes larger waves as the moving air pushes against the raised ridges of water. The waves reach their maximum height when the rate at which they are travelling nearly matches the speed of the wind. In open water, when the wind blows continuously as happens in the Southern Hemisphere in the Roaring Forties , long, organized masses of water called swell roll across
1134-480: A phospholipid outer membrane, and an intermembrane space. Enclosed by the membrane is an aqueous fluid called the stroma. Embedded within the stroma are stacks of thylakoids (grana), which are the site of photosynthesis. The thylakoids appear as flattened disks. The thylakoid itself is enclosed by the thylakoid membrane, and within the enclosed volume is a lumen or thylakoid space. Embedded in the thylakoid membrane are integral and peripheral membrane protein complexes of
1260-479: A photocomplex. When a photon is absorbed by a chromophore, it is converted into a quasiparticle referred to as an exciton , which jumps from chromophore to chromophore towards the reaction center of the photocomplex, a collection of molecules that traps its energy in a chemical form accessible to the cell's metabolism. The exciton's wave properties enable it to cover a wider area and try out several possible paths simultaneously, allowing it to instantaneously "choose"
1386-405: A photon by the antenna complex loosens an electron by a process called photoinduced charge separation . The antenna system is at the core of the chlorophyll molecule of the photosystem II reaction center. That loosened electron is taken up by the primary electron-acceptor molecule, pheophytin . As the electrons are shuttled through an electron transport chain (the so-called Z-scheme shown in
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#17327648004871512-422: A plant's photosynthetic response. Integrated chlorophyll fluorometer – gas exchange systems allow a more precise measure of photosynthetic response and mechanisms. While standard gas exchange photosynthesis systems can measure Ci, or substomatal CO 2 levels, the addition of integrated chlorophyll fluorescence measurements allows a more precise measurement of C C, the estimation of CO 2 concentration at
1638-405: A process that produces oxygen. Photosynthetic organisms store the chemical energy so produced within intracellular organic compounds (compounds containing carbon) like sugars, glycogen , cellulose and starches . To use this stored chemical energy, an organism's cells metabolize the organic compounds through cellular respiration . Photosynthesis plays a critical role in producing and maintaining
1764-450: A redox-active tyrosine residue that is oxidized by the energy of P680 . This resets the ability of P680 to absorb another photon and release another photo-dissociated electron. The oxidation of water is catalyzed in photosystem II by a redox-active structure that contains four manganese ions and a calcium ion ; this oxygen-evolving complex binds two water molecules and contains the four oxidizing equivalents that are used to drive
1890-610: A result of the Chamrousse back-arc basin . Torsvik & Cocks 2009 used the name 'Ran Ocean' for a similar concept, the Cambrian-Ordovician ocean that separated Baltica from Gondwana. Other geologists dispute the existence of such an ocean. The ocean formed when Pannotia disintegrated, Proto-Laurasia (Laurentia, Baltica, and Siberia) rifted away from a supercontinent that would become Gondwana . Proto-Tethys formed between these two supercontinents. The ocean
2016-496: A shallow area and this, coupled with a low pressure system, can raise the surface of the ocean dramatically above a typical high tide. The average depth of the oceans is about 4 km. More precisely the average depth is 3,688 meters (12,100 ft). Nearly half of the world's marine waters are over 3,000 meters (9,800 ft) deep. "Deep ocean," which is anything below 200 meters (660 ft), covers about 66% of Earth's surface. This figure does not include seas not connected to
2142-399: A simpler method that employs a pigment similar to those used for vision in animals. The bacteriorhodopsin changes its configuration in response to sunlight, acting as a proton pump. This produces a proton gradient more directly, which is then converted to chemical energy. The process does not involve carbon dioxide fixation and does not release oxygen, and seems to have evolved separately from
2268-481: A source of carbon atoms to carry out photosynthesis; photoheterotrophs use organic compounds, rather than carbon dioxide, as a source of carbon. In plants, algae, and cyanobacteria, photosynthesis releases oxygen. This oxygenic photosynthesis is by far the most common type of photosynthesis used by living organisms. Some shade-loving plants (sciophytes) produce such low levels of oxygen during photosynthesis that they use all of it themselves instead of releasing it to
2394-541: A subsequent sequence of light-independent reactions called the Calvin cycle . In this process, atmospheric carbon dioxide is incorporated into already existing organic compounds, such as ribulose bisphosphate (RuBP). Using the ATP and NADPH produced by the light-dependent reactions, the resulting compounds are then reduced and removed to form further carbohydrates, such as glucose . In other bacteria, different mechanisms like
2520-403: A wave-cut platform develops at the foot of the cliff and this has a protective effect, reducing further wave-erosion. Material worn from the margins of the land eventually ends up in the sea. Here it is subject to attrition as currents flowing parallel to the coast scour out channels and transport sand and pebbles away from their place of origin. Sediment carried to the sea by rivers settles on
2646-591: A wide variety of colors. These pigments are embedded in plants and algae in complexes called antenna proteins. In such proteins, the pigments are arranged to work together. Such a combination of proteins is also called a light-harvesting complex . Although all cells in the green parts of a plant have chloroplasts, the majority of those are found in specially adapted structures called leaves . Certain species adapted to conditions of strong sunlight and aridity , such as many Euphorbia and cactus species, have their main photosynthetic organs in their stems. The cells in
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#17327648004872772-406: A zone undergoes dramatic changes in temperature with depth, it contains a thermocline , a distinct boundary between warmer surface water and colder deep water. In tropical regions, the thermocline is typically deeper compared to higher latitudes. Unlike polar waters , where solar energy input is limited, temperature stratification is less pronounced, and a distinct thermocline is often absent. This
2898-439: Is a bay , a small bay with a narrow inlet is a cove and a large bay may be referred to as a gulf . Coastlines are influenced by several factors including the strength of the waves arriving on the shore, the gradient of the land margin, the composition and hardness of the coastal rock, the inclination of the off-shore slope and the changes of the level of the land due to local uplift or submergence. Normally, waves roll towards
3024-490: Is an endothermic redox reaction. In general outline, photosynthesis is the opposite of cellular respiration : while photosynthesis is a process of reduction of carbon dioxide to carbohydrates, cellular respiration is the oxidation of carbohydrates or other nutrients to carbon dioxide. Nutrients used in cellular respiration include carbohydrates, amino acids and fatty acids. These nutrients are oxidized to produce carbon dioxide and water, and to release chemical energy to drive
3150-444: Is an important reference point for oceanography and geography, particularly as mean sea level . The ocean surface has globally little, but measurable topography , depending on the ocean's volumes. The ocean surface is a crucial interface for oceanic and atmospheric processes. Allowing interchange of particles, enriching the air and water, as well as grounds by some particles becoming sediments . This interchange has fertilized life in
3276-502: Is both an evolutionary precursor to C 4 and a useful carbon-concentrating mechanism in its own right. Xerophytes , such as cacti and most succulents , also use PEP carboxylase to capture carbon dioxide in a process called Crassulacean acid metabolism (CAM). In contrast to C 4 metabolism, which spatially separates the CO 2 fixation to PEP from the Calvin cycle, CAM temporally separates these two processes. CAM plants have
3402-474: Is commonly measured in μmols /( m / s ), parts per million, or volume per million; and H 2 O is commonly measured in mmols /(m /s) or in mbars . By measuring CO 2 assimilation , ΔH 2 O, leaf temperature, barometric pressure , leaf area, and photosynthetically active radiation (PAR), it becomes possible to estimate, "A" or carbon assimilation, "E" or transpiration , "gs" or stomatal conductance , and "Ci" or intracellular CO 2 . However, it
3528-430: Is converted to CO 2 by an oxalate oxidase enzyme, and the produced CO 2 can support the Calvin cycle reactions. Reactive hydrogen peroxide (H 2 O 2 ), the byproduct of oxalate oxidase reaction, can be neutralized by catalase . Alarm photosynthesis represents a photosynthetic variant to be added to the well-known C4 and CAM pathways. However, alarm photosynthesis, in contrast to these pathways, operates as
3654-411: Is customarily divided into five principal oceans – listed below in descending order of area and volume: The ocean fills Earth's oceanic basins . Earth's oceanic basins cover different geologic provinces of Earth's oceanic crust as well as continental crust . As such it covers mainly Earth's structural basins , but also continental shelfs . In mid-ocean, magma is constantly being thrust through
3780-399: Is due to the fact that surface waters in polar latitudes are nearly as cold as deeper waters. Below the thermocline, water everywhere in the ocean is very cold, ranging from −1 °C to 3 °C. Because this deep and cold layer contains the bulk of ocean water, the average temperature of the world ocean is 3.9 °C. If a zone undergoes dramatic changes in salinity with depth, it contains
3906-419: Is freed from its locked position through a classic "hop". The movement of the electron towards the photo center is therefore covered in a series of conventional hops and quantum walks. Fossils of what are thought to be filamentous photosynthetic organisms have been dated at 3.4 billion years old. More recent studies also suggest that photosynthesis may have begun about 3.4 billion years ago, though
Proto-Tethys Ocean - Misplaced Pages Continue
4032-412: Is further excited by the light absorbed by that photosystem . The electron is then passed along a chain of electron acceptors to which it transfers some of its energy . The energy delivered to the electron acceptors is used to move hydrogen ions across the thylakoid membrane into the lumen . The electron is eventually used to reduce the coenzyme NADP with an H to NADPH (which has functions in
4158-508: Is irregular, unevenly dominating the Earth's surface . This leads to the distinction of the Earth's surface into a water and land hemisphere , as well as the division of the ocean into different oceans. Seawater covers about 361,000,000 km (139,000,000 sq mi) and the ocean's furthest pole of inaccessibility , known as " Point Nemo ", in a region known as spacecraft cemetery of
4284-451: Is more common to use chlorophyll fluorescence for plant stress measurement , where appropriate, because the most commonly used parameters FV/FM and Y(II) or F/FM' can be measured in a few seconds, allowing the investigation of larger plant populations. Gas exchange systems that offer control of CO 2 levels, above and below ambient , allow the common practice of measurement of A/Ci curves, at different CO 2 levels, to characterize
4410-406: Is passed through a food chain . The fixation or reduction of carbon dioxide is a process in which carbon dioxide combines with a five-carbon sugar , ribulose 1,5-bisphosphate , to yield two molecules of a three-carbon compound, glycerate 3-phosphate , also known as 3-phosphoglycerate. Glycerate 3-phosphate, in the presence of ATP and NADPH produced during the light-dependent stages,
4536-454: Is produced and magma is forced up creating underwater mountains, some of which may form chains of volcanic islands near to deep trenches. Near some of the boundaries between the land and sea, the slightly denser oceanic plates slide beneath the continental plates and more subduction trenches are formed. As they grate together, the continental plates are deformed and buckle causing mountain building and seismic activity. Every ocean basin has
4662-407: Is pushed across the surface of the ocean by the wind, but this represents a transfer of energy and not horizontal movement of water. As waves approach land and move into shallow water , they change their behavior. If approaching at an angle, waves may bend ( refraction ) or wrap around rocks and headlands ( diffraction ). When the wave reaches a point where its deepest oscillations of the water contact
4788-623: Is reduced to glyceraldehyde 3-phosphate . This product is also referred to as 3-phosphoglyceraldehyde (PGAL) or, more generically, as triose phosphate. Most (five out of six molecules) of the glyceraldehyde 3-phosphate produced are used to regenerate ribulose 1,5-bisphosphate so the process can continue. The triose phosphates not thus "recycled" often condense to form hexose phosphates, which ultimately yield sucrose , starch , and cellulose , as well as glucose and fructose . The sugars produced during carbon metabolism yield carbon skeletons that can be used for other metabolic reactions like
4914-423: Is reflected back out of the water. Red light is most easily absorbed and thus does not reach great depths, usually to less than 50 meters (164 ft). Blue light, in comparison, can penetrate up to 200 meters (656 ft). Second, water molecules and very tiny particles in ocean water preferentially scatter blue light more than light of other colors. Blue light scattering by water and tiny particles happens even in
5040-480: Is sometimes referred to as the World Ocean, global ocean or the great ocean . The concept of a continuous body of water with relatively unrestricted exchange between its components is critical in oceanography . The word ocean comes from the figure in classical antiquity , Oceanus ( / oʊ ˈ s iː ə n ə s / ; ‹See Tfd› Greek : Ὠκεανός Ōkeanós , pronounced [ɔːkeanós] ),
5166-429: Is then translocated to specialized bundle sheath cells where the enzyme RuBisCO and other Calvin cycle enzymes are located, and where CO 2 released by decarboxylation of the four-carbon acids is then fixed by RuBisCO activity to the three-carbon 3-phosphoglyceric acids . The physical separation of RuBisCO from the oxygen-generating light reactions reduces photorespiration and increases CO 2 fixation and, thus,
Proto-Tethys Ocean - Misplaced Pages Continue
5292-404: Is then converted into the final carbohydrate products. The simple carbon sugars photosynthesis produces are then used to form other organic compounds , such as the building material cellulose , the precursors for lipid and amino acid biosynthesis, or as a fuel in cellular respiration . The latter occurs not only in plants but also in animals when the carbon and energy from plants
5418-499: Is used by 16,000 species of plants. Calcium-oxalate -accumulating plants, such as Amaranthus hybridus and Colobanthus quitensis , show a variation of photosynthesis where calcium oxalate crystals function as dynamic carbon pools , supplying carbon dioxide (CO 2 ) to photosynthetic cells when stomata are partially or totally closed. This process was named alarm photosynthesis . Under stress conditions (e.g., water deficit ), oxalate released from calcium oxalate crystals
5544-474: Is vital for climate processes, as it captures carbon dioxide from the air and binds it into plants, harvested produce and soil. Cereals alone are estimated to bind 3,825 Tg or 3.825 Pg of carbon dioxide every year, i.e. 3.825 billion metric tons. Most photosynthetic organisms are photoautotrophs , which means that they are able to synthesize food directly from carbon dioxide and water using energy from light. However, not all organisms use carbon dioxide as
5670-786: The Cadomian orogenic events 550 Ma, the Proto-Tethys Ocean formed the eastern part of an oceanic domain (of which the Iapetus and Tornquist oceans formed the western parts) which subducted under the northern margins of Gondwana . In this model the Proto-Tethys separated the North China and Baltica continents from Gondwana. In the Early Ordovician 500-480 Ma, the Proto-Tethys was subducted under Cadomia as
5796-691: The North Sea or the Red Sea . There is no sharp distinction between seas and oceans, though generally seas are smaller, and are often partly (as marginal seas ) or wholly (as inland seas ) bordered by land. The contemporary concept of the World Ocean was coined in the early 20th century by the Russian oceanographer Yuly Shokalsky to refer to the continuous ocean that covers and encircles most of Earth. The global, interconnected body of salt water
5922-481: The South Pacific Ocean , at 48°52.6′S 123°23.6′W / 48.8767°S 123.3933°W / -48.8767; -123.3933 ( Point Nemo ) . This point is roughly 2,688 km (1,670 mi) from the nearest land. There are different customs to subdivide the ocean and are adjourned by smaller bodies of water such as, seas , gulfs , bays , bights , and straits . The ocean
6048-592: The Thames Barrier is designed to protect London from a storm surge, while the failure of the dykes and levees around New Orleans during Hurricane Katrina created a humanitarian crisis in the United States. Most of the ocean is blue in color, but in some places the ocean is blue-green, green, or even yellow to brown. Blue ocean color is a result of several factors. First, water preferentially absorbs red light, which means that blue light remains and
6174-410: The carbon cycle , and the water cycle by acting as a huge heat reservoir . Ocean scientists split the ocean into vertical and horizontal zones based on physical and biological conditions. The pelagic zone is the open ocean's water column from the surface to the ocean floor. The water column is further divided into zones based on depth and the amount of light present. The photic zone starts at
6300-406: The coastline and structure of the world ocean. A global ocean has existed in one form or another on Earth for eons. Since its formation the ocean has taken many conditions and shapes with many past ocean divisions and potentially at times covering the whole globe. During colder climatic periods, more ice caps and glaciers form, and enough of the global water supply accumulates as ice to lessen
6426-637: The light reaction of photosynthesis by using chlorophyll fluorometers . Actual plants' photosynthetic efficiency varies with the frequency of the light being converted, light intensity , temperature , and proportion of carbon dioxide in the atmosphere , and can vary from 0.1% to 8%. By comparison, solar panels convert light into electric energy at an efficiency of approximately 6–20% for mass-produced panels, and above 40% in laboratory devices. Scientists are studying photosynthesis in hopes of developing plants with increased yield . The efficiency of both light and dark reactions can be measured, but
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#17327648004876552-434: The light-independent (or "dark") reactions, the enzyme RuBisCO captures CO 2 from the atmosphere and, in a process called the Calvin cycle , uses the newly formed NADPH and releases three-carbon sugars , which are later combined to form sucrose and starch . The overall equation for the light-independent reactions in green plants is Carbon fixation produces the three-carbon sugar intermediate , which
6678-591: The ocean floor , they begin to slow down. This pulls the crests closer together and increases the waves' height , which is called wave shoaling . When the ratio of the wave's height to the water depth increases above a certain limit, it " breaks ", toppling over in a mass of foaming water. This rushes in a sheet up the beach before retreating into the ocean under the influence of gravity. Earthquakes , volcanic eruptions or other major geological disturbances can set off waves that can lead to tsunamis in coastal areas which can be very dangerous. The ocean's surface
6804-441: The oxygen content of the Earth's atmosphere, and it supplies most of the biological energy necessary for complex life on Earth. Some bacteria also perform anoxygenic photosynthesis , which uses bacteriochlorophyll to split hydrogen sulfide as a reductant instead of water, producing sulfur instead of oxygen. Archaea such as Halobacterium also perform a type of non- carbon-fixing anoxygenic photosynthesis, where
6930-417: The palisade mesophyll cells where most of the photosynthesis takes place. In the light-dependent reactions , one molecule of the pigment chlorophyll absorbs one photon and loses one electron . This electron is taken up by a modified form of chlorophyll called pheophytin , which passes the electron to a quinone molecule, starting the flow of electrons down an electron transport chain that leads to
7056-417: The photosynthetic capacity of the leaf . C 4 plants can produce more sugar than C 3 plants in conditions of high light and temperature . Many important crop plants are C 4 plants, including maize , sorghum , sugarcane , and millet . Plants that do not use PEP-carboxylase in carbon fixation are called C 3 plants because the primary carboxylation reaction , catalyzed by RuBisCO, produces
7182-462: The photosystems , quantum efficiency and the CO 2 assimilation rates. With some instruments, even wavelength dependency of the photosynthetic efficiency can be analyzed . A phenomenon known as quantum walk increases the efficiency of the energy transport of light significantly. In the photosynthetic cell of an alga , bacterium , or plant, there are light-sensitive molecules called chromophores arranged in an antenna-shaped structure called
7308-437: The plasma membrane . In these light-dependent reactions, some energy is used to strip electrons from suitable substances, such as water, producing oxygen gas. The hydrogen freed by the splitting of water is used in the creation of two important molecules that participate in energetic processes: reduced nicotinamide adenine dinucleotide phosphate (NADPH) and ATP. In plants, algae, and cyanobacteria, sugars are synthesized by
7434-402: The reverse Krebs cycle are used to achieve the same end. The first photosynthetic organisms probably evolved early in the evolutionary history of life using reducing agents such as hydrogen or hydrogen sulfide, rather than water, as sources of electrons. Cyanobacteria appeared later; the excess oxygen they produced contributed directly to the oxygenation of the Earth , which rendered
7560-433: The CO 2 concentration in the leaves under these conditions. Plants that use the C 4 carbon fixation process chemically fix carbon dioxide in the cells of the mesophyll by adding it to the three-carbon molecule phosphoenolpyruvate (PEP), a reaction catalyzed by an enzyme called PEP carboxylase , creating the four-carbon organic acid oxaloacetic acid . Oxaloacetic acid or malate synthesized by this process
7686-442: The Earth's biosphere . Oceanic evaporation , as a phase of the water cycle, is the source of most rainfall (about 90%), causing a global cloud cover of 67% and a consistent oceanic cloud cover of 72%. Ocean temperatures affect climate and wind patterns that affect life on land. One of the most dramatic forms of weather occurs over the oceans: tropical cyclones (also called "typhoons" and "hurricanes" depending upon where
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#17327648004877812-456: The Moon are 20x stronger than the Moon's tidal forces on the Earth.) The primary effect of lunar tidal forces is to bulge Earth matter towards the near and far sides of the Earth, relative to the moon. The "perpendicular" sides, from which the Moon appears in line with the local horizon, experience "tidal troughs". Since it takes nearly 25 hours for the Earth to rotate under the Moon (accounting for
7938-403: The Moon's 28 day orbit around Earth), tides thus cycle over a course of 12.5 hours. However, the rocky continents pose obstacles for the tidal bulges, so the timing of tidal maxima may not actually align with the Moon in most localities on Earth, as the oceans are forced to "dodge" the continents. Timing and magnitude of tides vary widely across the Earth as a result of the continents. Thus, knowing
8064-471: The Moon's gravity, oceanic tides are also substantially modulated by the Sun's tidal forces, by the rotation of the Earth, and by the shape of the rocky continents blocking oceanic water flow. (Tidal forces vary more with distance than the "base" force of gravity: the Moon's tidal forces on Earth are more than double the Sun's, despite the latter's much stronger gravitational force on Earth. Earth's tidal forces upon
8190-400: The Moon's position does not allow a local to predict tide timings, instead requiring precomputed tide tables which account for the continents and the Sun, among others. During each tidal cycle, at any given place the tidal waters rise to maximum height, high tide, before ebbing away again to the minimum level, low tide. As the water recedes, it gradually reveals the foreshore , also known as
8316-648: The Okeanos is represented with a dragon-tail on some early Greek vases. Scientists believe that a sizable quantity of water would have been in the material that formed Earth. Water molecules would have escaped Earth's gravity more easily when it was less massive during its formation. This is called atmospheric escape . During planetary formation , Earth possibly had magma oceans . Subsequently, outgassing , volcanic activity and meteorite impacts , produced an early atmosphere of carbon dioxide , nitrogen and water vapor , according to current theories. The gases and
8442-741: The World Ocean, such as the Caspian Sea . The deepest region of the ocean is at the Mariana Trench , located in the Pacific Ocean near the Northern Mariana Islands . The maximum depth has been estimated to be 10,971 meters (35,994 ft). The British naval vessel Challenger II surveyed the trench in 1951 and named the deepest part of the trench the " Challenger Deep ". In 1960, the Trieste successfully reached
8568-419: The action spectrum is blue-green light, which allows these algae to use the blue end of the spectrum to grow in the deeper waters that filter out the longer wavelengths (red light) used by above-ground green plants. The non-absorbed part of the light spectrum is what gives photosynthetic organisms their color (e.g., green plants, red algae, purple bacteria ) and is the least effective for photosynthesis in
8694-421: The amounts in other parts of the water cycle. The reverse is true during warm periods. During the last ice age, glaciers covered almost one-third of Earth's land mass with the result being that the oceans were about 122 m (400 ft) lower than today. During the last global "warm spell," about 125,000 years ago, the seas were about 5.5 m (18 ft) higher than they are now. About three million years ago
8820-479: The atmosphere are thought to have accumulated over millions of years. After Earth's surface had significantly cooled, the water vapor over time would have condensed, forming Earth's first oceans. The early oceans might have been significantly hotter than today and appeared green due to high iron content. Geological evidence helps constrain the time frame for liquid water existing on Earth. A sample of pillow basalt (a type of rock formed during an underwater eruption)
8946-521: The atmosphere. Although there are some differences between oxygenic photosynthesis in plants , algae , and cyanobacteria , the overall process is quite similar in these organisms. There are also many varieties of anoxygenic photosynthesis , used mostly by bacteria, which consume carbon dioxide but do not release oxygen. Carbon dioxide is converted into sugars in a process called carbon fixation ; photosynthesis captures energy from sunlight to convert carbon dioxide into carbohydrates . Carbon fixation
9072-407: The bottom of the trench, manned by a crew of two men. Oceanographers classify the ocean into vertical and horizontal zones based on physical and biological conditions. The pelagic zone consists of the water column of the open ocean, and can be divided into further regions categorized by light abundance and by depth. The ocean zones can be grouped by light penetration into (from top to bottom):
9198-483: The carboxysome quickly sponges it up. HCO 3 ions are made from CO 2 outside the cell by another carbonic anhydrase and are actively pumped into the cell by a membrane protein. They cannot cross the membrane as they are charged, and within the cytosol they turn back into CO 2 very slowly without the help of carbonic anhydrase. This causes the HCO 3 ions to accumulate within the cell from where they diffuse into
9324-491: The carboxysomes. Pyrenoids in algae and hornworts also act to concentrate CO 2 around RuBisCO. The overall process of photosynthesis takes place in four stages: Plants usually convert light into chemical energy with a photosynthetic efficiency of 3–6%. Absorbed light that is unconverted is dissipated primarily as heat , with a small fraction (1–2%) reemitted as chlorophyll fluorescence at longer (redder) wavelengths . This fact allows measurement of
9450-409: The conditions of non-cyclic electron flow in green plants is: Not all wavelengths of light can support photosynthesis. The photosynthetic action spectrum depends on the type of accessory pigments present. For example, in green plants , the action spectrum resembles the absorption spectrum for chlorophylls and carotenoids with absorption peaks in violet-blue and red light. In red algae ,
9576-425: The details may differ between species , the process always begins when light energy is absorbed by the reaction centers , proteins that contain photosynthetic pigments or chromophores . In plants, these proteins are chlorophylls (a porphyrin derivative that absorbs the red and blue spectrums of light, thus reflecting green) held inside chloroplasts , abundant in leaf cells. In bacteria, they are embedded in
9702-514: The diagram), a chemiosmotic potential is generated by pumping proton cations ( H ) across the membrane and into the thylakoid space . An ATP synthase enzyme uses that chemiosmotic potential to make ATP during photophosphorylation , whereas NADPH is a product of the terminal redox reaction in the Z-scheme . The electron enters a chlorophyll molecule in Photosystem I . There it
9828-509: The elder of the Titans in classical Greek mythology . Oceanus was believed by the ancient Greeks and Romans to be the divine personification of an enormous river encircling the world. The concept of Ōkeanós has an Indo-European connection. Greek Ōkeanós has been compared to the Vedic epithet ā-śáyāna-, predicated of the dragon Vṛtra-, who captured the cows/rivers. Related to this notion,
9954-505: The equation for this process is: This equation emphasizes that water is both a reactant in the light-dependent reaction and a product of the light-independent reaction , but canceling n water molecules from each side gives the net equation: Other processes substitute other compounds (such as arsenite ) for water in the electron-supply role; for example some microbes use sunlight to oxidize arsenite to arsenate : The equation for this reaction is: Photosynthesis occurs in two stages. In
10080-518: The evolution of complex life possible. The average rate of energy captured by global photosynthesis is approximately 130 terawatts , which is about eight times the total power consumption of human civilization . Photosynthetic organisms also convert around 100–115 billion tons (91–104 Pg petagrams , or billions of metric tons), of carbon into biomass per year. Photosynthesis was discovered in 1779 by Jan Ingenhousz . He showed that plants need light, not just air, soil, and water. Photosynthesis
10206-582: The first stage, light-dependent reactions or light reactions capture the energy of light and use it to make the hydrogen carrier NADPH and the energy-storage molecule ATP . During the second stage, the light-independent reactions use these products to capture and reduce carbon dioxide. Most organisms that use oxygenic photosynthesis use visible light for the light-dependent reactions, although at least three use shortwave infrared or, more specifically, far-red radiation. Some organisms employ even more radical variants of photosynthesis. Some archaea use
10332-441: The first step of the Z-scheme , requires an external source of electrons to reduce its oxidized chlorophyll a reaction center. The source of electrons for photosynthesis in green plants and cyanobacteria is water. Two water molecules are oxidized by the energy of four successive charge-separation reactions of photosystem II to yield a molecule of diatomic oxygen and four hydrogen ions. The electrons yielded are transferred to
10458-412: The formation of unusually high rogue waves . Most waves are less than 3 m (10 ft) high and it is not unusual for strong storms to double or triple that height. Rogue waves, however, have been documented at heights above 25 meters (82 ft). The top of a wave is known as the crest, the lowest point between waves is the trough and the distance between the crests is the wavelength. The wave
10584-421: The interface between water and air is called swell – a term used in sailing , surfing and navigation . These motions profoundly affect ships on the surface of the ocean and the well-being of people on those ships who might suffer from sea sickness . Wind blowing over the surface of a body of water forms waves that are perpendicular to the direction of the wind. The friction between air and water caused by
10710-399: The interior of a cell, giving the membrane a very large surface area and therefore increasing the amount of light that the bacteria can absorb. In plants and algae, photosynthesis takes place in organelles called chloroplasts . A typical plant cell contains about 10 to 100 chloroplasts. The chloroplast is enclosed by a membrane. This membrane is composed of a phospholipid inner membrane,
10836-433: The interior tissues of a leaf, called the mesophyll , can contain between 450,000 and 800,000 chloroplasts for every square millimeter of leaf. The surface of the leaf is coated with a water-resistant waxy cuticle that protects the leaf from excessive evaporation of water and decreases the absorption of ultraviolet or blue light to minimize heating . The transparent epidermis layer allows light to pass through to
10962-482: The intertidal zone. The difference in height between the high tide and low tide is known as the tidal range or tidal amplitude. When the sun and moon are aligned (full moon or new moon), the combined effect results in the higher "spring tides", while the sun and moon misaligning (half moons) result in lesser tidal ranges. In the open ocean tidal ranges are less than 1 meter, but in coastal areas these tidal ranges increase to more than 10 meters in some areas. Some of
11088-412: The large bodies of water into which the world ocean is conventionally divided. The following names describe five different areas of the ocean: Pacific , Atlantic , Indian , Antarctic/Southern , and Arctic . The ocean contains 97% of Earth's water and is the primary component of Earth's hydrosphere and is thereby essential to life on Earth. The ocean influences climate and weather patterns,
11214-739: The largest tidal ranges in the world occur in the Bay of Fundy and Ungava Bay in Canada, reaching up to 16 meters. Other locations with record high tidal ranges include the Bristol Channel between England and Wales, Cook Inlet in Alaska, and the Río Gallegos in Argentina. Tides are not to be confused with storm surges , which can occur when high winds pile water up against the coast in
11340-551: The late Devonian , the microcontinent of Kazakhstania collided with Siberia, shrinking the ocean even more. The ocean closed when the North China craton collided with Siberia-Kazakstania continent in the Carboniferous , while the Paleo-Tethys Ocean expanded. Ocean The ocean is the body of salt water that covers approximately 70.8% of Earth . In English , the term ocean also refers to any of
11466-434: The light reaction, and infrared gas analyzers can measure the dark reaction . An integrated chlorophyll fluorometer and gas exchange system can investigate both light and dark reactions when researchers use the two separate systems together. Infrared gas analyzers and some moisture sensors are sensitive enough to measure the photosynthetic assimilation of CO 2 and of Δ H 2 O using reliable methods . CO 2
11592-433: The light-independent reaction); at that point, the path of that electron ends. The cyclic reaction is similar to that of the non-cyclic but differs in that it generates only ATP, and no reduced NADP (NADPH) is created. The cyclic reaction takes place only at photosystem I. Once the electron is displaced from the photosystem, the electron is passed down the electron acceptor molecules and returns to photosystem I, from where it
11718-471: The majority of Earth's surface. It includes the Pacific , Atlantic , Indian , Southern/Antarctic , and Arctic oceans. As a general term, "the ocean" and "the sea" are often interchangeable. Strictly speaking, a "sea" is a body of water (generally a division of the world ocean) partly or fully enclosed by land. The word "sea" can also be used for many specific, much smaller bodies of seawater, such as
11844-432: The more common types of photosynthesis. In photosynthetic bacteria, the proteins that gather light for photosynthesis are embedded in cell membranes . In its simplest form, this involves the membrane surrounding the cell itself. However, the membrane may be tightly folded into cylindrical sheets called thylakoids , or bunched up into round vesicles called intracytoplasmic membranes . These structures can fill most of
11970-439: The most efficient route, where it will have the highest probability of arriving at its destination in the minimum possible time. Because that quantum walking takes place at temperatures far higher than quantum phenomena usually occur, it is only possible over very short distances. Obstacles in the form of destructive interference cause the particle to lose its wave properties for an instant before it regains them once again after it
12096-408: The ocean faces many environmental threats, such as marine pollution , overfishing , and the effects of climate change . Those effects include ocean warming , ocean acidification and sea level rise . The continental shelf and coastal waters are most affected by human activity. The terms "the ocean" or "the sea" used without specification refer to the interconnected body of salt water covering
12222-693: The ocean, deep ocean temperatures range between −2 °C (28 °F) and 5 °C (41 °F). Constant circulation of water in the ocean creates ocean currents . Those currents are caused by forces operating on the water, such as temperature and salinity differences, atmospheric circulation (wind), and the Coriolis effect . Tides create tidal currents, while wind and waves cause surface currents. The Gulf Stream , Kuroshio Current , Agulhas Current and Antarctic Circumpolar Current are all major ocean currents. Such currents transport massive amounts of water, gases, pollutants and heat to different parts of
12348-508: The ocean, on land and air. All these processes and components together make up ocean surface ecosystems . Tides are the regular rise and fall in water level experienced by oceans, primarily driven by the Moon 's gravitational tidal forces upon the Earth. Tidal forces affect all matter on Earth, but only fluids like the ocean demonstrate the effects on human timescales. (For example, tidal forces acting on rock may produce tidal locking between two planetary bodies.) Though primarily driven by
12474-469: The ocean. If the wind dies down, the wave formation is reduced, but already-formed waves continue to travel in their original direction until they meet land. The size of the waves depends on the fetch , the distance that the wind has blown over the water and the strength and duration of that wind. When waves meet others coming from different directions, interference between the two can produce broken, irregular seas. Constructive interference can lead to
12600-419: The oceans absorb CO 2 from the atmosphere , a higher concentration leads to ocean acidification (a drop in pH value ). The ocean provides many benefits to humans such as ecosystem services , access to seafood and other marine resources , and a means of transport . The ocean is known to be the habitat of over 230,000 species , but may hold considerably more – perhaps over two million species. Yet,
12726-425: The oceans could have been up to 50 m (165 ft) higher. The entire ocean, containing 97% of Earth's water, spans 70.8% of Earth 's surface, making it Earth's global ocean or world ocean . This makes Earth, along with its vibrant hydrosphere a "water world" or " ocean world ", particularly in Earth's early history when the ocean is thought to have possibly covered Earth completely. The ocean's shape
12852-434: The oceans have been mapped. The zone where land meets sea is known as the coast , and the part between the lowest spring tides and the upper limit reached by splashing waves is the shore . A beach is the accumulation of sand or shingle on the shore. A headland is a point of land jutting out into the sea and a larger promontory is known as a cape . The indentation of a coastline, especially between two headlands,
12978-537: The oceans may have always been on the Earth since the beginning of the planet's formation. In this model, atmospheric greenhouse gases kept the oceans from freezing when the newly forming Sun had only 70% of its current luminosity . The origin of Earth's oceans is unknown. Oceans are thought to have formed in the Hadean eon and may have been the cause for the emergence of life . Plate tectonics , post-glacial rebound , and sea level rise continually change
13104-410: The organism's metabolism . Photosynthesis and cellular respiration are distinct processes, as they take place through different sequences of chemical reactions and in different cellular compartments (cellular respiration in mitochondria ). The general equation for photosynthesis as first proposed by Cornelis van Niel is: Since water is used as the electron donor in oxygenic photosynthesis,
13230-451: The photic zone, the mesopelagic zone and the aphotic deep ocean zone: The pelagic part of the aphotic zone can be further divided into vertical regions according to depth and temperature: Distinct boundaries between ocean surface waters and deep waters can be drawn based on the properties of the water. These boundaries are called thermoclines (temperature), haloclines (salinity), chemoclines (chemistry), and pycnoclines (density). If
13356-556: The photosynthetic system. Plants absorb light primarily using the pigment chlorophyll . The green part of the light spectrum is not absorbed but is reflected, which is the reason that most plants have a green color. Besides chlorophyll, plants also use pigments such as carotenes and xanthophylls . Algae also use chlorophyll, but various other pigments are present, such as phycocyanin , carotenes , and xanthophylls in green algae , phycoerythrin in red algae (rhodophytes) and fucoxanthin in brown algae and diatoms resulting in
13482-418: The power of a storm wave impacting on the foot of a cliff has a shattering effect as air in cracks and crevices is compressed and then expands rapidly with release of pressure. At the same time, sand and pebbles have an erosive effect as they are thrown against the rocks. This tends to undercut the cliff, and normal weathering processes such as the action of frost follows, causing further destruction. Gradually,
13608-476: The production of amino acids and lipids . In hot and dry conditions , plants close their stomata to prevent water loss. Under these conditions, CO 2 will decrease and oxygen gas , produced by the light reactions of photosynthesis, will increase, causing an increase of photorespiration by the oxygenase activity of ribulose-1,5-bisphosphate carboxylase/oxygenase (RuBisCO) and decrease in carbon fixation. Some plants have evolved mechanisms to increase
13734-486: The relationship between the two can be complex. For example, the light reaction creates ATP and NADPH energy molecules , which C 3 plants can use for carbon fixation or photorespiration . Electrons may also flow to other electron sinks. For this reason, it is not uncommon for authors to differentiate between work done under non-photorespiratory conditions and under photorespiratory conditions . Chlorophyll fluorescence of photosystem II can measure
13860-462: The respective organisms . In plants , light-dependent reactions occur in the thylakoid membranes of the chloroplasts where they drive the synthesis of ATP and NADPH . The light-dependent reactions are of two forms: cyclic and non-cyclic . In the non-cyclic reaction, the photons are captured in the light-harvesting antenna complexes of photosystem II by chlorophyll and other accessory pigments (see diagram at right). The absorption of
13986-455: The seabed between adjoining plates to form mid-oceanic ridges and here convection currents within the mantle tend to drive the two plates apart. Parallel to these ridges and nearer the coasts, one oceanic plate may slide beneath another oceanic plate in a process known as subduction . Deep trenches are formed here and the process is accompanied by friction as the plates grind together. The movement proceeds in jerks which cause earthquakes, heat
14112-407: The seabed causing deltas to form in estuaries. All these materials move back and forth under the influence of waves, tides and currents. Dredging removes material and deepens channels but may have unexpected effects elsewhere on the coastline. Governments make efforts to prevent flooding of the land by the building of breakwaters , seawalls , dykes and levees and other sea defences. For instance,
14238-401: The shore at the rate of six to eight per minute and these are known as constructive waves as they tend to move material up the beach and have little erosive effect. Storm waves arrive on shore in rapid succession and are known as destructive waves as the swash moves beach material seawards. Under their influence, the sand and shingle on the beach is ground together and abraded. Around high tide,
14364-495: The simpler photopigment retinal and its microbial rhodopsin derivatives are used to absorb green light and power proton pumps to directly synthesize adenosine triphosphate (ATP), the "energy currency" of cells. Such archaeal photosynthesis might have been the earliest form of photosynthesis that evolved on Earth, as far back as the Paleoarchean , preceding that of cyanobacteria (see Purple Earth hypothesis ). While
14490-479: The site of carboxylation in the chloroplast, to replace Ci. CO 2 concentration in the chloroplast becomes possible to estimate with the measurement of mesophyll conductance or g m using an integrated system. Photosynthesis measurement systems are not designed to directly measure the amount of light the leaf absorbs, but analysis of chlorophyll fluorescence , P700 - and P515-absorbance, and gas exchange measurements reveal detailed information about, e.g.,
14616-399: The source of the food supply which sustains most of the ocean ecosystem . Ocean photosynthesis also produces half of the oxygen in the Earth's atmosphere. Light can only penetrate a few hundred more meters; the rest of the deeper ocean is cold and dark (these zones are called mesopelagic and aphotic zones). The continental shelf is where the ocean meets dry land. It is more shallow, with
14742-412: The surface and is defined to be "the depth at which light intensity is only 1% of the surface value" (approximately 200 m in the open ocean). This is the zone where photosynthesis can occur. In this process plants and microscopic algae (free floating phytoplankton ) use light, water, carbon dioxide, and nutrients to produce organic matter. As a result, the photic zone is the most biodiverse and
14868-456: The system forms). As the world's ocean is the principal component of Earth's hydrosphere , it is integral to life on Earth, forms part of the carbon cycle and water cycle , and – as a huge heat reservoir – influences climate and weather patterns. The motions of the ocean surface, known as undulations or wind waves , are the partial and alternate rising and falling of the ocean surface. The series of mechanical waves that propagate along
14994-468: The three-carbon 3-phosphoglyceric acids directly in the Calvin-Benson cycle . Over 90% of plants use C 3 carbon fixation, compared to 3% that use C 4 carbon fixation; however, the evolution of C 4 in over sixty plant lineages makes it a striking example of convergent evolution . C 2 photosynthesis , which involves carbon-concentration by selective breakdown of photorespiratory glycine,
15120-424: The ultimate reduction of NADP to NADPH . In addition, this creates a proton gradient (energy gradient) across the chloroplast membrane , which is used by ATP synthase in the synthesis of ATP . The chlorophyll molecule ultimately regains the electron it lost when a water molecule is split in a process called photolysis , which releases oxygen . The overall equation for the light-dependent reactions under
15246-434: The very clearest ocean water, and is similar to blue light scattering in the sky . Ocean water represents the largest body of water within the global water cycle (oceans contain 97% of Earth's water ). Evaporation from the ocean moves water into the atmosphere to later rain back down onto land and the ocean. Oceans have a significant effect on the biosphere . The ocean as a whole is thought to cover approximately 90% of
15372-408: The water-oxidizing reaction (Kok's S-state diagrams). The hydrogen ions are released in the thylakoid lumen and therefore contribute to the transmembrane chemiosmotic potential that leads to ATP synthesis . Oxygen is a waste product of light-dependent reactions, but the majority of organisms on Earth use oxygen and its energy for cellular respiration , including photosynthetic organisms . In
15498-476: The world, and from the surface into the deep ocean. All this has impacts on the global climate system . Ocean water contains dissolved gases, including oxygen , carbon dioxide and nitrogen . An exchange of these gases occurs at the ocean's surface. The solubility of these gases depends on the temperature and salinity of the water. The carbon dioxide concentration in the atmosphere is rising due to CO 2 emissions , mainly from fossil fuel combustion. As
15624-710: Was bordered by Panthalassic Ocean to the north, separating it from Panthalassa by island arcs and Kazakhstania . The Proto-Tethys expanded during the Cambrian . The ocean was at its widest during the Late Ordovician to Middle Silurian . The ocean was situated between the Siberia to the west, and Gondwana to the east. The ocean began to shrink during the Late Silurian, when North China , and South China moved away from Gondwana and headed north. In
15750-462: Was emitted, hence the name cyclic reaction . Linear electron transport through a photosystem will leave the reaction center of that photosystem oxidized . Elevating another electron will first require re-reduction of the reaction center. The excited electrons lost from the reaction center ( P700 ) of photosystem I are replaced by transfer from plastocyanin , whose electrons come from electron transport through photosystem II . Photosystem II, as
15876-775: Was recovered from the Isua Greenstone Belt and provides evidence that water existed on Earth 3.8 billion years ago. In the Nuvvuagittuq Greenstone Belt , Quebec , Canada, rocks dated at 3.8 billion years old by one study and 4.28 billion years old by another show evidence of the presence of water at these ages. If oceans existed earlier than this, any geological evidence either has yet to be discovered, or has since been destroyed by geological processes like crustal recycling . However, in August 2020, researchers reported that sufficient water to fill
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