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Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.

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54-494: See text Verbena ( / v ər ˈ b iː n ə / ), also known as vervain or verveine , is a genus in the family Verbenaceae . It contains about 150 species of annual and perennial herbaceous or semi-woody flowering plants . The majority of the species are native to the Americas and Asia; however, Verbena officinalis , the common vervain or common verbena, is the type species and native to Europe. In English,

108-531: A diploid North American and a polyploid South American lineage, both with a base chromosome number of seven. The European species is derived from the North American lineage. It seems that verbena as well as the related mock vervains ( Glandularia ) evolved from the assemblage provisionally treated under the genus name Junellia ; both other genera were usually included in the Verbenaceae until

162-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

216-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

270-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

324-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

378-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

432-617: A twice-monthly professional medical journal , the British Journal of Cancer . The charity worked to bring about the smoking ban in England and continues to campaign for further action on smoking. The charity lobbies for better screening programmes and advises on access to new cancer medicines. Income sources include: On 18 July 2012, it was announced that Cancer Research UK was to receive its largest single donation of £10 million from an anonymous donor. The money went towards

486-584: A unique clinical trials database. A team of nurses provides a confidential telephone service, the Cancer Chat forum provides a place for users to talk to others affected by cancer, and mobile cancer awareness units deliver health information to locations of high cancer incidence and mortality. It provides statistical information via the Cancer Stats section. It also provides publications for the public to order and download. Cancer Research UK publishes

540-475: Is awarded to locations performing the highest quality cancer research, to provide funds for equipment and training. Centre status has been designated to: Drugs developed by the organisation's scientists include: Several of the organisation's scientists have won major prizes, including: Through Cancer Health UK, a website written in Plain English , it provides information on cancer and cancer care, and

594-399: Is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of

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648-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

702-512: Is employed in the preparation of a mandragora charm. The book also describes its antiseptic capabilities (p. 336), and use as a protection against spells (pp. 339, 414). Romani people use vervain for love and good luck. While common vervain is not native to North America, it has been introduced there and for example the Pawnee have adopted it as an entheogen enhancer and in oneiromancy (dream divination), much as Calea zacatechichi

756-524: Is no high-quality evidence for its effectiveness. Verbena has been listed as one of the 38 plants used to prepare Bach flower remedies , a kind of alternative medicine promoted for its effect on health. According to Cancer Research UK , "essence therapists believe that using essences can help to increase your mental, emotional and spiritual wellbeing. However, essences are not used to prevent, control, or cure cancer or any other physical condition." The essential oil of various species, mainly common vervain,

810-589: Is so renowned among the Romans as hiera botane ('sacred plant'). Some call it aristereon, and Latin writers verbenaca. This is the plant which I mentioned as carried to the enemy by envoys. With this the table of Jupiter is swept, and homes are cleansed and purified. There are two kinds of it; one has many leaves and is thought to be female, the other, the male, has fewer leaves. Pliny the Elder notes "the Magi especially make

864-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

918-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

972-597: Is the Latin term for a plant sacred to the ancient Romans. Pliny the Elder describes verbena presented on Jupiter altars; it is not entirely clear if this referred to a verbena rather than the general term for prime sacrificial herbs. Nulla tamen Romae nobilitatis plus habet quam hiera botane. aliqui aristereon, nostri verbenacam vocant. haec est quam legatos ferre ad hostes indicavimus; hac Iovis mensa verritur, domus purgantur lustranturque. genera eius duo: foliosa, quam feminam putant, mas rarioribus foliis. No plant however

1026-404: Is traded as "Spanish verbena oil". Considered inferior to oil of lemon verbena ( Aloysia citrodora ) in perfumery , it is of some commercial importance for herbalism. Verbena has long been associated with divine and other supernatural forces. It was called "tears of Isis " in ancient Egypt, and later called "Hera's tears". In ancient Greece it was dedicated to Eos Erigineia . The generic name

1080-580: Is used in Mexico. An indeterminate vervain is among the plants on the eighth panel of the New World Tapestry ( Expedition to Cape Cod ). In the Victorian language of flowers , verbena held the dual meaning of enchantment and sensibility. The following species are accepted: (See also Aloysia and Junellia for species formerly placed here.) Genus The composition of a genus

1134-814: The Hummingbird hawk-moth , Chocolate albatross , or the Pipevine swallowtail , and also hummingbirds, especially V. officinalis , which is also grown as a honey plant . The hybrid cultivars "Silver Anne" and " Sissinghurst " have gained the Royal Horticultural Society 's Award of Garden Merit . For some verbena pathogens, see List of verbena diseases . Cultivated verbenas are sometimes parasitized by sweet potato whitefly ( Bemisia tabaci ) and spread this pest to other crops. Although verbena ("vervain") has been used in herbalism and traditional medicine , usually as an herbal tonic , there

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1188-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

1242-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

1296-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

1350-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

1404-426: The 1990s. Intergeneric chloroplast gene transfer by an undetermined mechanism – though probably not hybridization – has occurred at least twice from vervains to Glandularia , between the ancestors of the present-day South American lineages and once more recently, between V. orcuttiana or V. hastata and G. bipinnatifida . In addition, several species of verbena are of natural hybrid origin;

1458-656: The Cancer Research UK London Research Institute , are now part of the Francis Crick Institute . The British Empire Cancer Campaign (BECC) was founded in 1923, and initially drew a hostile response from ICRF and the Medical Research Council , who considered it a rival. "The Campaign", as it was colloquially known, became a very successful and powerful grant-giving body. In 1970, the charity

1512-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

1566-570: The early Christian era, folk legend stated that V. officinalis was used to stanch Jesus' wounds after his removal from the cross. It was consequently called "holy herb" or (e.g. in Wales) "Devil's bane". According to the Wiccan writer Doreen Valiente , Vervain flowers signify the goddess Diana and are often depicted on cimaruta , traditional Italian amulets. In the 1870 The History and Practice of Magic by "Paul Christian" (Jean-Baptiste Pitois) it

1620-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

1674-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

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1728-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

1782-473: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Cancer Research UK Cancer Research UK ( CRUK ) is the world's largest independent cancer research organisation. It is registered as a charity in the United Kingdom and Isle of Man , and was formed on 4 February 2002 by

1836-558: The maddest statements about the plant: that [among other things] a circle must be drawn with iron round the plant". The common names of verbena in many Central and Eastern European languages often associate it with iron . These include for example the Dutch IJzerhard ("iron-hard"), Danish Læge-Jernurt ("medical ironwort"), German Echtes Eisenkraut ("true ironherb"), Slovak Železník lekársky ("medical ironherb"), and Hungarian vasfű ("iron grass"). In

1890-650: The merger of The Cancer Research Campaign and the Imperial Cancer Research Fund . Cancer Research UK conducts research using both its own staff and grant-funded researchers. It also provides information about cancer and runs campaigns aimed at raising awareness and influencing public policy. The organisation's work is almost entirely funded by the public. It raises money through donations, legacies, community fundraising, events, retail and corporate partnerships. Over 25,000 people are regular volunteers. The Imperial Cancer Research Fund (ICRF)

1944-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

1998-698: The name Verbena is usually used in the United States and the United Kingdom, whereas elsewhere the terms verveine or vervain are in use. Verbena is an herbaceous flowering plant, belonging to the Verbenaceae family , and may be annual or perennial depending on the species . The leaves are usually opposite, simple, and in many species hairy, often densely so. The flowers are small, with five petals, and borne in dense spikes. Typically some shade of blue, they may also be white, pink, or purple, especially in cultivars . The genus can be divided into

2052-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

2106-557: The period between January 2015 to August 2019, Nature listed Cancer Research UK in the top 150 of the Top 200 institutions in cancer research in the world. CRUK had an income of £718,793,138 and expenditure of £640,845,146 for the financial year ending in 31 March 2023. On 30 April 2024, King Charles III was announced as patron of the charity. In the financial year 2014/15, the charity spent £422.67 million on cancer research projects (67% of its total income for that year). The bulk of

2160-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

2214-644: The remaining costs were spent on trading and fundraising costs with a small amount spent on information services, campaigning, advocacy, administration and other activities or was held in reserve. Around 40% of its research expenditure (27% of its total spending) is on basic laboratory research into the molecular basis of cancer . The remainder supports research into over 100 specific cancer types, focusing on drug discovery and development; prevention, early detection and imaging ; surgery and radiotherapy ; and cancers where survival rates are still low, such as oesophageal, lung and pancreatic cancers. The charity funds

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2268-470: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or

2322-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in

2376-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

2430-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

2484-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

2538-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

2592-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

2646-613: The well-known garden vervain/verbena has an entirely muddy history. The relationships of this close-knit group are therefore hard to resolve with standard methods of computational phylogenetics . Some species, hybrids and cultivars of verbena are used as ornamental plants . They are drought-resistant, tolerating full to partial sun, and enjoy well-drained, average soils. Plants are usually grown from seed. Some species and hybrids are not hardy and are treated as half-hardy annuals in bedding schemes. They are valued in butterfly gardening in suitable climates, attracting Lepidoptera such as

2700-476: The work of over 4,000 researchers, doctors and nurses throughout the UK, supports over 200 clinical trials and studies cancer and cancer risk in over a million people in the UK. The charity participates in numerous citizen-science projects including: The charity funds networks in seven locations across the UK, to drive collaborations between universities, NHS hospitals, and other research organisations. Centre status

2754-615: The £100 million funding for the Francis Crick Institute in London. During the COVID-19 pandemic in 2020, CRUK closed its shops and cancelled mass participation fundraising events. They predicted that this, coupled with economic uncertainty affecting people's ability or willingness to donate, would lead to a 30% fall in income that year and a reduction in income lasting at least 3 years. In June 2011, Cancer Research UK

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2808-586: Was founded in 1902 as the Cancer Research Fund, changing its name to the Imperial Cancer Research Fund in 1904. It grew over the next twenty years to become one of the world's leading cancer research charities. Its executive committee was chaired by Sir William Church from its inception in 1902 until 1923. Its flagship laboratories formerly at Lincoln's Inn Fields , London , and Clare Hall, Hertfordshire , and known as

2862-721: Was one of several health charities (along with the British Heart Foundation , the Alzheimer's Society and Parkinson's UK ) targeted by the animal rights organisation Animal Aid in a series of advertisements in British newspapers urging members of the public to stop giving donations to organisations that fund medical research involving animal experiments. In April 2017, the Information Commissioner's Office fined eleven charities that breached

2916-701: Was renamed The Cancer Research Campaign (CRC). Incorporated on 20 November 2001, the two organisations officially merged on 4 February 2002 to form Cancer Research UK, the largest independent cancer research organisation in the world (the largest, the National Cancer Institute , is funded by the US Government). At the time of the merger, the ICRF had an annual income of £124m, while the CRC had an income of £101m. Based on article share during

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