Ploidy ( / ˈ p l ɔɪ d i / ) is the number of complete sets of chromosomes in a cell , and hence the number of possible alleles for autosomal and pseudoautosomal genes . Here sets of chromosomes refers to the number of maternal and paternal chromosome copies, respectively, in each homologous chromosome pair—the form in which chromosomes naturally exist. Somatic cells , tissues , and individual organisms can be described according to the number of sets of chromosomes present (the "ploidy level"): monoploid (1 set), diploid (2 sets), triploid (3 sets), tetraploid (4 sets), pentaploid (5 sets), hexaploid (6 sets), heptaploid or septaploid (7 sets), etc. The generic term polyploid is often used to describe cells with three or more sets of chromosomes.
126-479: Abacá ( / ɑː b ə ˈ k ɑː / ah-bə- KAH ; Filipino : abaka [ɐbɐˈka] ), also known as Manila hemp , is a species of banana , Musa textilis , endemic to the Philippines . The plant grows to 13–22 feet (4.0–6.7 m), and averages about 12 feet (3.7 m). The plant has great economic importance, being harvested for its fiber extracted from the leaf-stems. The lustrous fiber
252-535: A ploidy series , featuring diploid ( X. tropicalis , 2n=20), tetraploid ( X. laevis , 4n=36), octaploid ( X. wittei , 8n=72), and dodecaploid ( X. ruwenzoriensis , 12n=108) species. Over evolutionary time scales in which chromosomal polymorphisms accumulate, these changes become less apparent by karyotype – for example, humans are generally regarded as diploid, but the 2R hypothesis has confirmed two rounds of whole genome duplication in early vertebrate ancestors. Ploidy can also vary between individuals of
378-417: A syncytium , though usually the ploidy of each nucleus is described individually. For example, a fungal dikaryon with two separate haploid nuclei is distinguished from a diploid cell in which the chromosomes share a nucleus and can be shuffled together. It is possible on rare occasions for ploidy to increase in the germline , which can result in polyploid offspring and ultimately polyploid species. This
504-675: A zygote with n pairs of chromosomes, i.e. 2 n chromosomes in total. The chromosomes in each pair, one of which comes from the sperm and one from the egg, are said to be homologous . Cells and organisms with pairs of homologous chromosomes are called diploid. For example, most animals are diploid and produce haploid gametes. During meiosis , sex cell precursors have their number of chromosomes halved by randomly "choosing" one member of each pair of chromosomes, resulting in haploid gametes. Because homologous chromosomes usually differ genetically, gametes usually differ genetically from one another. All plants and many fungi and algae switch between
630-556: A "Modernizing the Language Approach Movement" (MOLAM). Lacuesta hosted a number of "anti-purist" conferences and promoted a "Manila Lingua Franca" which would be more inclusive of loanwords of both foreign and local languages. Lacuesta managed to get nine congressmen to propose a bill aiming to abolish the SWP with an Akademia ng Wikang Filipino , to replace the balarila with a Gramatica ng Wikang Filipino , to replace
756-468: A "false trunk" or pseudostem about 6–15 inches (15–38 cm) in diameter. The leaf stalks ( petioles ) are expanded at the base to form sheaths that are tightly wrapped together to form the pseudostem. There are from 12 to 25 leaves, dark green on the top and pale green on the underside, sometimes with large brown patches. They are oblong in shape with a deltoid base. They grow in succession. The petioles grow to at least 1 foot (30 cm) in length. When
882-607: A 39% share of Philippine abacá production of which an overwhelming 92% came from Catanduanes Island . Eastern Visayas, the second largest producer had 24% and the Davao Region, the third largest producer had 11% of the total production. Around 42 percent of the total abacá fiber shipments from the Philippines went to the United Kingdom in 2014, making it the top importer. Germany imported 37.1 percent abacá pulp from
1008-467: A chromosome copy number of 1 to 4, and that number is commonly fractional, counting portions of the chromosome partly replicated at a given time. This is because under exponential growth conditions the cells are able to replicate their DNA faster than they can divide. In ciliates, the macronucleus is called ampliploid , because only part of the genome is amplified. Mixoploidy is the case where two cell lines, one diploid and one polyploid, coexist within
1134-517: A common Malayo-Polynesian language due to the Austronesian migration from Taiwan. The common Malayo-Polynesian language split into different languages, and usually through the Malay language, the lingua franca of maritime Southeast Asia, these were able to adopt terms that ultimately originate from other languages such as Japanese , Hokkien , Sanskrit , Tamil , and Arabic . The Malay language
1260-585: A common national language, termed Filipino , to replace Pilipino. Neither the original nor the amended version specified either Tagalog or Pilipino as the basis for Filipino; Instead, tasking the National Assembly to: take steps toward the development and formal adoption of a common national language to be known as Filipino. In 1987, a new constitution designated Filipino as the national language and, along with English, as an official language. That constitution included several provisions related to
1386-424: A germ cell with an uneven number of chromosomes undergoes meiosis, the chromosomes cannot be evenly divided between the daughter cells, resulting in aneuploid gametes. Triploid organisms, for instance, are usually sterile. Because of this, triploidy is commonly exploited in agriculture to produce seedless fruit such as bananas and watermelons. If the fertilization of human gametes results in three sets of chromosomes,
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#17327730148271512-629: A haploid and a diploid state, with one of the stages emphasized over the other. This is called alternation of generations . Most fungi and algae are haploid during the principal stage of their life cycle, as are some primitive plants like mosses . More recently evolved plants, like the gymnosperms and angiosperms , spend the majority of their life cycle in the diploid stage. Most animals are diploid, but male bees , wasps , and ants are haploid organisms because they develop from unfertilized, haploid eggs, while females (workers and queens) are diploid, making their system haplodiploid . In some cases there
1638-653: A large scale in Sumatra in 1925 under the Dutch , who had observed its cultivation in the Philippines for cordage since the nineteenth century, followed up by plantings in Central America in 1929 sponsored by the U.S. Department of Agriculture . It also was transplanted into India and Guam . Commercial planting began in 1930 in British North Borneo ; at the onset of World War II , the supply from
1764-452: A major trade commodity in the colonial era for this reason. The abaca industry declined sharply in the mid-20th century when abaca plantations were decimated by World War II and plant diseases, as well as the invention of nylon in the 1930s. Today, abaca is mostly used in a variety of specialized paper products including tea bags , filter paper and banknotes . Manila envelopes and Manila paper derive their name from this fiber. Abaca
1890-649: A native Cebuano name). They were exported to other Spanish colonies since the 16th century. A waistcoat of a native Quechua man in Peru was recorded as being made of medriñaque as early as 1584. Abacá cloth also appear in English records, spelled variously as medrinacks , medrianacks , medrianackes , and medrinacles , among other names. They were used as canvas for sails and for stiffening clothing like skirts, collars, and doublets . Philippine indigenous tribes still weave abacá-based textiles like t'nalak , made by
2016-451: A number of chromosomes that is an exact multiple of the number of chromosomes in a normal gamete; and having any other number, respectively. For example, a person with Turner syndrome may be missing one sex chromosome (X or Y), resulting in a (45,X) karyotype instead of the usual (46,XX) or (46,XY). This is a type of aneuploidy and cells from the person may be said to be aneuploid with a (diploid) chromosome complement of 45. The term ploidy
2142-591: A period of time when exposed to saltwater. Manila hemp rope was previously the favoured variety of rope used for executions by hanging, both in the U.K. and USA. Usually 3/4 to 1 inch diameter, boiled prior to use to take out any overelasticity. It was also used in the 19th century as whaling line. Abacá fiber was once used primarily for rope, but this application is now of minor significance. Filipino language Filipino ( English: / ˌ f ɪ l ɪ ˈ p iː n oʊ / , FIH-lih-PEE-noh ; Wikang Filipino , [ˈwi.kɐŋ fi.liˈpi.no̞] )
2268-540: A preferred growing environment. Harvesting generally includes several operations involving the leaf sheaths: When the processing is complete, the bundles of fiber are pale and lustrous with a length of 6–12 feet (1.8–3.7 m). In Costa Rica, more modern harvest and drying techniques are being developed to accommodate the very high yields obtained there. According to the Philippine Fiber Industry Development Authority ,
2394-425: A process called endoreduplication , where duplication of the genome occurs without mitosis (cell division). The extreme in polyploidy occurs in the fern genus Ophioglossum , the adder's-tongues, in which polyploidy results in chromosome counts in the hundreds, or, in at least one case, well over one thousand. It is possible for polyploid organisms to revert to lower ploidy by haploidisation . Polyploidy
2520-627: A resolution on November 9, 1937 recommending Tagalog to be basis of the national language. On December 30, President Quezon issued Executive Order No. 134, s. 1937, approving the adoption of Tagalog as the language of the Philippines, and proclaimed the national language of the Philippines so based on the Tagalog language. Quezon himself was born and raised in Baler, Aurora , which is a native Tagalog-speaking area. The order stated that it would take effect two years from its promulgation. On December 31 of
2646-401: A single nucleus rather than in the cell as a whole. Because in most situations there is only one nucleus per cell, it is commonplace to speak of the ploidy of a cell, but in cases in which there is more than one nucleus per cell, more specific definitions are required when ploidy is discussed. Authors may at times report the total combined ploidy of all nuclei present within the cell membrane of
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#17327730148272772-437: A single chromosome and diploid individuals have two chromosomes. In Entamoeba , the ploidy level varies from 4 n to 40 n in a single population. Alternation of generations occurs in most plants, with individuals "alternating" ploidy level between different stages of their sexual life cycle. In large multicellular organisms, variations in ploidy level between different tissues, organs, or cell lineages are common. Because
2898-495: A single copy of each chromosome – that is, one and only one set of chromosomes. In this case, the nucleus of a eukaryotic cell is said to be haploid only if it has a single set of chromosomes , each one not being part of a pair. By extension a cell may be called haploid if its nucleus has one set of chromosomes, and an organism may be called haploid if its body cells (somatic cells) have one set of chromosomes per cell. By this definition haploid therefore would not be used to refer to
3024-484: A single parent, without the involvement of gametes and fertilization, and all the offspring are genetically identical to each other and to the parent, including in chromosome number. The parents of these vegetative clones may still be capable of producing haploid gametes in preparation for sexual reproduction, but these gametes are not used to create the vegetative offspring by this route. Some eukaryotic genome-scale or genome size databases and other sources which may list
3150-429: A week and was known as Linggo ng Wika (Language Week). The celebration coincides with the month of birth of President Manuel L. Quezon, regarded as the "Ama ng Wikang Pambansa" (Father of the national language). In 1946, Proclamation No. 35 of March 26 provided for a week-long celebration of the national language. this celebration would last from March 27 until April 2 each year, the last day coinciding with birthday of
3276-490: Is 24. The monoploid number equals the total chromosome number divided by the ploidy level of the somatic cells: 48 chromosomes in total divided by a ploidy level of 4 equals a monoploid number of 12. Hence, the monoploid number (12) and haploid number (24) are distinct in this example. However, commercial potato crops (as well as many other crop plants) are commonly propagated vegetatively (by asexual reproduction through mitosis), in which case new individuals are produced from
3402-402: Is 7. The gametes of common wheat are considered to be haploid, since they contain half the genetic information of somatic cells, but they are not monoploid, as they still contain three complete sets of chromosomes ( n = 3 x ). In the case of wheat, the origin of its haploid number of 21 chromosomes from three sets of 7 chromosomes can be demonstrated. In many other organisms, although
3528-519: Is a back-formation from haploidy and diploidy . "Ploid" is a combination of Ancient Greek -πλόος (-plóos, "-fold") and -ειδής (- eidḗs ), from εἶδος ( eîdos , "form, likeness"). The principal meaning of the Greek word ᾰ̔πλόος (haplóos) is "single", from ἁ- (ha-, "one, same"). διπλόος ( diplóos ) means "duplex" or "two-fold". Diploid therefore means "duplex-shaped" (compare "humanoid", "human-shaped"). Polish-German botanist Eduard Strasburger coined
3654-413: Is a characteristic of the bacterium Deinococcus radiodurans and of the archaeon Halobacterium salinarum . These two species are highly resistant to ionizing radiation and desiccation , conditions that induce DNA double-strand breaks. This resistance appears to be due to efficient homologous recombinational repair. Depending on growth conditions, prokaryotes such as bacteria may have
3780-522: Is a language under the Austronesian language family . It is the national language ( Wikang pambansa / Pambansang wika ) of the Philippines , lingua franca (Karaniwang wika), and one of the two official languages ( Wikang opisyal / Opisyal na wika ) of the country, with English . It is a standardized variety of the native language Tagalog , spoken and written in Metro Manila ,
3906-801: Is a translation of Article 1 of the Universal Declaration of Human Rights . Usually, the diacritics are not written, and the syntax and grammar are based on that of Tagalog . the General Assembly proclaims this UNIVERSAL DECLARATION OF HUMAN RIGHTS as a common standard of achievement for all peoples and all nations, to the end that every individual and every organ of society, keeping this Declaration constantly in mind, shall strive by teaching and education to promote respect for these rights and freedoms and by progressive measures, national and international, to secure their universal and effective recognition and observance, both among
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4032-462: Is also common among Austronesian languages. It has head-initial directionality. It is an agglutinative language but can also display inflection . It is not a tonal language and can be considered a pitch-accent language and a syllable-timed language. It has nine basic parts of speech . The Philippines is a multilingual state with 175 living languages originating and spoken by various ethno-linguistic groups. Many of these languages descend from
4158-412: Is an important evolutionary mechanism in both plants and animals and is known as a primary driver of speciation . As a result, it may become desirable to distinguish between the ploidy of a species or variety as it presently breeds and that of an ancestor. The number of chromosomes in the ancestral (non-homologous) set is called the monoploid number ( x ), and is distinct from the haploid number ( n ) in
4284-600: Is argued that current state of the Filipino language is contrary to the intention of Republic Act (RA) No. 7104 that requires that the national language be developed and enriched by the lexicon of the country's other languages. It is further argued that, while the official view (shared by the government, the Komisyon sa Wikang Filipino , and a number of educators) is that Filipino and Tagalog are considered separate languages, in practical terms, Filipino may be considered
4410-462: Is associated with a 14% lower risk of being endangered, and a 20% greater chance of being invasive. Polyploidy may be associated with increased vigor and adaptability. Some studies suggest that selection is more likely to favor diploidy in host species and haploidy in parasite species. However, polyploidization is associated with an increase in transposable element content and relaxed purifying selection on recessive deleterious alleles. When
4536-433: Is classified as a hard fiber , along with coir , henequin and sisal . Abaca is grown as a commercial crop in the Philippines, Ecuador, Costa Rica. The abacá plant is stoloniferous , meaning that the plant produces runners or shoots along the ground that then root at each segment. Cutting and transplanting rooted runners is the primary technique for creating new plants, since seed growth is substantially slower. Abacá has
4662-525: Is evidence that the n chromosomes in a haploid set have resulted from duplications of an originally smaller set of chromosomes. This "base" number – the number of apparently originally unique chromosomes in a haploid set – is called the monoploid number , also known as basic or cardinal number , or fundamental number . As an example, the chromosomes of common wheat are believed to be derived from three different ancestral species, each of which had 7 chromosomes in its haploid gametes. The monoploid number
4788-471: Is faster than diploid under high nutrient conditions. The NLH is also tested in haploid, diploid, and polyploid fungi by Gerstein et al. 2017. This result is also more complex: On the one hand, under phosphorus and other nutrient limitation, lower ploidy is selected as expected. However under normal nutrient levels or under limitation of only nitrogen , higher ploidy was selected. Thus the NLH – and more generally,
4914-401: Is frequently used. While the word Tagalista literally means "one who specializes in Tagalog language or culture" or a "Tagalog specialist", in the context of the debates on the national language and " Imperial Manila ", the word Tagalista is used as a reference to "people who promote or would promote the primacy of Tagalog at the expense of [the] other [Philippine] indigenous tongues". This
5040-770: Is generally known as sinamáy in most of the islands. Abacá cloth with a more delicate texture is called tinampipi . While especially fine lace-like abacá cloth is called nipis or lupis . Fine abacá fibers may also be woven with piña , silk , or fine cotton to create a fabric called jusi . Traditional abacá textiles were often dyed in various colors from various natural dyes . These include blue from indigo ( tarum , dagum , tayum , etc.); black from ebony ( knalum or batulinao ) leaves; red from noni roots and sapang ; yellow from turmeric ( kalawag , kuning , etc.); and so on. They were often woven into specific patterns, and further ornamented with embroidery, beadwork, and other decorations. Most clothing made from abacá took
5166-437: Is normally grown in well-drained loamy soil, using rhizomes planted at the start of the rainy season. In addition, new plants can be started by seeds. Growers harvest abacá fields every three to eight months after an initial growth period of 12–25 months. Harvesting is done by removing the leaf-stems after flowering but before the fruit appears. The plant loses productivity between 15 to 40 years. The slopes of volcanoes provide
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5292-483: Is not a wanted feature, it should be well taken into account. Since shrinkage is more pronounced the first time the rope becomes wet, new rope is usually immersed into water and put to dry before use so that the shrinkage is less than it would be if the rope had never been wet. A major disadvantage in this shrinkage is that many knots made with manila rope became harder and more difficult to untie when wet, thus becoming subject of increased stress. Manila rope will rot after
5418-600: Is placed in section Callimusa (now including the former section Australimusa ), members of which have a diploid chromosome number of 2n = 20. The Philippines, especially the Bicol region in Luzon, has the most abaca genotypes and cultivars. Genetic analysis using simple sequence repeats (SSR) markers revealed that the Philippines' abaca germplasm is genetically diverse. Abaca genotypes in Luzon had higher genetic diversity than Visayas and Mindanao. Ninety-five (95) percent
5544-485: Is that speech variety spoken in Metro Manila and other urban centers where different ethnic groups meet. It is the most prestigious variety of Tagalog and the language used by the national mass media. The other yardstick for distinguishing a language from a dialect is: different grammar, different language. "Filipino", "Pilipino" and "Tagalog" share identical grammar. They have the same determiners (ang, ng and sa);
5670-414: Is the indigenous written and spoken language of Metro Manila and other urban centers in the Philippines used as the language of communication of ethnic groups . However, as with the 1973 and 1987 Constitutions, 92-1 went neither so far as to categorically identify, nor so far as to dis-identify this language as Tagalog. Definite, absolute, and unambiguous interpretation of 92–1 is the prerogative of
5796-572: Is the simplest to illustrate in diagrams of genetics concepts. But this definition also allows for haploid gametes with more than one set of chromosomes. As given above, gametes are by definition haploid, regardless of the actual number of sets of chromosomes they contain. An organism whose somatic cells are tetraploid (four sets of chromosomes), for example, will produce gametes by meiosis that contain two sets of chromosomes. These gametes might still be called haploid even though they are numerically diploid. An alternative usage defines "haploid" as having
5922-426: Is the state in which the chromosomes are paired and can undergo meiosis. The zygoid state of a species may be diploid or polyploid. In the azygoid state the chromosomes are unpaired. It may be the natural state of some asexual species or may occur after meiosis. In diploid organisms the azygoid state is monoploid. (See below for dihaploidy.) In the strictest sense, ploidy refers to the number of sets of chromosomes in
6048-731: Is the state where all cells have multiple sets of chromosomes beyond the basic set, usually 3 or more. Specific terms are triploid (3 sets), tetraploid (4 sets), pentaploid (5 sets), hexaploid (6 sets), heptaploid or septaploid (7 sets), octoploid (8 sets), nonaploid (9 sets), decaploid (10 sets), undecaploid (11 sets), dodecaploid (12 sets), tridecaploid (13 sets), tetradecaploid (14 sets), etc. Some higher ploidies include hexadecaploid (16 sets), dotriacontaploid (32 sets), and tetrahexacontaploid (64 sets), though Greek terminology may be set aside for readability in cases of higher ploidy (such as "16-ploid"). Polytene chromosomes of plants and fruit flies can be 1024-ploid. Ploidy of systems such as
6174-463: Is the strongest of the natural fibers. It is used by the paper industry for such specialty uses such as tea bags , banknotes and decorative papers. It can be used to make handcrafts such as hats, bags, carpets, clothing and furniture. Lupis is the finest quality of abacá. Sinamay is woven chiefly from abacá. Abacá fibers were traditionally woven into sturdy textiles and clothing in the Philippines since pre-colonial times. Along with cotton , they were
6300-402: Is thus 7 and the haploid number is 3 × 7 = 21. In general n is a multiple of x . The somatic cells in a wheat plant have six sets of 7 chromosomes: three sets from the egg and three sets from the sperm which fused to form the plant, giving a total of 42 chromosomes. As a formula, for wheat 2 n = 6 x = 42, so that the haploid number n is 21 and the monoploid number x
6426-428: Is traditionally hand-loomed into various indigenous textiles ( abaca cloth or medriñaque ) in the Philippines. They are still featured prominently as the traditional material of the barong tagalog , the national male attire of the Philippines, as well as in sheer lace-like fabrics called nipis used in various clothing components. Native abaca textiles also survive into the modern era among various ethnic groups, like
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#17327730148276552-475: Is very durable, flexible, and resistant to salt water damage, allowing its use in rope, hawsers , ships' lines, and fishing nets . A 1 inch (2.5 cm) rope can require 4 metric tons (8,800 lb) to break. Manilla rope is still the only material specified for lifeboat falls (the ropes with which a ship's lifeboat is lowered) in the United Kingdom. Manila ropes shrink when they become wet. This effect can be advantageous under certain circumstances, but if it
6678-417: Is well established in this original sense, but it has also been used for doubled monoploids or doubled haploids , which are homozygous and used for genetic research. Euploidy ( Greek eu , "true" or "even") is the state of a cell or organism having one or more than one set of the same set of chromosomes, possibly excluding the sex-determining chromosomes . For example, most human cells have 2 of each of
6804-650: The Komisyon sa Wikang Filipino (Commission on the Filipino Language, or KWF), superseding the Institute of Philippine Languages. The KWF reports directly to the President and was tasked to undertake, coordinate and promote researches for the development, propagation and preservation of Filipino and other Philippine languages. On May 13, 1992, the commission issued Resolution 92-1, specifying that Filipino
6930-509: The t'nalak of the T'boli people and the dagmay of the Bagobo people . Abaca is also used in traditional Philippine millinery , as well as for bags, shawls, and other decorative items. The hatmaking straw made from Manila hemp is called tagal or tagal straw . The fiber is also exceptionally strong, stronger than hemp and naturally salt-resistant, making it ideal for making twines and ropes (especially for maritime shipping). It became
7056-479: The American colonial period , English became an additional official language of the Philippines alongside Spanish; however, the number of speakers of Spanish steadily decreased. The United States initiated policies that led to the gradual removal of Spanish from official use in the Philippines. This was not done through an outright ban, but rather through a strategic shift in language policy that promoted English as
7182-697: The Balarílà ng Wikang Pambansâ (English: Grammar of the National Language ) of grammarian Lope K. Santos introduced the 20-letter Abakada alphabet which became the standard of the national language. The alphabet was officially adopted by the Institute for the Tagalog-Based National Language. In 1959, the language became known as Pilipino in an effort to disassociate it from the Tagalog ethnic group . The changing of
7308-547: The Mariano Marcos State University in Batac, Ilocos Norte, that Filipino was simply Tagalog in syntax and grammar, with as yet no grammatical element or lexicon coming from Ilokano , Cebuano , Hiligaynon , or any of the other Philippine languages . He said further that this is contrary to the intention of Republic Act No. 7104, which requires that the national language be developed and enriched by
7434-468: The Spanish colonial era , it was referred to as " medriñaque " cloth. By 1897, the Philippines were exporting almost 100,000 tons of abacá, and it was one of the three biggest cash crops, along with tobacco and sugar. In fact, from 1850 through the end of the 19th century, sugar or abacá alternated with each other as the biggest export crop of the Philippines. This 19th-century trade was predominantly with
7560-713: The Supreme Court in the absence of directives from the KWF, otherwise the sole legal arbiter of the Filipino language. Filipino was presented and registered with the International Organization for Standardization (ISO), by Ateneo de Manila University student Martin Gomez, and was added to the ISO registry of languages on September 21, 2004, with it receiving the ISO 639-2 code fil . On August 22, 2007, it
7686-619: The Tiboli tribe of South Cotabato , and dagmay , made by the Bagobo people. Abacá cloth is found in museum collections around the world, like the Boston Museum of Fine Arts and the Textile Museum of Canada. The inner fibers are also used in the making of hats, including the "Manila hats", hammocks, matting, cordage , ropes, coarse twines, and types of canvas. Manila rope is a type of rope made from manila hemp. Manila rope
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#17327730148277812-657: The United States and the making of ropes was done mainly in New England , although in time rope-making shifted back to the Philippines. From 1898 to 1946, the United States colonized the Philippines following the Spanish-American War . The Guggenheim claims the "colonial government found ways to prevent Filipinos from profiting off of the abaca crops, instead favoring the businesses of American expats and Japanese immigrants, as well as ensuring that
7938-423: The monoploid number ( x ). The haploid number ( n ) refers to the total number of chromosomes found in a gamete (a sperm or egg cell produced by meiosis in preparation for sexual reproduction). Under normal conditions, the haploid number is exactly half the total number of chromosomes present in the organism's somatic cells, with one paternal and maternal copy in each chromosome pair. For diploid organisms,
8064-425: The salivary gland , elaiosome , endosperm , and trophoblast can exceed this, up to 1048576-ploid in the silk glands of the commercial silkworm Bombyx mori . The chromosome sets may be from the same species or from closely related species. In the latter case, these are known as allopolyploids (or amphidiploids, which are allopolyploids that behave as if they were normal diploids). Allopolyploids are formed from
8190-466: The 20-letter Abakada with a 32-letter alphabet, and to prohibit the creation of neologisms and the respelling of loanwords. This movement quietened down following the death of Lacuesta. The national language issue was revived once more during the 1971 Constitutional Convention . While there was a sizable number of delegates in favor of retaining the Tagalog-based national language, majority of
8316-453: The 23 homologous monoploid chromosomes, for a total of 46 chromosomes. A human cell with one extra set of the 23 normal chromosomes (functionally triploid) would be considered euploid. Euploid karyotypes would consequentially be a multiple of the haploid number , which in humans is 23. Aneuploidy is the state where one or more individual chromosomes of a normal set are absent or present in more than their usual number of copies (excluding
8442-407: The 23 homologous pairs of chromosomes that humans normally have. This results in two homologous pairs within each of the 23 homologous pairs, providing a full complement of 46 chromosomes. This total number of individual chromosomes (counting all complete sets) is called the chromosome number or chromosome complement . The number of chromosomes found in a single complete set of chromosomes is called
8568-523: The Filipino language. Article XIV, Section 6, omits any mention of Tagalog as the basis for Filipino, and states that: as Filipino evolves, it shall be further developed and enriched on the basis of existing Philippine and other languages. And also states in the article: Subject to provisions of law and as the Congress may deem appropriate, the Government shall take steps to initiate and sustain
8694-422: The Filipino writer Francisco Baltazar , author of the Tagalog epic Florante at Laura . In 1954, Proclamation No. 12 of March 26 provided that the week of celebration would be from March 29 to April 4 every year. This proclamation was amended the following year by President Ramon Magsaysay by Proclamation No. 186 of September 23, moving the dates of celebration to August 13–19, every year. Now coinciding with
8820-500: The Institute of National Language (later the Surián ng Wikang Pambansâ or SWP) and tasking it with making a study and survey of each existing native language, hoping to choose which was to be the base for a standardized national language. Later, President Manuel L. Quezon later appointed representatives for each major regional language to form the NLI. Led by Jaime C. De Veyra , who sat as
8946-489: The National Capital Region, and in other urban centers of the archipelago. The 1987 Constitution mandates that Filipino be further enriched and developed by the other languages of the Philippines . Filipino, like other Austronesian languages, commonly uses verb-subject-object order, but can also use subject-verb-object order. Filipino follows the trigger system of morphosyntactic alignment that
9072-599: The Philippine islands started in 1565 with the fall of Cebu. The eventual capital established by Spain for its settlement in the Philippines was Manila , situated in a Tagalog-speaking region, after the capture of Manila from the Muslim Kingdom of Luzon ruled by Raja Matanda with the heir apparent Raja Sulayman and the Hindu-Buddhist Kingdom of Tondo ruled by Lakan Dula . After its fall to
9198-480: The Philippines provided that: The National Assembly shall take steps toward the development and adoption of a common national language based on one of the existing native languages. Until otherwise provided by law, English and Spanish shall continue as official languages. On November 13, 1936, the first National Assembly of the Philippine Commonwealth approved Commonwealth Act No. 184; creating
9324-594: The Philippines provided 87.4% of the world's abacá in 2014, earning the Philippines US$ 111.33 million. The demand is still greater than the supply. The remainder came from Ecuador (12.5%) and Costa Rica (0.1%). The Bicol region in the Philippines produced 27,885 metric tons of abacá in 2014, the largest of any Philippine region . The Philippine Rural Development Program (PRDP) and the Department of Agriculture reported that in 2009–2013, Bicol Region had
9450-535: The Philippines was eliminated by the Empire of Japan . After the war, the U.S. Department of Agriculture started production in Panama , Costa Rica , Honduras , and Guatemala . Today, abacá is produced primarily in the Philippines and Ecuador. The Philippines produces between 85% and 95% of the world's abacá, and the production employs 1.5 million people. Production has declined because of virus diseases. The plant
9576-477: The Philippines, importing around 7,755 metric tons (MT). Sales of abacá cordage surged 20 percent in 2014 to a total of 5,093 MT from 4,240 MT, with the United States holding around 68 percent of the market. Abacá is vulnerable to a number of pathogens, notably abaca bunchy top virus , abaca bract mosaic virus , and abaca mosaic virus . Due to its strength, it is a sought after product and
9702-924: The Spaniards, Manila was made the capital of the Spanish settlement in Asia due to the city's commercial wealth and influence, its strategic location, and Spanish fears of raids from the Portuguese and the Dutch. The first dictionary of Tagalog, published as the Vocabulario de la lengua tagala , was written by the Franciscan Pedro de San Buenaventura, and published in 1613 by the "Father of Filipino Printing" Tomás Pinpin in Pila , Laguna . A latter book of
9828-572: The absence or presence of complete sets, which is considered euploidy). Unlike euploidy, aneuploid karyotypes will not be a multiple of the haploid number. In humans, examples of aneuploidy include having a single extra chromosome (as in Down syndrome , where affected individuals have three copies of chromosome 21) or missing a chromosome (as in Turner syndrome , where affected individuals have only one sex chromosome). Aneuploid karyotypes are given names with
9954-476: The birthday of President Manuel L. Quezon. The reason for the move being given that the original celebration was a period "outside of the school year, thereby precluding the participation of schools in its celebration". In 1988, President Corazon Aquino signed Proclamation No. 19, reaffirming the celebration every August 13 to 19. In 1997, the celebration was extended from a week to a month by Proclamation 1041 of July 15 signed by President Fidel V. Ramos . It
10080-409: The bulk of the abaca harvests were exported to the United States" for use in military initiatives. In the early 1900s, a train running from Danao to Argao would transport Philippine abacá from the plantations to Cebu City for export. The railway system was destroyed during World War II; the abaca continues to be transported to Cebu by road. Outside the Philippines, abacá was first cultivated on
10206-747: The chair of the Institute and as the representative of Samar-Leyte-Visayans , the Institute's members were composed of Santiago A. Fonacier (representing the Ilokano-speaking regions ), Filemon Sotto (the Cebu-Visayans ), Casimiro Perfecto (the Bikolanos ), Felix S. Sales Rodriguez (the Panay-Visayans ), Hadji Butu (the languages of Muslim Filipinos ), and Cecilio Lopez (the Tagalogs ). The Institute of National Language adopted
10332-437: The chromosome constitution. Dihaploids (which are diploid) are important for selective breeding of tetraploid crop plants (notably potatoes), because selection is faster with diploids than with tetraploids. Tetraploids can be reconstituted from the diploids, for example by somatic fusion. The term "dihaploid" was coined by Bender to combine in one word the number of genome copies (diploid) and their origin (haploid). The term
10458-453: The chromosome number is generally reduced only by the specialized process of meiosis, the somatic cells of the body inherit and maintain the chromosome number of the zygote by mitosis. However, in many situations somatic cells double their copy number by means of endoreduplication as an aspect of cellular differentiation . For example, the hearts of two-year-old human children contain 85% diploid and 15% tetraploid nuclei, but by 12 years of age
10584-499: The condition is called triploid syndrome . In unicellular organisms the ploidy nutrient limitation hypothesis suggests that nutrient limitation should encourage haploidy in preference to higher ploidies. This hypothesis is due to the higher surface-to-volume ratio of haploids, which eases nutrient uptake, thereby increasing the internal nutrient-to-demand ratio. Mable 2001 finds Saccharomyces cerevisiae to be somewhat inconsistent with this hypothesis however, as haploid growth
10710-427: The cultivated varieties of Catanduanes even more competitive in local and international markets. This results in the optimum production of the island which had a consistent highest production throughout the archipelago. Europeans first came into contact with Abacá fibre when Ferdinand Magellan landed in the Philippines in 1521, as the natives were already cultivating it and utilizing it in bulk for textiles. Throughout
10836-461: The delegates who were non-Tagalogs were even in favor of scrapping the idea of a "national language" altogether. A compromise was reached and the wording on the 1973 constitution made no mention of dropping the national language Pilipino or made any mention of Tagalog. Instead, the 1973 Constitution , in both its original form and as amended in 1976, designated English and Pilipino as official languages and provided for development and formal adoption of
10962-423: The diploid stage are under less efficient natural selection than those genes expressed in the haploid stage is referred to as the “masking theory”. Evidence in support of this masking theory has been reported in studies of the single-celled yeast Saccharomyces cerevisiae . In further support of the masking theory, evidence of strong purifying selection in haploid tissue-specific genes has been reported for
11088-476: The form of the baro (also barú or bayú , literally "shirt" or "clothing"), a simple collar-less shirt or jacket with close-fitting long sleeves worn by both men and women in most ethnic groups in the pre-colonial Philippines . These were paired with wraparound sarong-like skirts (for both men and women), close-fitting pants, or loincloths ( bahag ). During the Spanish colonial era , abacá cloth became known as medriñaque in Spanish (apparently derived from
11214-425: The gametes produced by the tetraploid organism in the example above, since these gametes are numerically diploid. The term monoploid is often used as a less ambiguous way to describe a single set of chromosomes; by this second definition, haploid and monoploid are identical and can be used interchangeably. Gametes ( sperm and ova ) are haploid cells. The haploid gametes produced by most organisms combine to form
11340-419: The haploid number n = 21). The gametes are haploid for their own species, but triploid, with three sets of chromosomes, by comparison to a probable evolutionary ancestor, einkorn wheat . Tetraploidy (four sets of chromosomes, 2 n = 4 x ) is common in many plant species, and also occurs in amphibians , reptiles , and insects . For example, species of Xenopus (African toads) form
11466-532: The hybridization of two separate species. In plants, this probably most often occurs from the pairing of meiotically unreduced gametes , and not by diploid–diploid hybridization followed by chromosome doubling. The so-called Brassica triangle is an example of allopolyploidy, where three different parent species have hybridized in all possible pair combinations to produce three new species. Polyploidy occurs commonly in plants, but rarely in animals. Even in diploid organisms, many somatic cells are polyploid due to
11592-583: The idea that haploidy is selected by harsher conditions – is cast into doubt by these results. Older WGDs have also been investigated. Only as recently as 2015 was the ancient whole genome duplication in Baker's yeast proven to be allopolyploid , by Marcet-Houben and Gabaldón 2015. It still remains to be explained why there are not more polyploid events in fungi, and the place of neopolyploidy and mesopolyploidy in fungal history . The concept that those genes of an organism that are expressed exclusively in
11718-551: The latest edition being published in 2013 in Manila. Spanish served in an official capacity as language of the government during the Spanish period. Spanish played a significant role in unifying the Philippines, a country made up of over 7,000 islands with a multitude of ethnicities, languages, and cultures. Before Spanish rule, the archipelago was not a unified nation, but rather a collection of independent kingdoms, sultanates, and tribes, each with its own language and customs. During
11844-468: The latter national. This is similar to the comparison between Castilian and Spanish , or Mandarin and Chinese . Political designations aside, Tagalog and Filipino are linguistically the same, sharing, among other things, the same grammatical structure. On May 23, 2007, Ricardo Maria Nolasco, KWF chair and a linguistics expert, acknowledged in a keynote speech during the NAKEM Conference at
11970-408: The lexicon of the country's other languages, something toward which the commission was working. On August 24, 2007, Nolasco elaborated further on the relationship between Tagalog and Filipino in a separate article, as follows: Are "Tagalog," "Pilipino" and "Filipino" different languages? No, they are mutually intelligible varieties, and therefore belong to one language. According to the KWF, Filipino
12096-425: The main source of textile fibers used for clothing in the pre-colonial Philippines. Abacá cloth was often compared to calico in terms of texture and was a major trade commodity in the pre-colonial maritime trade and the Spanish colonial era . There are multiple traditional types and names of abaca cloth among the different ethnic groups of the Philippines . Undyed plain abacá cloth, woven from fine fibers of abaca,
12222-428: The modern abacá are thought to have originated from the eastern Philippines, where there is significant rainfall throughout the year. Wild varieties of abacá can still be found in the interior forests of the island province of Catanduanes , away from cultivated areas. Today, Catanduanes has many other modern kinds of abacá which are more competitive. For many years, breeders from various research institutions have made
12348-455: The monoploid number and haploid number are equal; in humans, both are equal to 23. When a human germ cell undergoes meiosis, the diploid 46 chromosome complement is split in half to form haploid gametes. After fusion of a male and a female gamete (each containing 1 set of 23 chromosomes) during fertilization , the resulting zygote again has the full complement of 46 chromosomes: 2 sets of 23 chromosomes. Euploidy and aneuploidy describe having
12474-418: The most generic sense, haploid refers to having the number of sets of chromosomes normally found in a gamete . Because two gametes necessarily combine during sexual reproduction to form a single zygote from which somatic cells are generated, healthy gametes always possess exactly half the number of sets of chromosomes found in the somatic cells, and therefore "haploid" in this sense refers to having exactly half
12600-431: The name did not, however, result in universal acceptance among non- Tagalogs , especially Cebuanos who had previously not accepted the 1937 selection. The 1960s saw the rise of the purist movement where new words were being coined to replace loanwords. This era of "purism" by the SWP sparked criticisms by a number of persons. Two counter-movements emerged during this period of "purism": one campaigning against Tagalog and
12726-565: The number of chromosomes may have originated in this way, this is no longer clear, and the monoploid number is regarded as the same as the haploid number. Thus in humans, x = n = 23. Diploid cells have two homologous copies of each chromosome , usually one from the mother and one from the father . All or nearly all mammals are diploid organisms. The suspected tetraploid (possessing four-chromosome sets) plains viscacha rat ( Tympanoctomys barrerae ) and golden viscacha rat ( Pipanacoctomys aureus ) have been regarded as
12852-406: The number of sets of chromosomes found in a somatic cell. By this definition, an organism whose gametic cells contain a single copy of each chromosome (one set of chromosomes) may be considered haploid while the somatic cells, containing two copies of each chromosome (two sets of chromosomes), are diploid. This scheme of diploid somatic cells and haploid gametes is widely used in the animal kingdom and
12978-401: The official name of Tagalog, or even a synonym of it. Today's Filipino language is best described as "Tagalog-based". The language is usually called Tagalog within the Philippines and among Filipinos to differentiate it from other Philippine languages, but it has also come to be known as Filipino to differentiate it from the languages of other countries; the former implies a regional origin,
13104-694: The only known exceptions (as of 2004). However, some genetic studies have rejected any polyploidism in mammals as unlikely, and suggest that amplification and dispersion of repetitive sequences best explain the large genome size of these two rodents. All normal diploid individuals have some small fraction of cells that display polyploidy . Human diploid cells have 46 chromosomes (the somatic number, 2n ) and human haploid gametes (egg and sperm) have 23 chromosomes ( n ). Retroviruses that contain two copies of their RNA genome in each viral particle are also said to be diploid. Examples include human foamy virus , human T-lymphotropic virus , and HIV . Polyploidy
13230-409: The organism as it now reproduces. Common wheat ( Triticum aestivum ) is an organism in which x and n differ. Each plant has a total of six sets of chromosomes (with two sets likely having been obtained from each of three different diploid species that are its distant ancestors). The somatic cells are hexaploid, 2 n = 6 x = 42 (where the monoploid number x = 7 and
13356-556: The other campaigning for more inclusiveness in the national language. In 1963, Negros Occidental congressman Innocencio V. Ferrer took a case reaching the Supreme Court questioning the constitutionality of the choice of Tagalog as the basis of the national language (a case ruled in favor of the national language in 1970). Accusing the national language as simply being Tagalog and lacking any substantial input from other Philippine languages, Congressman Geruncio Lacuesta eventually led
13482-1005: The peoples of Member States themselves and among the peoples of territories under their jurisdiction. ang Pangkalahatáng Kapulungán ay nagpapahayág ng PANGKALAHATÁNG PAGPAPAHAYÁG NA ITÓ NG MGÁ KARAPATÁN NG TÁO bílang pangkalahatáng pamantáyang maisasagawâ pára sa lahát ng táo at bansâ, sa layúning ang báwat táo at báwat galamáy ng lipúnan, na láging nása ísip ang Pahayág na itó, ay magsíkap sa pamamagítan ng pagtutúrò at edukasyón na maitagúyod ang paggálang sa mgá karapatán at kalayáang itó at sa pamamagítan ng mgá hakbáng na pagsúlong na pambansâ at pandaigdíg, ay makamtán ang pangkalahatán at mabísang pagkilála at pagtalíma sa mgá itó, magíng ng mgá mamamayán ng mgá Kasáping Estádo at ng mgá mamamayán ng mgá teritóryo na nása ilálim ng kaniláng nasasakúpan. Diploid Virtually all sexually reproducing organisms are made up of somatic cells that are diploid or greater, but ploidy level may vary widely between different organisms, between different tissues within
13608-403: The plant Scots Pine . The common potato ( Solanum tuberosum ) is an example of a tetraploid organism, carrying four sets of chromosomes. During sexual reproduction, each potato plant inherits two sets of 12 chromosomes from the pollen parent, and two sets of 12 chromosomes from the ovule parent. The four sets combined provide a full complement of 48 chromosomes. The haploid number (half of 48)
13734-420: The plant fruits, grows to about 2–3 inches (5.1–7.6 cm) in length and 1 inch (2.5 cm) in diameter. It has black turbinate seeds that are 0.167 inches (0.42 cm) in diameter. The abacá plant belongs to the banana family, Musaceae ; it resembles the closely related wild seeded bananas, Musa acuminata and Musa balbisiana . Its scientific name is Musa textilis . Within the genus Musa , it
13860-426: The plant is mature, the flower stalk grows up inside the pseudostem. The male flower has five petals, each about 1.5 inches (3.8 cm) long. The leaf sheaths contain the valuable fiber. After harvesting, the coarse fibers range in length from 6–12 feet (180–370 cm) long. They are composed primarily of cellulose , lignin , and pectin . The fruit, which is inedible and is rarely seen as harvesting occurs before
13986-525: The primary language for education, governance, and law. At present, Spanish was designated an optional and voluntary language under the 1987 Constitution, along with Arabic. While Spanish and English were considered "official languages" during the American colonial period, there existed no "national language" initially. Article XIII, section 3 of the 1935 constitution establishing the Commonwealth of
14112-400: The proportions become approximately equal, and adults examined contained 27% diploid, 71% tetraploid and 2% octaploid nuclei. There is continued study and debate regarding the fitness advantages or disadvantages conferred by different ploidy levels. A study comparing the karyotypes of endangered or invasive plants with those of their relatives found that being polyploid as opposed to diploid
14238-684: The rest of the body being diploid (as in the mammalian liver ). For many organisms, especially plants and fungi, changes in ploidy level between generations are major drivers of speciation . In mammals and birds, ploidy changes are typically fatal. There is, however, evidence of polyploidy in organisms now considered to be diploid, suggesting that polyploidy has contributed to evolutionary diversification in plants and animals through successive rounds of polyploidization and rediploidization. Humans are diploid organisms, normally carrying two complete sets of chromosomes in their somatic cells: one copy of paternal and maternal chromosomes, respectively, in each of
14364-501: The same name was written by Czech Jesuit missionary Paul Klein (known locally as Pablo Clain) at the beginning of the 18th century. Klein spoke Tagalog and used it actively in several of his books. He wrote a dictionary, which he later passed to Francisco Jansens and José Hernández. Further compilation of his substantial work was prepared by Juan de Noceda and Pedro de Sanlúcar and published as Vocabulario de la lengua tagala in Manila in 1754 and then repeatedly re-edited, with
14490-450: The same organism . Though polyploidy in humans is not viable, mixoploidy has been found in live adults and children. There are two types: diploid-triploid mixoploidy, in which some cells have 46 chromosomes and some have 69, and diploid-tetraploid mixoploidy, in which some cells have 46 and some have 92 chromosomes. It is a major topic of cytology. Dihaploid and polyhaploid cells are formed by haploidisation of polyploids, i.e., by halving
14616-462: The same organism, and at different stages in an organism's life cycle. Half of all known plant genera contain polyploid species, and about two-thirds of all grasses are polyploid. Many animals are uniformly diploid, though polyploidy is common in invertebrates, reptiles, and amphibians. In some species, ploidy varies between individuals of the same species (as in the social insects ), and in others entire tissues and organ systems may be polyploid despite
14742-398: The same personal pronouns (siya, ako, niya, kanila, etc.); the same demonstrative pronouns (ito, iyan, doon, etc.); the same linkers (na, at and ay); the same particles (na and pa); and the same verbal affixes -in, -an, i- and -um-. In short, same grammar, same language. In connection with the use of Filipino, or specifically the promotion of the national language, the related term Tagalista
14868-500: The same species or at different stages of the life cycle . In some insects it differs by caste . In humans, only the gametes are haploid, but in many of the social insects , including ants , bees , and termites , males develop from unfertilized eggs, making them haploid for their entire lives, even as adults. In the Australian bulldog ant, Myrmecia pilosula , a haplodiploid species, haploid individuals of this species have
14994-636: The same year, Quezon proclaimed Tagalog as the basis of the Wikang Pambansâ (National Language) giving the following factors: On June 7, 1940, the Philippine National Assembly passed Commonwealth Act No. 570 declaring that the Filipino national language would be considered an official language effective July 4, 1946 (coinciding with the country's expected date of independence from the United States). That same year,
15120-547: The suffix -somy (rather than -ploidy , used for euploid karyotypes), such as trisomy and monosomy . Homoploid means "at the same ploidy level", i.e. having the same number of homologous chromosomes . For example, homoploid hybridization is hybridization where the offspring have the same ploidy level as the two parental species. This contrasts with a common situation in plants where chromosome doubling accompanies or occurs soon after hybridization. Similarly, homoploid speciation contrasts with polyploid speciation . Zygoidy
15246-436: The terms haploid and diploid in 1905. Some authors suggest that Strasburger based the terms on August Weismann 's conception of the id (or germ plasm ), hence haplo- id and diplo- id . The two terms were brought into the English language from German through William Henry Lang 's 1908 translation of a 1906 textbook by Strasburger and colleagues. The term haploid is used with two distinct but related definitions. In
15372-518: The use of Filipino as a medium of official communication and as language of instruction in the educational system. and: The regional languages are the auxiliary official languages in the regions and shall serve as auxiliary media of instruction therein. Section 17(d) of Executive Order 117 of January 30, 1987 renamed the Institute of National Language as Institute of Philippine Languages . Republic Act No. 7104, approved on August 14, 1991, created
15498-409: Was attributed to molecular variance within the population, and only 5% of the molecular variance to variation among populations. Genetic analysis by Unweighted Pair Group Method with Arithmetic Mean (UPGMA) revealed several clusters irrespective of geographical origin. Before synthetic textiles came into use, M. textilis was a major source of high quality fiber: soft, silky and fine. Ancestors of
15624-443: Was generally used by the ruling classes and the merchants from the states and various cultures in the Philippine archipelago for international communication as part of maritime Southeast Asia. In fact, Filipinos first interacted with the Spaniards using the Malay language. In addition to this, 16th-century chroniclers of the time noted that the kings and lords in the islands usually spoke around five languages. Spanish intrusion into
15750-463: Was reported that three Malolos City regional trial courts in Bulacan decided to use Filipino, instead of English , in order to promote the national language. Twelve stenographers from Branches 6, 80 and 81, as model courts, had undergone training at Marcelo H. del Pilar College of Law of Bulacan State University following a directive from the Supreme Court of the Philippines . De la Rama said it
15876-416: Was the dream of Chief Justice Reynato Puno to implement the program in other areas such as Laguna , Cavite , Quezon , Aurora , Nueva Ecija , Batangas , Rizal , and Metro Manila , all of which mentioned are natively Tagalog-speaking. Since 1997, a month-long celebration of the national language occurs during August, known in Filipino as Buwan ng Wika (Language Month). Previously, this lasted only
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