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98-555: Aceh Besar Regency produces cloves , nutmeg and palm oil and rice , and also small quantities of maize , cassava , sweet potatoes and beans . Aceh Besar Regency contains several museums. The Museum dan Rumoh Aceh is the State Museum, located in Banda Aceh . The museum's main building is built in the style of a traditional Acehnese house. It was installed by Dutch Governor Van Swart in 1915. Museum Ali Hasymi contains

196-459: A bloom or blossom , is the reproductive structure found in flowering plants (plants of the division Angiospermae ). Flowers consist of a combination of vegetative organs – sepals that enclose and protect the developing flower. These petals attract pollinators, and reproductive organs that produce gametophytes , which in flowering plants produce gametes . The male gametophytes, which produce sperm, are enclosed within pollen grains produced in

294-665: A boron -based wood preservative. Cloves can be used to make a fragrant pomander when combined with an orange. When given as a gift in Victorian England, such a pomander indicated warmth of feeling. The use of clove for any medicinal purpose has not been approved by the US Food and Drug Administration , and its use may cause adverse effects if taken orally by people with liver disease , blood clotting and immune system disorders, or food allergies . Cloves are used in traditional medicine as an essential oil , which

392-526: A diploid (two copies of each chromosome ) cell. Whereas in fertilization only plasmogamy, or the fusion of the whole sex cells, results, in Angiosperms (flowering plants) a process known as double fertilization, which involves both karyogamy and plasmogamy, occurs. In double fertilization the second sperm cell subsequently also enters the synergid and fuses with the two polar nuclei of the central cell. Since all three nuclei are haploid , they result in

490-411: A filament , or stalk. The anther contains microsporocytes which become pollen , the male gametophyte , after undergoing meiosis . Although they exhibit the widest variation among floral organs, the androecium is usually confined just to one whorl and to two whorls only in rare cases. Stamens range in number, size, shape, orientation, and in their point of connection to the flower. In general, there

588-459: A spice , flavoring , or fragrance in consumer products , such as toothpaste , soaps, or cosmetics . Cloves are available throughout the year owing to different harvest seasons across various countries. The word clove , first used in English in the 15th century , derives via Middle English clow of gilofer , Anglo-French clowes de gilofre and Old French clou de girofle , from

686-411: A broad base, stomata and chlorophyll and may have stipules . Sepals are often waxy and tough, and grow quickly to protect the flower as it develops. They may be deciduous , but will more commonly grow on to assist in fruit dispersal. If the calyx is fused it is called gamosepalous. The petals , or corolla, are almost or completely fiberless leaf-like structures that form the innermost whorl of

784-588: A common element in spice blends (as part of the Malay rempah empat beradik –"four sibling spices"– besides cinnamon, cardamom and star anise for example ), including pumpkin pie spice and speculaas spices. In Mexican cuisine , cloves are best known as clavos de olor , and often accompany cumin and cinnamon . They are also used in Peruvian cuisine, in a wide variety of dishes such as carapulcra and arroz con leche . A major component of clove's taste

882-404: A different flower of the same plant, but others have mechanisms to prevent self-pollination and rely on cross-pollination , when pollen is transferred from the anther of one flower to the stigma of another flower on a different individual of the same species. Self-pollination happens in flowers where the stamen and carpel mature at the same time, and are positioned so that the pollen can land on

980-418: A different plant of the same species. Because the genetic make-up of the sperm contained within the pollen from the other plant is different, their combination will result in a new, genetically distinct, plant, through the process of sexual reproduction . Since each new plant is genetically distinct, the different plants show variation in their physiological and structural adaptations and so the population as

1078-425: A food source for pollinators. In this way, many flowering plants have co-evolved with pollinators to be mutually dependent on services they provide to one another—in the plant's case, a means of reproduction; in the pollinator's case, a source of food. When pollen from the anther of a flower is deposited on the stigma , this is called pollination. Some flowers may self-pollinate , producing seed using pollen from

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1176-432: A large endosperm nucleus which is triploid . Following the formation of zygote it begins to grow through nuclear and cellular divisions, called mitosis , eventually becoming a small group of cells. One section of it becomes the embryo , while the other becomes the suspensor; a structure which forces the embryo into the endosperm and is later undetectable. Two small primordia also form at this time, that later become

1274-1340: A mate). In pursuing this attractant from many flowers of the same species, the pollinator transfers pollen to the stigmas—arranged with equally pointed precision—of all of the flowers it visits. Many flowers rely on simple proximity between flower parts to ensure pollination, while others have elaborate designs to ensure pollination and prevent self-pollination . Flowers use animals including: insects ( entomophily ), birds ( ornithophily ), bats ( chiropterophily ), lizards, and even snails and slugs ( malacophilae ). Plants cannot move from one location to another, thus many flowers have evolved to attract animals to transfer pollen between individuals in dispersed populations. Most commonly, flowers are insect-pollinated, known as entomophilous ; literally "insect-loving" in Greek. To attract these insects flowers commonly have glands called nectaries on various parts that attract animals looking for nutritious nectar . Some flowers have glands called elaiophores , which produce oils rather than nectar. Birds and bees have color vision , enabling them to seek out colorful flowers. Some flowers have patterns, called nectar guides , that show pollinators where to look for nectar; they may be visible only under ultraviolet light, which

1372-458: A noticeable scent. Because of this, plants typically have many thousands of tiny flowers which have comparatively large, feathery stigmas; to increase the chance of pollen being received. Whereas the pollen of entomophilous flowers is usually large, sticky, and rich in protein (to act as a "reward" for pollinators), anemophilous flower pollen is typically small-grained, very light, smooth, and of little nutritional value to insects . In order for

1470-572: A pale hue, gradually turn green, then transition to a bright red when ready for harvest. Cloves are harvested at 1.5–2 centimetres ( 5 ⁄ 8 – 3 ⁄ 4  in) long, and consist of a long calyx that terminates in four spreading sepals , and four unopened petals that form a small central ball. Clove stalks are slender stems of the inflorescence axis that show opposite decussate branching. Externally, they are brownish, rough, and irregularly wrinkled longitudinally with short fracture and dry, woody texture. Mother cloves (anthophylli) are

1568-470: A plant can interpret important endogenous and environmental cues such as changes in levels of plant hormones and seasonable temperature and photoperiod changes. Many perennial and most biennial plants require vernalization to flower. The molecular interpretation of these signals is through the transmission of a complex signal known as florigen , which involves a variety of genes , including Constans, Flowering Locus C, and Flowering Locus T. Florigen

1666-404: A sonar-reflecting petal above its flowers, which helps the bat find them, and one species, the cactus Espostoa frutescens , has flowers that are surrounded by an area of sound-absorbent and woolly hairs called the cephalium, which absorbs the bat's ultrasound instead. Flowers are also specialized in shape and have an arrangement of the stamens that ensures that pollen grains are transferred to

1764-407: A third in 1944. Knuth named his two groups 'Hyphydrogamy' and the more common 'Ephydrogamy'. In hyphydrogamy pollination occurs below the surface of the water and so the pollen grains are typically negatively buoyant . For marine plants that exhibit this method, the stigmas are usually stiff, while freshwater species have small and feathery stigmas. In ephydrogamy pollination occurs on the surface of

1862-435: A whole is better prepared for an adverse occurrence in the environment. Cross-pollination, therefore, increases the survival of the species and is usually preferred by flowers for this reason. The principal adaptive function of flowers is the promotion of cross-pollination or outcrossing, a process that allows the masking of deleterious mutations in the genome of progeny. The masking effect of outcrossing sexual reproduction

1960-416: Is a loss of B gene function, mutant flowers are produced with sepals in the first whorl as usual, but also in the second whorl instead of the normal petal formation. In the third whorl, the lack of the B function but the presence of the C function mimics the fourth whorl, leading to the formation of carpels also in the third whorl. The principal purpose of a flower is the reproduction of the individual and

2058-422: Is a much rarer method, occurring in only around 2% of abiotically pollinated flowers. Common examples of this include Calitriche autumnalis , Vallisneria spiralis and some sea-grasses . One characteristic which most species in this group share is a lack of an exine , or protective layer, around the pollen grain. Paul Knuth identified two types of hydrophilous pollination in 1906 and Ernst Schwarzenbach added

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2156-417: Is a way to represent the structure of a flower using specific letters, numbers, and symbols, presenting substantial information about the flower in a compact form. It can represent a taxon , usually giving ranges of the numbers of different organs, or particular species. Floral formulae have been developed in the early 19th century and their use has declined since. Prenner et al. (2010) devised an extension of

2254-404: Is an example of coevolution , as the flower and pollinator have developed together over a long period to match each other's needs. This close relationship compounds the negative effects of extinction , however, since the extinction of either member in such a relationship would almost certainly mean the extinction of the other member as well. Flowers that use abiotic, or non-living, vectors use

2352-427: Is because it is directly connected to the method of seed dispersal; that being the purpose of fruit - to encourage or enable the seed's dispersal and protect the seed while doing so. Following the pollination of a flower, fertilization, and finally the development of a seed and fruit, a mechanism is typically used to disperse the fruit away from the plant. In Angiosperms (flowering plants) seeds are dispersed away from

2450-462: Is cultivated in three separate ways, a monoculture , agricultural parklands, and agroforestry systems. Archaeologist Giorgio Buccellati found cloves in Terqa , Syria, in a burned-down house which was dated to 1720 BC during the kingdom of Khana . This was the first evidence of cloves being used in the west before Roman times. The discovery was first reported in 1978. They reached Rome by

2548-440: Is divided administratively into twenty-three districts ( Indonesian : kecamatan ), which comprise 604 villages (Indonesian: gampong ). The areas of the districts and their populations at the 2010 Census and the 2020 Census, together with the official estimates as at mid 2023, are listed below. These are grouped below into three geographical sections for convenience, which have no administrative significance. The table also includes

2646-622: Is from the Middle English flour , which referred to both the ground grain and the reproductive structure in plants, before splitting off in the 17th century. It comes originally from the Latin name of the Italian goddess of flowers, Flora . The early word for flower in English was blossom , though it now refers to flowers only of fruit trees . The morphology of a flower, or its form and structure, can be considered in two parts:

2744-541: Is highly reduced or absent). The stem or stalk subtending a flower, or an inflorescence of flowers, is called a peduncle . If a peduncle supports more than one flower, the stems connecting each flower to the main axis are called pedicels . The apex of a flowering stem forms a terminal swelling which is called the torus or receptacle. In the majority of species, individual flowers have both carpels and stamens. These flowers are described by botanists as being perfect, bisexual, or hermaphrodite . In some species of plants,

2842-487: Is imparted by the chemical eugenol , and the quantity of the spice required is typically small. It pairs well with cinnamon, allspice , vanilla , red wine , basil , onion , citrus peel , star anise , and peppercorns . It is often added to betel quids to enhance aroma while chewing. The spice is used in a type of cigarette called kretek in Indonesia. Clove cigarettes were smoked throughout Europe, Asia, and

2940-494: Is known as "genetic complementation". This beneficial effect of outcrossing on progeny is also recognized as hybrid vigour or heterosis. Once outcrossing is established due to the benefits of genetic complementation, subsequent switching to inbreeding becomes disadvantageous because it allows the expression of the previously masked deleterious recessive mutations, usually referred to as inbreeding depression. Charles Darwin in his 1889 book The Effects of Cross and Self-Fertilization in

3038-491: Is located about 20 km south of Medan . Indra Puri was a Hindu kingdom and it was a place of worship before Islam arrived. Later, Sultan Iskandar Muda introduced Islam to the public. After the people embraced Islam, the previous temple was converted into a mosque. The mosque site covers an area of 33,875 m, and is located approximately 150 metres from the edge of the Krueng Aceh River . Indra Patra Fortress

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3136-407: Is often present in a root canal treatment failure. Studies to determine its effectiveness for fever reduction, as a mosquito repellent, and to prevent premature ejaculation have been inconclusive. It remains unproven whether blood sugar levels are reduced by cloves or clove oil. The essential oil may be used in aromatherapy . Until the colonial era , cloves only grew on a few islands in

3234-408: Is only one type of stamen, but there are plant species where the flowers have two types; a "normal" one and one with anthers that produce sterile pollen meant to attract pollinators. The gynoecium , or the carpels , is the female part of the flower found on the innermost whorl. Each carpel consists of a stigma , which receives pollen, a style , which acts as a stalk, and an ovary , which contains

3332-443: Is preferred because it allows for genetic variation , which contributes to the survival of the species. Many flowers depend on external factors for pollination, such as the wind, water, animals, and especially insects . Larger animals such as birds, bats, and even some pygmy possums , however, can also be employed. To accomplish this, flowers have specific designs which encourage the transfer of pollen from one plant to another of

3430-429: Is produced in the leaves in reproductively favorable conditions and acts in buds and growing tips to induce several different physiological and morphological changes. The first step of the transition is the transformation of the vegetative stem primordia into floral primordia. This occurs as biochemical changes take place to change the cellular differentiation of leaf, bud and stem tissues into tissue that will grow into

3528-612: Is situated 19 km from Banda Aceh at Krueng Raya . According to history it was built under the Indra Puri Hindu kingdom, although some say that the fort was built during the Sultanate of Aceh Darussalam in an effort to resist the Portuguese. The fort had a very important function at the time of Sultan Iskandar Muda. The Tomb of Admiral Malahayati is located about 32 km from the city of Banda Aceh. The regency

3626-418: Is typically another part. In some families, such as the grasses , the petals are greatly reduced; in many species, the sepals are colorful and petal-like. Other flowers have modified petal-like stamens; the double flowers of peonies and roses are mostly petaloid stamens. Many flowers have symmetry. When the perianth is bisected through the central axis from any point and symmetrical halves are produced,

3724-411: Is used as an anodyne ( analgesic ) mainly for dental emergencies and other disorders. There is evidence that clove oil containing eugenol is effective for toothache pain and other types of pain, and one review reported the efficacy of eugenol combined with zinc oxide as an analgesic for alveolar osteitis . Clove essential oil may prevent the growth of Enterococcus faecalis bacteria which

3822-574: Is visible to bees and some other insects. Flowers also attract pollinators by scent , though not all flower scents are appealing to humans; several flowers are pollinated by insects that are attracted to rotten flesh and have flowers that smell like dead animals. These are often called carrion flowers , including plants in the genus Rafflesia , and the titan arum . Flowers pollinated by night visitors, including bats and moths, are likely to concentrate on scent to attract pollinators and so most such flowers are white. Some plants pollinated by bats have

3920-732: The Moluccas (historically called the Spice Islands ), including Bacan , Makian , Moti , Ternate , and Tidore . Cloves were first traded by the Austronesian peoples in the Austronesian maritime trade network (which began around 1500 BC, later becoming the Maritime Silk Road and part of the Spice Trade ). The first notable example of modern clove farming developed on the east coast of Madagascar , and

4018-903: The Yuan dynasty (1271 to 1368 AD) Chinese merchants began sending ships directly to the Moluccas to trade for cloves, and other spices. In the Western Classical literature cloves are mentioned by Pliny the Elders' Natural History . Dioscorides mentions cloves in his book De materia medica . The Liber Pontifcalis records an endowment made by Passinopolis under Pope Sylvester I . This endowment included an Egyptian estate, its annual revenues, 150 libra (around 50 kg or 108 lb) of cloves, and other amounts of spices and papyrus. Cosmas Indicopleustis in his book Topographia Christiana outlined his travels to Sri Lanka, and recounted that

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4116-400: The anthers . The female gametophytes are contained within the ovules produced in the ovary . Most flowering plants depend on animals, such as bees, moths, and butterflies, to transfer their pollen between different flowers, and have evolved to attract these pollinators by various strategies, including brightly colored, conspicuous petals, attractive scents, and the production of nectar ,

4214-414: The calyx , corolla , androecium , and gynoecium . Together the calyx and corolla make up the non-reproductive part of the flower called the perianth , and in some cases may not be differentiated. If this is the case, then they are described as tepals . The sepals , collectively called the calyx, are modified leaves that occur on the outermost whorl of the flower. They are leaf-like, in that they have

4312-499: The cotyledon , which is used as an energy store. Plants which grow out one of these primordia are called monocotyledons , while those that grow out two are dicotyledons . The next stage is called the Torpedo stage and involves the growth of several key structures, including: the radicle (embryotic root), the epicotyl (embryotic stem), and the hypocotyl , (the root/shoot junction). In the final step vascular tissue develops around

4410-401: The flavonoids eugenin , kaempferol , rhamnetin , and eugenitin , tri terpenoids such as oleanolic acid , stigmasterol , and campesterol and several sesquiterpenes . Although eugenol has not been classified for its potential toxicity , it was shown to be toxic to test organisms in concentrations of 50, 75, and 100 mg per liter. Flower A flower , also known as

4508-439: The gizzard of animals or even to germinate better after passing through them. They can be eaten by birds ( ornithochory) , bats ( chiropterochory) , rodents , primates, ants ( myrmecochory ), non-bird sauropsids ( saurochory) , mammals in general (mammaliochory) , and even fish . Typically their fruit are fleshy, have a high nutritional value, and may have chemical attractants as an additional "reward" for dispersers. This

4606-518: The wind or, much less commonly, water , to move pollen from one flower to the next. In wind-dispersed ( anemophilous ) species, the tiny pollen grains are carried, sometimes many thousands of kilometers, by the wind to other flowers. Common examples include the grasses , birch trees , along with many other species in the order Fagales , ragweeds , and many sedges . They do not need to attract pollinators and therefore tend not to grow large, showy, or colorful flowers, and do not have nectaries, nor

4704-478: The 2004 tsunami struck Bunta island, 45 minutes by traditional motorised boat from Ujong Pancu Village, Peukan Bada, there are no longer any inhabitants living there. Nowadays some people use the island as a coconut plantation, but they live in Banda Aceh. As the tsunami washed out all of the living creatures there, nowadays there are no squirrels, monkeys or snakes on the island. The island has been popularized by

4802-888: The Indians said that cloves, among other products, came in from unspecified places along sea trade routes. Cloves were also present in records in China , Sri Lanka , Southern India , Persia , and Oman by around the third century to second century BC. These mentions of "cloves" reported in China, South Asia, and the Middle East come from before the establishment of Southeast Asian maritime trade. But all of these are misidentifications that referred to other plants (like cassia buds, cinnamon , or nutmeg ); or are imports from Maritime Southeast Asia mistakenly identified as being natively produced in these regions. Archaeologists recovered

4900-432: The Latin word clavus "nail". The related English word gillyflower , originally meaning "clove", derives via said Old French girofle and Latin caryophyllon , from the Greek karyophyllon "clove", literally "nut leaf". The clove tree is an evergreen that grows up to 8–12 metres (26–39 ft) tall, with large leaves and crimson flowers grouped in terminal clusters. The flower buds initially have

4998-461: The Moluccas, and the trade in cloves was beyond the limited policing powers of the corporation". One clove tree named Afo that experts believe is the oldest in the world on Ternate may be 350–400 years old. Tourists are told that seedlings from this very tree were stolen by a Frenchman named Pierre Poivre in 1770, transferred to the Isle de France ( Mauritius ), and then later to Zanzibar , which

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5096-682: The Officialdom of the Han Dynasty, dating to around 200 BC). The book states a rule that ministers should suck cloves to sweeten their breath before speaking to the emperor. From Chinese records during the Song Dynasty (960 to 1279 AD) cloves were primarily imported by private ventures, called Merchant Shipping Offices, who bought goods from middlemen in the Austronesian polities of Java , Srivijaya , Champa , and Butuan . During

5194-564: The United States. Clove cigarettes are currently classified in the United States as cigars , the result of a ban on flavored cigarettes in September 2009. Clove essential oil may be used to inhibit mold growth on various types of foods. In addition to these non-culinary uses of clove, it can be used to protect wood in a system for cultural heritage conservation, and showed the efficacy of clove essential oil to be higher than

5292-470: The Vegetable Kingdom at the beginning of chapter XII noted, "The first and most important of the conclusions which may be drawn from the observations given in this volume, is that generally cross-fertilisation is beneficial and self-fertilisation often injurious, at least with the plants on which I experimented." Self-pollination is the pollination of the carpel of a flower by pollen from either

5390-490: The apical meristem, which becomes a whorl of sepals. In the second whorl, both A and B genes are expressed, leading to the formation of petals. In the third whorl, B and C genes interact to form stamens and in the center of the flower C genes alone give rise to carpels. The model is based upon studies of aberrant flowers and mutations in Arabidopsis thaliana and the snapdragon, Antirrhinum majus . For example, when there

5488-582: The bodies of the pollinator when it lands in search of its attractant. Other flowers use mimicry or pseudocopulation to attract pollinators. Many orchids, for example, produce flowers resembling female bees or wasps in color, shape, and scent. Males move from one flower to the next in search of a mate, pollinating the flowers. Many flowers have close relationships with one or a few specific pollinating organisms. Many flowers, for example, attract only one specific species of insect and therefore rely on that insect for successful reproduction. This close relationship

5586-625: The burial practices of Europeans from the late middle ages into the early modern period. During renovations on the Grote Kerk of Breda a tomb was rediscovered that was used between 1475 and 1526 AD by eight members of the house of Nassau . These burials had to be moved, but before being re-interred these burials were studied for botanical remains. The burial of Cimberga van Baden contained pollen from cloves. The Dutch Physician Pieter Van Foreest wrote down multiple recipes for embalming some of which included cloves. One of these recipes he wrote down

5684-517: The collective cluster of flowers is called an inflorescence . Some inflorescences are composed of many small flowers arranged in a formation that resembles a single flower. A common example of this is most members of the very large composite ( Asteraceae ) group. A single daisy or sunflower , for example, is not a flower but a flower head —an inflorescence composed of numerous flowers (or florets). An inflorescence may include specialized stems and modified leaves known as bracts . A floral formula

5782-558: The documentary film Hikayat di Ujung Pesisir and is ideal for camping, but with no facilities at all, and modest snorkeling can be done there. Clove Cloves are the aromatic flower buds of a tree in the family Myrtaceae , Syzygium aromaticum ( / s ɪ ˈ z ɪ dʒ iː ə m ˌ ær ə ˈ m æ t ɪ k ə m / ). They are native to the Maluku Islands , or Moluccas, in Indonesia , and are commonly used as

5880-550: The earliest known example of macro-botanical cloves in northwest Europe from the wreck of the Danish-Norwegian flagship, Gribshunden . The ship sank near Ronneby , Sweden in June 1495 while King Hans was sailing to political summit at Kalmar, Sweden. Exotic luxuries including cloves, ginger, peppercorns, and saffron would have impressed the noblemen and high church officials at the summit. Cloves have been documented in

5978-484: The existing model to broaden the descriptive capability of the formula. The format of floral formulae differs in different parts of the world, yet they convey the same information. The structure of a flower can also be expressed by the means of floral diagrams . The use of schematic diagrams can replace long descriptions or complicated drawings as a tool for understanding both floral structure and evolution. Such diagrams may show important features of flowers, including

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6076-572: The first century AD. Other archeological finds of cloves include: At the Batujaya site a single clove was found in a waterlogged layer dating to between the 100s BC to 200s BC corresponding to the Buni culture phase of this site. A study at the site of Óc Eo in the Mekong Delta of Vietnam found starch grains of cloves on stone implements used in food processing. This site was occupied from

6174-664: The first to eighth century BC, and was a trading center for the kingdom of Funnan . Two cloves were found during archaeological excavations at the Sri Lankan city of Mantai dated to around 900–1100 AD. Cloves are mentioned in the Ramayana . Cloves are also mentioned in the Charaka Samhita . One of the earliest examples of literary evidence of cloves in China is from the book the Han Guan Yi (Etiquettes of

6272-413: The flower is said to be actinomorphic or regular. This is an example of radial symmetry . When flowers are bisected and produce only one line that produces symmetrical halves, the flower is said to be irregular or zygomorphic . If, in rare cases, they have no symmetry at all they are called asymmetric. Flowers may be directly attached to the plant at their base ( sessile —the supporting stalk or stem

6370-497: The flower's stigma. This pollination does not require an investment from the plant to provide nectar and pollen as food for pollinators. Some flowers produce diaspores without fertilization ( parthenocarpy ). After fertilization, the ovary of the flower develops into fruit containing seeds . Flowers have long been appreciated for their beauty and pleasant scents, and also hold cultural significance as religious, ritual, or symbolic objects, or sources of medicine and food. Flower

6468-511: The flowers are imperfect or unisexual: having only either male (stamen) or female (carpel) parts. If unisexual male and female flowers appear on the same plant, the species is called monoecious . However, if an individual plant is either female or male, the species is called dioecious . Many flowers have nectaries , which are glands that produce a sugary fluid used to attract pollinators. They are not considered as an organ on their own. In those species that have more than one flower on an axis,

6566-424: The following two broad groups of pollination methods: Flowers that use biotic vectors attract and use insects , bats , birds , or other animals to transfer pollen from one flower to the next. Often they are specialized in shape and have an arrangement of the stamens that ensures that pollen grains are transferred to the bodies of the pollinator when it lands in search of its attractant (such as nectar, pollen, or

6664-451: The gametophytes also develop inside the spores, i.e., they are endosporic. Since the flowers are the reproductive organs of the plant, they mediate the joining of the sperm, contained within pollen, to the ovules — contained in the ovary. Pollination is the movement of pollen from the anthers to the stigma. Normally pollen is moved from one plant to another, known as cross-pollination , but many plants can self-pollinate. Cross-pollination

6762-594: The island of Sulawesi produced over 40% collectively. Eugenol comprises 72–90% of the essential oil extracted from cloves, and is the compound most responsible for clove aroma. Complete extraction occurs at 80 minutes in pressurized water at 125 °C (257 °F). Ultrasound-assisted and microwave-assisted extraction methods provide more rapid extraction rates with lower energy costs. Other phytochemicals of clove oil include acetyl eugenol, beta- caryophyllene , vanillin , crategolic acid , tannins , such as bicornin , gallotannic acid , methyl salicylate ,

6860-623: The locations of the district administrative centres, the number of administrative villages ( gampong ) in each district, and its post code. Notes: (a) comprises a group of islands off the north-western tip of Sumatra, of which the largest are Pulau Breuh , Pulau Nasi and Pulau Teunom. (b) includes some smaller islands off the north-western tip of Sumatra but closer to the mainland than Pulo Aceh District - Pulau Bunta and Pulau Batee. (c) comprises southern suburbs of Banda Aceh city, with 113,408 inhabitants in mid 2023. Rusa Island in Lhoong district

6958-400: The ovary. After penetrating the center-most part of the ovary it enters the egg apparatus and into one synergid . At this point the end of the pollen tube bursts and releases the two sperm cells, one of which makes its way to an egg, while also losing its cell membrane and much of its protoplasm . The sperm's nucleus then fuses with the egg's nucleus, resulting in the formation of a zygote ,

7056-504: The ovule is fertilized by pollen from the same flower or plant, occurs in flowers that always self-fertilize, such as many dandelions . Some flowers are self-pollinated and have flowers that never open or are self-pollinated before the flowers open; these flowers are called cleistogamous ; many species in the genus Viola exhibit this, for example. Conversely, many species of plants have ways of preventing self-pollination and hence, self-fertilization. Unisexual male and female flowers on

7154-524: The ovules. Carpels may occur in one to several whorls, and when fused are often described as a pistil . Inside the ovary, the ovules are attached to the placenta by structures called funiculi . Although this arrangement is considered "typical", plant species show a wide variation in floral structure. The four main parts of a flower are generally defined by their positions on the receptacle and not by their function. Many flowers lack some parts or parts may be modified into other functions or look like what

7252-534: The perianth. They are often delicate and thin and are usually colored, shaped, or scented to encourage pollination. Although similar to leaves in shape, they are more comparable to stamens in that they form almost simultaneously with one another, but their subsequent growth is delayed. If the corolla is fused together it is called sympetalous. The androecium , or stamens, is the whorl of pollen-producing male parts. Stamens consist typically of an anther , made up of four pollen sacs arranged in two thecae , connected to

7350-430: The personal collection of Ali Hasymi , a former governor of Aceh and artist and include books by the scholars of Acehnese past, ancient ceramics, the typical weapons of Aceh, souvenirs from all over the world, etc. Museum Cut Nyak Dhien was originally the home of the heroine Cut Nyak Dhien . The ancient Abee Tanoh Library , located at the foot of Mount Seulawa contains some important manuscripts. Indra Puri Old Mosque

7448-451: The plant so as to not force competition between the mother and the daughter plants, as well as to enable the colonization of new areas. They are often divided into two categories, though many plants fall in between or in one or more of these: In allochory, plants use an external vector , or carrier, to transport their seeds away from them. These can be either biotic (living), such as by birds and ants, or abiotic (non-living), such as by

7546-400: The process of pollination. Fertilization , also called Synagmy, occurs following pollination, which is the movement of pollen from the stamen to the carpel. It encompasses both plasmogamy , the fusion of the protoplasts , and karyogamy , the fusion of the nuclei . When pollen lands on the stigma of the flower it begins creating a pollen tube which runs down through the style and into

7644-600: The relative positions of the various organs, including the presence of fusion and symmetry, as well as structural details. A flower develops on a modified shoot or axis from a determinate apical meristem ( determinate meaning the axis grows to a set size). It has compressed internodes, bearing structures that in classical plant morphology are interpreted as highly modified leaves . Detailed developmental studies, however, have shown that stamens are often initiated more or less like modified stems (caulomes) that in some cases may even resemble branchlets . Taking into account

7742-420: The reproductive organs. Growth of the central part of the stem tip stops or flattens out and the sides develop protuberances in a whorled or spiral fashion around the outside of the stem end. These protuberances develop into the sepals, petals, stamens, and carpels . Once this process begins, in most plants, it cannot be reversed and the stems develop flowers, even if the initial start of the flower formation event

7840-742: The ripe fruits of cloves that are ovoid, brown berries, unilocular and one-seeded. Blown cloves are expanded flowers from which both corollae and stamens have been detached. Exhausted cloves have most or all the oil removed by distillation. They yield no oil and are darker in color. Cloves are used in the cuisine of Asian , African , Mediterranean , and the Near and Middle East countries, lending flavor to meats (such as baked ham ), curries , and marinades , as well as fruit (such as apples, pears, and rhubarb ). Cloves may be used to give aromatic and flavor qualities to hot beverages, often combined with other ingredients such as lemon and sugar. They are

7938-490: The same flower or another flower on the same plant, leading to the creation of a genetic clone through asexual reproduction . This increases the reliability of producing seeds, the rate at which they can be produced, and lowers the amount energy needed. But, most importantly, it limits genetic variation . In addition, self-pollination causes inbreeding depression , due largely to the expression of recessive deleterious mutations . The extreme case of self-fertilization, when

8036-412: The same plant may not appear or mature at the same time, or pollen from the same plant may be incapable of fertilizing its ovules. The latter flower types, which have chemical barriers to their own pollen, are referred to as self-incompatible. In Clianthus puniceus , self-pollination is used strategically as an "insurance policy". When a pollinator, in this case a bird, visits C. puniceus , it rubs off

8134-769: The same species. The period during which this process can take place (when the flower is fully expanded and functional) is called anthesis , hence the study of pollination biology is called anthecology . Flowering plants usually face evolutionary pressure to optimize the transfer of their pollen , and this is typically reflected in the morphology of the flowers and the behavior of the plants. Pollen may be transferred between plants via several 'vectors,' or methods. Around 80% of flowering plants make use of biotic or living vectors. Others use abiotic, or non-living, vectors and some plants make use of multiple vectors, but most are highly specialized. Though some fit between or outside of these groups, most flowers can be divided between

8232-482: The seed. The ovary, inside which the seed is forming from the ovule, grows into a fruit . All the other main floral parts die during this development, including: the style, stigma, sepals, stamens, and petals. The fruit contains three structures: the exocarp , or outer layer, the mesocarp , or the fleshy part, and the endocarp , or innermost layer, while the fruit wall is called the pericarp . The size, shape, toughness, and thickness varies among different fruit. This

8330-439: The species. All flowering plants are heterosporous , that is, every individual plant produces two types of spores . Microspores are produced by meiosis inside anthers and megaspores are produced inside ovules that are within an ovary. Anthers typically consist of four microsporangia and an ovule is an integumented megasporangium. Both types of spores develop into gametophytes inside sporangia. As with all heterosporous plants,

8428-428: The stigmatic covering and allows for pollen from the bird to enter the stigma. If no pollinators visit, however, then the stigmatic covering falls off naturally to allow for the flower's own anthers to pollinate the flower through self-pollination. Pollen is a large contributor to asthma and other respiratory allergies which combined affect between 10 and 50% of people worldwide. This number appears to be growing, as

8526-463: The temperature increases due to climate change mean that plants are producing more pollen , which is also more allergenic. Pollen is difficult to avoid, however, because of its small size and prevalence in the natural environment. Most of the pollen which causes allergies is that produced by wind-dispersed pollinators such as the grasses , birch trees , oak trees, and ragweeds ; the allergens in pollen are proteins which are thought to be necessary in

8624-428: The vegetative part, consisting of non-reproductive structures such as petals ; and the reproductive or sexual parts. A stereotypical flower is made up of four kinds of structures attached to the tip of a short stalk or axis, called a receptacle . Each of these parts or floral organs is arranged in a spiral called a whorl . The four main whorls (starting from the base of the flower or lowest node and working upwards) are

8722-619: The water and so the pollen has a low density to enable floating, though many also use rafts, and are hydrophobic . Marine flowers have floating thread-like stigmas and may have adaptations for the tide, while freshwater species create indentations in the water. The third category, set out by Schwarzenbach, is those flowers which transport pollen above the water through conveyance. This ranges from floating plants, ( Lemnoideae ), to staminate flowers ( Vallisneria ). Most species in this group have dry, spherical pollen which sometimes forms into larger masses, and female flowers which form depressions in

8820-401: The water; the method of transport varies. Flowers can be pollinated by two mechanisms; cross-pollination and self-pollination. No mechanism is indisputably better than the other as they each have their advantages and disadvantages. Plants use one or both of these mechanisms depending on their habitat and ecological niche . Cross-pollination is the pollination of the carpel by pollen from

8918-480: The whole diversity in the development of the androecium of flowering plants, we find a continuum between modified leaves (phyllomes), modified stems (caulomes), and modified branchlets (shoots). The transition to flowering is one of the major phase changes that a plant makes during its life cycle. The transition must take place at a time that is favorable for fertilization and the formation of seeds , hence ensuring maximal reproductive success. To meet these needs

9016-460: The wind or water. Many plants use biotic vectors to disperse their seeds away from them. This method falls under the umbrella term zoochory , while endozoochory , also known as fruigivory, refers specifically to plants adapted to grow fruit in order to attract animals to eat them. Once eaten they go through typically go through animal's digestive system and are dispersed away from the plant. Some seeds are specially adapted either to last in

9114-419: The wind to effectively pick up and transport the pollen, the flowers typically have anthers loosely attached to the end of long thin filaments, or pollen forms around a catkin which moves in the wind. Rarer forms of this involve individual flowers being moveable by the wind ( pendulous ), or even less commonly; the anthers exploding to release the pollen into the wind. Pollination through water ( hydrophily )

9212-490: Was buried in 1737 AD contained whole cloves. During the colonial era, cloves were traded like oil, with an enforced limit on exportation. As the Dutch East India Company consolidated its control of the spice trade in the 17th century, they sought to gain a monopoly in cloves as they had in nutmeg. However, "unlike nutmeg and mace , which were limited to the minute Bandas , clove trees grew all over

9310-416: Was dependent on some environmental cue. The ABC model is a simple model that describes the genes responsible for the development of flowers. Three gene activities interact in a combinatorial manner to determine the developmental identities of the primordia organ within the floral apical meristem . These gene functions are called A, B, and C. Genes are expressed in only the outer and lower most section of

9408-492: Was once the world's largest producer of cloves. Current leaders in clove production are Indonesia , Madagascar , Tanzania , Sri Lanka , and Comoros . Indonesia is the largest clove producer, but only about 10-15% of its cloves production is exported, and domestic shortfalls must sometimes be filled with imports from Madagascar. The modern province of Maluku remains the largest source of cloves in Indonesia with around 15% of national production, although provinces comprising

9506-439: Was shaped like Rusa (deer) before the tsunami struck the island on 26 December 2004, but now that much of it has been washed away, the form of the island has changed and become smaller like a lamb embryo. The island is very important for marking Indonesia's boundaries due to its location as the most westerly island in Indonesia with Titik Dasar TD175 and Titik Referensi TR175 (Base Point TD175 and Reference Point TR175). Since

9604-493: Was that used by his fellow physicians Spierinck and Goethals. An embalming jar associated with Vittoria della Rovere also contained clove pollen. This probably came from her ingestion of clove oil as a medicine in her final days. When burials needed to be moved from the church of Saint Germain in Flers , France they were also studied for botanical remains. The body and coffin of Philippe René de la Motte Ango, count of Flers who

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