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95-432: From north to south, the islands consist of: Namua and Fanuatapu lie at the outer edge of the fringing reef of Upolu, at a distance of 0.7 km and 2.5 km, respectively, from Upolu Island itself. Nuʻutele and Nuʻulua are 4 to 6 km further south, outside the fringing reef, and lie 1.4 km 3.5 km off Cape Tapaga, which is the southeastern headland of Upolu. Only Namua is open for visitors, who can enjoy

190-749: A Darwin Point is reached. One such point exists at the northwestern end of the Hawaiian Archipelago; see Evolution of Hawaiian volcanoes#Coral atoll stage . However, reefs in the Persian Gulf have adapted to temperatures of 13 °C (55 °F) in winter and 38 °C (100 °F) in summer. 37 species of scleractinian corals inhabit such an environment around Larak Island . Deep-water coral inhabits greater depths and colder temperatures at much higher latitudes, as far north as Norway. Although deep water corals can form reefs, little

285-644: A continental shelf, and not around a volcanic island, Darwin's principles apply. Development stopped at the barrier reef stage, since Australia is not about to submerge. It formed the world's largest barrier reef, 300–1,000 m (980–3,280 ft) from shore, stretching for 2,000 km (1,200 mi). Healthy tropical coral reefs grow horizontally from 1 to 3 cm (0.39 to 1.18 in) per year, and grow vertically anywhere from 1 to 25 cm (0.39 to 9.84 in) per year; however, they grow only at depths shallower than 150 m (490 ft) because of their need for sunlight, and cannot grow above sea level. As

380-1378: A lesser extent in the Southern Hemisphere. They have different names, historically developed and depending on their geographic distributions: Fraser (in the Pacific Cordillera of North America), Pinedale (in the Central Rocky Mountains ), Wisconsinan or Wisconsin (in central North America), Devensian (in the British Isles), Midlandian (in Ireland), Würm (in the Alps ), Mérida (in Venezuela ), Weichselian or Vistulian (in Northern Europe and northern Central Europe), Valdai in Russia and Zyryanka in Siberia , Llanquihue in Chile , and Otira in New Zealand. The geochronological Late Pleistocene includes

475-452: A nesting place for critically-endangered Hawksbill sea turtles , and birds including the Samoan flycatcher and Manumean . In 2009 an attempt was made to eradicate the introduced Polynesian rat from the southern islands using air-dropped Brodifacoum to provide a pest-free refuge. The attempt was unsuccessful, with rats being detected again in 2011. Coral reef A coral reef

570-412: A pinhead to 12 inches (30 cm) across. Reef-building or hermatypic corals live only in the photic zone (above 70 m), the depth to which sufficient sunlight penetrates the water. Coral polyps do not photosynthesize, but have a symbiotic relationship with microscopic algae ( dinoflagellates ) of the genus Symbiodinium , commonly referred to as zooxanthellae . These organisms live within

665-496: A pseudo-atoll. These can be distinguished from real atolls only by detailed investigation, possibly including core drilling. Some platform reefs of the Laccadives are U-shaped, due to wind and water flow. Atolls or atoll reefs are a more or less circular or continuous barrier reef that extends all the way around a lagoon without a central island. They are usually formed from fringing reefs around volcanic islands. Over time,

760-433: A rapidly subsiding bottom can overwhelm coral growth, killing the coral and the reef, due to what is called coral drowning . Corals that rely on zooxanthellae can die when the water becomes too deep for their symbionts to adequately photosynthesize , due to decreased light exposure. The two main variables determining the geomorphology , or shape, of coral reefs are the nature of the substrate on which they rest, and

855-527: A result of melting ice, the land has continued to rise yearly in Scandinavia, mostly in northern Sweden and Finland, where the land is rising at a rate of as much as 8–9 mm per year, or 1 m in 100 years. This is important for archaeologists, since a site that was coastal in the Nordic Stone Age now is inland and can be dated by its relative distance from the present shore. The term Würm

950-458: A single ice age given the continual presence of ice sheets near both poles. Glacials are somewhat better defined, as colder phases during which glaciers advance, separated by relatively warm interglacials . The end of the last glacial period, which was about 10,000 years ago, is often called the end of the ice age, although extensive year-round ice persists in Antarctica and Greenland . Over

1045-797: A still lesser extent, glaciers existed in Africa, for example in the High Atlas , the mountains of Morocco , the Mount Atakor massif in southern Algeria , and several mountains in Ethiopia . Just south of the equator, an ice cap of several hundred square kilometers was present on the east African mountains in the Kilimanjaro massif , Mount Kenya , and the Rwenzori Mountains , which still bear relic glaciers today. Glaciation of

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1140-423: A zone extending from approximately 30° N to 30° S of the equator. Tropical corals do not grow at depths of over 50 meters (160 ft). The optimum temperature for most coral reefs is 26–27 °C (79–81 °F), and few reefs exist in waters below 18 °C (64 °F). When the net production by reef building corals no longer keeps pace with relative sea level and the reef structure permanently drowns

1235-454: Is an underwater ecosystem characterized by reef-building corals . Reefs are formed of colonies of coral polyps held together by calcium carbonate . Most coral reefs are built from stony corals , whose polyps cluster in groups. Coral belongs to the class Anthozoa in the animal phylum Cnidaria , which includes sea anemones and jellyfish . Unlike sea anemones, corals secrete hard carbonate exoskeletons that support and protect

1330-578: Is constantly changing. Each reef is made up of irregular patches of algae, sessile invertebrates, and bare rock and sand. The size, shape and relative abundance of these patches change from year to year in response to the various factors that favor one type of patch over another. Growing coral, for example, produces constant change in the fine structure of reefs. On a larger scale, tropical storms may knock out large sections of reef and cause boulders on sandy areas to move. Coral reefs are estimated to cover 284,300 km (109,800 sq mi), just under 0.1% of

1425-605: Is derived from a river in the Alpine foreland, roughly marking the maximum glacier advance of this particular glacial period. The Alps were where the first systematic scientific research on ice ages was conducted by Louis Agassiz at the beginning of the 19th century. Here, the Würm glaciation of the LGP was intensively studied. Pollen analysis , the statistical analyses of microfossilized plant pollens found in geological deposits, chronicled

1520-445: Is directly attached to a shore, or borders it with an intervening narrow, shallow channel or lagoon. It is the most common reef type. Fringing reefs follow coastlines and can extend for many kilometres. They are usually less than 100 metres wide, but some are hundreds of metres wide. Fringing reefs are initially formed on the shore at the low water level and expand seawards as they grow in size. The final width depends on where

1615-523: Is known about them. The northernmost coral reef on Earth is located near Eilat , Israel . Coral reefs are rare along the west coasts of the Americas and Africa , due primarily to upwelling and strong cold coastal currents that reduce water temperatures in these areas (the Humboldt , Benguela , and Canary Currents , respectively). Corals are seldom found along the coastline of South Asia —from

1710-418: Is often the reef's most diverse area. Coral and calcareous algae provide complex habitats and areas that offer protection, such as cracks and crevices. Invertebrates and epiphytic algae provide much of the food for other organisms. A common feature on this forereef zone is spur and groove formations that serve to transport sediment downslope. The reef flat is the sandy-bottomed flat, which can be behind

1805-655: Is ongoing. The glaciation and the current Quaternary Period both began with the formation of the Arctic ice cap . The Antarctic ice sheet began to form earlier, at about 34 Mya, in the mid- Cenozoic ( Eocene–Oligocene extinction event ), and the term Late Cenozoic Ice Age is used to include this early phase with the current glaciation. The previous ice age within the Quaternary is the Penultimate Glacial Period , which ended about 128,000 years ago,

1900-695: The Last glacial cycle , occurred from the end of the Last Interglacial to the beginning of the Holocene , c.  115,000  – c.  11,700 years ago, and thus corresponds to most of the timespan of the Late Pleistocene . The LGP is part of a larger sequence of glacial and interglacial periods known as the Quaternary glaciation which started around 2,588,000 years ago and

1995-475: The Late Cretaceous (100–66 Ma), when reefs formed by a group of bivalves called rudists existed; one of the valves formed the main conical structure and the other, much smaller valve acted as a cap. Measurements of the oxygen isotopic composition of the aragonitic skeleton of coral reefs, such as Porites , can indicate changes in sea surface temperature and sea surface salinity conditions during

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2090-607: The Lesotho Highlands and parts of the Drakensberg . The development of glaciers was likely aided in part due to shade provided by adjacent cliffs. Various moraines and former glacial niches have been identified in the eastern Lesotho Highlands a few kilometres west of the Great Escarpment , at altitudes greater than 3,000 m on south-facing slopes. Studies suggest that the annual average temperature in

2185-739: The Mesoamerican Barrier Reef System and the New Caledonian Barrier Reef . Barrier reefs are also found on the coasts of Providencia , Mayotte , the Gambier Islands , on the southeast coast of Kalimantan , on parts of the coast of Sulawesi , southeastern New Guinea and the south coast of the Louisiade Archipelago . Platform reefs, variously called bank or table reefs, can form on the continental shelf , as well as in

2280-592: The Middle Cambrian (513–501 Ma ), Devonian (416–359 Ma) and Carboniferous (359–299 Ma), owing to extinct order Rugosa corals, and Late Cretaceous (100–66 Ma) and Neogene (23 Ma–present), owing to order Scleractinia corals. Not all reefs in the past were formed by corals: those in the Early Cambrian (542–513 Ma) resulted from calcareous algae and archaeocyathids (small animals with conical shape, probably related to sponges ) and in

2375-727: The Owen Stanley Range , and the Saruwaged Range . Mount Giluwe in the Central Cordillera had a "more or less continuous ice cap covering about 188 km and extending down to 3200-3500 m". In Western New Guinea , remnants of these glaciers are still preserved atop Puncak Jaya and Ngga Pilimsit . Small glaciers developed in a few favorable places in Southern Africa during the last glacial period. These small glaciers would have been located in

2470-720: The Sierra Nevada in northern California . In northern Eurasia, the Scandinavian ice sheet once again reached the northern parts of the British Isles , Germany , Poland , and Russia, extending as far east as the Taymyr Peninsula in western Siberia. The maximum extent of western Siberian glaciation was reached by about 18,000 to 17,000 BP, later than in Europe (22,000–18,000 BP). Northeastern Siberia

2565-732: The Upper Mississippi River , which in turn was formed during an earlier glacial period. In its retreat, the Wisconsin episode glaciation left terminal moraines that form Long Island , Block Island , Cape Cod , Nomans Land , Martha's Vineyard , Nantucket , Sable Island , and the Oak Ridges Moraine in south-central Ontario, Canada. In Wisconsin itself, it left the Kettle Moraine . The drumlins and eskers formed at its melting edge are landmarks of

2660-537: The Younger Dryas , began around 12,800 years ago and ended around 11,700 years ago, also marking the end of the LGP and the Pleistocene epoch. It was followed by the Holocene , the current geological epoch . The LGP is often colloquially referred to as the "last ice age", though the term ice age is not strictly defined, and on a longer geological perspective, the last few million years could be termed

2755-584: The grooves left by these glaciers can be easily observed. In southwestern Saskatchewan and southeastern Alberta, a suture zone between the Laurentide and Cordilleran ice sheets formed the Cypress Hills , which is the northernmost point in North America that remained south of the continental ice sheets. The Great Lakes are the result of glacial scour and pooling of meltwater at the rim of

2850-586: The isostatically depressed area, a temporary marine incursion that geologists dub the Yoldia Sea . Then, as postglacial isostatic rebound lifted the region about 9500 BP, the deepest basin of the Baltic became a freshwater lake, in palaeological contexts referred to as Ancylus Lake , which is identifiable in the freshwater fauna found in sediment cores. The lake was filled by glacial runoff, but as worldwide sea level continued rising, saltwater again breached

2945-417: The tropics where the water is agitated, but can only live within a limited depth range, starting just below low tide . Where the level of the underlying earth allows, the corals grow around the coast to form fringing reefs, and can eventually grow to become a barrier reef. Where the bottom is rising, fringing reefs can grow around the coast, but coral raised above sea level dies. If the land subsides slowly,

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3040-778: The Alpine foreland . Local ice fields or small ice sheets could be found capping the highest massifs of the Pyrenees , the Carpathian Mountains , the Balkan Mountains , the Caucasus , and the mountains of Turkey and Iran . In the Himalayas and the Tibetan Plateau , there is evidence that glaciers advanced considerably, particularly between 47,000 and 27,000 BP, but the exact ages, as well as

3135-745: The Center for Arctic Gas Hydrate, Environment and Climate at the University of Tromsø , published a study in June 2017 describing over a hundred ocean sediment craters, some 3,000 m wide and up to 300 m deep, formed by explosive eruptions of methane from destabilized methane hydrates , following ice-sheet retreat during the LGP, around 12,000 years ago. These areas around the Barents Sea still seep methane today. The study hypothesized that existing bulges containing methane reservoirs could eventually have

3230-598: The Greenland climate was dry during the LGP, with precipitation reaching perhaps only 20% of today's value. The name Mérida glaciation is proposed to designate the alpine glaciation that affected the central Venezuelan Andes during the Late Pleistocene. Two main moraine levels have been recognized - one with an elevation of 2,600–2,700 m (8,500–8,900 ft), and another with an elevation of 3,000–3,500 m (9,800–11,500 ft). The snow line during

3325-738: The Indian Ocean, for example, in the Maldives , the Chagos Islands , the Seychelles and around Cocos Island . The entire Maldives consist of 26 atolls. Coral reef ecosystems contain distinct zones that host different kinds of habitats. Usually, three major zones are recognized: the fore reef, reef crest, and the back reef (frequently referred to as the reef lagoon). The three zones are physically and ecologically interconnected. Reef life and oceanic processes create opportunities for

3420-1010: The LGP as the Devensian . Irish geologists, geographers, and archaeologists refer to the Midlandian glaciation, as its effects in Ireland are largely visible in the Irish Midlands . The name Devensian is derived from the Latin Dēvenses , people living by the Dee ( Dēva in Latin), a river on the Welsh border near which deposits from the period are particularly well represented. The effects of this glaciation can be seen in many geological features of England, Wales, Scotland, and Northern Ireland . Its deposits have been found overlying material from

3515-618: The Namua Island Resort and the beaches. Fanuatapu, being the easternmost island, has a lighthouse . The islands are uninhabited and relatively undisturbed, and provide an important refuge for native plant and animal species. The northern islands are mostly bare rock and coastal forest, while the larger southern group are mostly covered by coastal and lowland forest. The islands are home to the largest population of Tongan ground doves in Samoa, as well as Coconut crabs , and provides

3610-868: The North American Laurentide ice sheet. At the height of glaciation, the Bering land bridge potentially permitted migration of mammals, including people, to North America from Siberia . It radically altered the geography of North America north of the Ohio River . At the height of the Wisconsin episode glaciation, ice covered most of Canada, the Upper Midwest , and New England , as well as parts of Montana and Washington . On Kelleys Island in Lake Erie or in New York's Central Park ,

3705-410: The Polish River Vistula or its German name Weichsel). Evidence suggests that the ice sheets were at their maximum size for only a short period, between 25,000 and 13,000 BP. Eight interstadials have been recognized in the Weichselian, including the Oerel, Glinde, Moershoofd, Hengelo, and Denekamp. Correlation with isotope stages is still in process. During the glacial maximum in Scandinavia, only

3800-437: The Southern Hemisphere was less extensive. Ice sheets existed in the Andes ( Patagonian Ice Sheet ), where six glacier advances between 33,500 and 13,900 BP in the Chilean Andes have been reported. Antarctica was entirely glaciated, much like today, but unlike today the ice sheet left no uncovered area. In mainland Australia only a very small area in the vicinity of Mount Kosciuszko was glaciated, whereas in Tasmania glaciation

3895-551: The UV radiation and allowing them to better adapt to warmer water temperatures. In the event of UV or thermal damage, if and when repair occurs, it will increase the likelihood of survival of the host and symbiont. This leads to the idea that, evolutionarily, clade A is more UV resistant and thermally resistant than the other clades. Clades B and C are found more frequently in deeper water, which may explain their higher vulnerability to increased temperatures. Terrestrial plants that receive less sunlight because they are found in

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3990-452: The Wisconsin episode. It began about 30,000 years ago, reached its greatest advance 21,000 years ago, and ended about 10,000 years ago. In northwest Greenland, ice coverage attained a very early maximum in the LGP around 114,000. After this early maximum, ice coverage was similar to today until the end of the last glacial period. Towards the end, glaciers advanced once more before retreating to their present extent. According to ice core data,

4085-1188: The area of France, yet they provide a home for at least 25% of all marine species , including fish , mollusks , worms , crustaceans , echinoderms , sponges , tunicates and other cnidarians . Coral reefs flourish in ocean waters that provide few nutrients. They are most commonly found at shallow depths in tropical waters, but deep water and cold water coral reefs exist on smaller scales in other areas. Shallow tropical coral reefs have declined by 50% since 1950, partly because they are sensitive to water conditions. They are under threat from excess nutrients (nitrogen and phosphorus), rising ocean heat content and acidification , overfishing (e.g., from blast fishing , cyanide fishing , spearfishing on scuba ), sunscreen use, and harmful land-use practices, including runoff and seeps (e.g., from injection wells and cesspools). Coral reefs deliver ecosystem services for tourism, fisheries and shoreline protection . The annual global economic value of coral reefs has been estimated at anywhere from US$ 30–375 billion (1997 and 2003 estimates) to US$ 2.7 trillion (a 2020 estimate) to US$ 9.9 trillion (a 2014 estimate). Though

4180-448: The coast. The fringing reefs of the Red Sea are "some of the best developed in the world" and occur along all its shores except off sandy bays. Barrier reefs are separated from a mainland or island shore by a deep channel or lagoon . They resemble the later stages of a fringing reef with its lagoon but differ from the latter mainly in size and origin. Their lagoons can be several kilometres wide and 30 to 70 metres deep. Above all,

4275-416: The coastal plains had become continental islands . As sea level rise continued, water topped most of the continental islands. The corals could then overgrow the hills, forming cays and reefs. Sea level on the Great Barrier Reef has not changed significantly in the last 6,000 years. The age of living reef structure is estimated to be between 6,000 and 8,000 years. Although the Great Barrier Reef formed along

4370-451: The coral animals. Coral that loses a large fraction of its zooxanthellae becomes white (or sometimes pastel shades in corals that are pigmented with their own proteins) and is said to be bleached , a condition which, unless corrected, can kill the coral. There are eight clades of Symbiodinium phylotypes . Most research has been conducted on clades A–D. Each clade contributes their own benefits as well as less compatible attributes to

4465-412: The coral. Most reefs grow best in warm, shallow, clear, sunny and agitated water. Coral reefs first appeared 485 million years ago, at the dawn of the Early Ordovician , displacing the microbial and sponge reefs of the Cambrian . Sometimes called rainforests of the sea , shallow coral reefs form some of Earth's most diverse ecosystems. They occupy less than 0.1% of the world's ocean area, about half

4560-415: The corals shelter the zooxanthellae, averaging one million for every cubic centimetre of coral, and provide a constant supply of the carbon dioxide they need for photosynthesis. The varying pigments in different species of zooxanthellae give them an overall brown or golden-brown appearance and give brown corals their colors. Other pigments such as reds, blues, greens, etc. come from colored proteins made by

4655-477: The deep sea away from continental shelves , around oceanic islands and atolls . The majority of these islands are volcanic in origin. Others have tectonic origins where plate movements lifted the deep ocean floor. In The Structure and Distribution of Coral Reefs , Charles Darwin set out his theory of the formation of atoll reefs, an idea he conceived during the voyage of the Beagle . He theorized that uplift and subsidence of Earth's crust under

4750-400: The dramatic changes in the European environment during the Würm glaciation. During the height of Würm glaciation, c.  24,000  – c.  10,000  BP, most of western and central Europe and Eurasia was open steppe-tundra, while the Alps presented solid ice fields and montane glaciers. Scandinavia and much of Britain were under ice. During the Würm, the Rhône Glacier covered

4845-527: The eastern tip of India ( Chennai ) to the Bangladesh and Myanmar borders —as well as along the coasts of northeastern South America and Bangladesh, due to the freshwater release from the Amazon and Ganges Rivers respectively. Significant coral reefs include: When alive, corals are colonies of small animals embedded in calcium carbonate shells. Coral heads consist of accumulations of individual animals called polyps , arranged in diverse shapes. Polyps are usually tiny, but they can range in size from

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4940-411: The exchange of seawater , sediments , nutrients and marine life. Most coral reefs exist in waters less than 50 m deep. Some inhabit tropical continental shelves where cool, nutrient-rich upwelling does not occur, such as the Great Barrier Reef . Others are found in the deep ocean surrounding islands or as atolls, such as in the Maldives . The reefs surrounding islands form when islands subside into

5035-401: The floor often supports seagrass meadows which are important foraging areas for reef fish. The reef drop-off is, for its first 50 m, habitat for reef fish who find shelter on the cliff face and plankton in the water nearby. The drop-off zone applies mainly to the reefs surrounding oceanic islands and atolls. The reef face is the zone above the reef floor or the reef drop-off. This zone

5130-423: The formation of a single contiguous ice sheet on the Tibetan Plateau, is controversial. Other areas of the Northern Hemisphere did not bear extensive ice sheets, but local glaciers were widespread at high altitudes. Parts of Taiwan , for example, were repeatedly glaciated between 44,250 and 10,680 BP as well as the Japanese Alps . In both areas, maximum glacier advance occurred between 60,000 and 30,000 BP. To

5225-403: The fringing reef, barrier reef, atoll and table reef based on Stoddart, D.R. (1969). Spalding et al. list four main reef types that can be clearly illustrated – the fringing reef, barrier reef, atoll, and "bank or platform reef"—and notes that many other structures exist which do not conform easily to strict definitions, including the "patch reef". A fringing reef, also called a shore reef,

5320-463: The fringing reefs keep pace by growing upwards on a base of older, dead coral, forming a barrier reef enclosing a lagoon between the reef and the land. A barrier reef can encircle an island, and once the island sinks below sea level a roughly circular atoll of growing coral continues to keep up with the sea level, forming a central lagoon. Barrier reefs and atolls do not usually form complete circles but are broken in places by storms. Like sea level rise ,

5415-419: The growth of the coral. This technique is often used by climate scientists to infer a region's paleoclimate . Since Darwin's identification of the three classical reef formations – the fringing reef around a volcanic island becoming a barrier reef and then an atoll – scientists have identified further reef types. While some sources find only three, Thomas lists "Four major forms of large-scale coral reefs" –

5510-489: The history of the change in sea level relative to that substrate. The approximately 20,000-year-old Great Barrier Reef offers an example of how coral reefs formed on continental shelves. Sea level was then 120 m (390 ft) lower than in the 21st century. As sea level rose, the water and the corals encroached on what had been hills of the Australian coastal plain. By 13,000 years ago, sea level had risen to 60 m (200 ft) lower than at present, and many hills of

5605-411: The island erodes away and sinks below sea level. Atolls may also be formed by the sinking of the seabed or rising of the sea level. A ring of reefs results, which enclose a lagoon. Atolls are numerous in the South Pacific, where they usually occur in mid-ocean, for example, in the Caroline Islands , the Cook Islands , French Polynesia , the Marshall Islands and Micronesia . Atolls are found in

5700-447: The last glacial advance was lowered approximately 1,200 m (3,900 ft) below the present snow line, which is 3,700 m (12,100 ft). The glaciated area in the Cordillera de Mérida was about 600 km (230 sq mi); this included these high areas, from southwest to northeast: Páramo de Tamá, Páramo Batallón, Páramo Los Conejos, Páramo Piedras Blancas, and Teta de Niquitao. Around 200 km (77 sq mi) of

5795-444: The last glacial maximum, the Patagonian ice sheet extended over the Andes from about 35°S to Tierra del Fuego at 55°S. The western part appears to have been very active, with wet basal conditions, while the eastern part was cold-based. Cryogenic features such as ice wedges , patterned ground , pingos , rock glaciers , palsas , soil cryoturbation , and solifluction deposits developed in unglaciated extra-Andean Patagonia during

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5890-442: The late glacial (Weichselian) and the immediately preceding penultimate interglacial ( Eemian ) period. Canada was almost completely covered by ice, as was the northern part of the United States , both blanketed by the huge Laurentide Ice Sheet . Alaska remained mostly ice free due to arid climate conditions. Local glaciations existed in the Rocky Mountains and the Cordilleran ice sheet and as ice fields and ice caps in

5985-401: The lower Connecticut River Valley . In the Sierra Nevada , three stages of glacial maxima, sometimes incorrectly called ice ages , were separated by warmer periods. These glacial maxima are called, from oldest to youngest, Tahoe, Tenaya, and Tioga. The Tahoe reached its maximum extent perhaps about 70,000 years ago. Little is known about the Tenaya. The Tioga was the least severe and last of

6080-448: The main Wisconsin glacial advance. The upper level probably represents the last glacial advance (Late Wisconsin). The Llanquihue glaciation takes its name from Llanquihue Lake in southern Chile , which is a fan-shaped piedmont glacial lake. On the lake's western shores, large moraine systems occur, of which the innermost belong to the LGP. Llanquihue Lake's varves are a node point in southern Chile's varve geochronology . During

6175-418: The main reef, containing chunks of coral. This zone may border a lagoon and serve as a protective area, or it may lie between the reef and the shore, and in this case is a flat, rocky area. Fish tend to prefer it when it is present. The reef lagoon is an entirely enclosed region, which creates an area less affected by wave action and often contains small reef patches. However, the topography of coral reefs

6270-484: The mainland. Unlike fringing and barrier reefs which extend only seaward, platform reefs grow in all directions. They are variable in size, ranging from a few hundred metres to many kilometres across. Their usual shape is oval to elongated. Parts of these reefs can reach the surface and form sandbanks and small islands around which may form fringing reefs. A lagoon may form In the middle of a platform reef. Platform reefs are typically situated within atolls, where they adopt

6365-430: The massive Missoula Floods . USGS geologists estimate that the cycle of flooding and reformation of the lake lasted an average of 55 years and that the floods occurred about 40 times over the 2,000-year period starting 15,000 years ago. Glacial lake outburst floods such as these are not uncommon today in Iceland and other places. The Wisconsin glacial episode was the last major advance of continental glaciers in

6460-502: The mountains of Southern Africa was about 6 °C colder than at present, in line with temperature drops estimated for Tasmania and southern Patagonia during the same time. This resulted in an environment of relatively arid periglaciation without permafrost , but with deep seasonal freezing on south-facing slopes. Periglaciation in the eastern Drakensberg and Lesotho Highlands produced solifluction deposits and blockfields ; including blockstreams and stone garlands. Scientists from

6555-400: The name "patch reefs" and often span a diameter of just a few dozen meters. In instances where platform reefs develop along elongated structures, such as old and weathered barrier reefs, they tend to arrange themselves in a linear formation. This is the case, for example, on the east coast of the Red Sea near Jeddah . In old platform reefs, the inner part can be so heavily eroded that it forms

6650-528: The name implies, coral reefs are made up of coral skeletons from mostly intact coral colonies. As other chemical elements present in corals become incorporated into the calcium carbonate deposits, aragonite is formed. However, shell fragments and the remains of coralline algae such as the green-segmented genus Halimeda can add to the reef's ability to withstand damage from storms and other threats. Such mixtures are visible in structures such as Eniwetok Atoll . The times of maximum reef development were in

6745-415: The ocean, and atolls form when an island subsides below the surface of the sea. Alternatively, Moyle and Cech distinguish six zones, though most reefs possess only some of the zones. The reef surface is the shallowest part of the reef. It is subject to surge and tides . When waves pass over shallow areas, they shoal , as shown in the adjacent diagram. This means the water is often agitated. These are

6840-528: The oceans formed the atolls. Darwin set out a sequence of three stages in atoll formation. A fringing reef forms around an extinct volcanic island as the island and ocean floor subside. As the subsidence continues, the fringing reef becomes a barrier reef and ultimately an atoll reef. Darwin predicted that underneath each lagoon would be a bedrock base, the remains of the original volcano. Subsequent research supported this hypothesis. Darwin's theory followed from his understanding that coral polyps thrive in

6935-589: The oceans' surface area. The Indo-Pacific region (including the Red Sea , Indian Ocean , Southeast Asia and the Pacific ) account for 91.9% of this total. Southeast Asia accounts for 32.3% of that figure, while the Pacific including Australia accounts for 40.8%. Atlantic and Caribbean coral reefs account for 7.6%. Although corals exist both in temperate and tropical waters, shallow-water reefs form only in

7030-490: The offshore outer reef edge formed in open water rather than next to a shoreline. Like an atoll, it is thought that these reefs are formed either as the seabed lowered or sea level rose. Formation takes considerably longer than for a fringing reef, thus barrier reefs are much rarer. The best known and largest example of a barrier reef is the Australian Great Barrier Reef . Other major examples are

7125-451: The open ocean, in fact anywhere where the seabed rises close enough to the surface of the ocean to enable the growth of zooxanthemic, reef-forming corals. Platform reefs are found in the southern Great Barrier Reef, the Swain and Capricorn Group on the continental shelf, about 100–200 km from the coast. Some platform reefs of the northern Mascarenes are several thousand kilometres from

7220-592: The past few million years, the glacial-interglacial cycles have been "paced" by periodic variations in the Earth's orbit via Milankovitch cycles . The LGP has been intensively studied in North America, northern Eurasia, the Himalayas , and other formerly glaciated regions around the world. The glaciations that occurred during this glacial period covered many areas, mainly in the Northern Hemisphere and to

7315-429: The polyps' tissues and provide organic nutrients that nourish the polyp in the form of glucose , glycerol and amino acids . Because of this relationship, coral reefs grow much faster in clear water, which admits more sunlight. Without their symbionts, coral growth would be too slow to form significant reef structures. Corals get up to 90% of their nutrients from their symbionts. In return, as an example of mutualism ,

7410-745: The preceding Ipswichian stage and lying beneath those from the following Holocene , which is the current stage. This is sometimes called the Flandrian interglacial in Britain. The latter part of the Devensian includes pollen zones I–IV, the Allerød oscillation and Bølling oscillation , and the Oldest Dryas , Older Dryas , and Younger Dryas cold periods. Alternative names include Weichsel glaciation or Vistulian glaciation (referring to

7505-420: The precise condition under which corals flourish. The light is sufficient for photosynthesis by the symbiotic zooxanthellae, and agitated water brings plankton to feed the coral. The off-reef floor is the shallow sea floor surrounding a reef. This zone occurs next to reefs on continental shelves. Reefs around tropical islands and atolls drop abruptly to great depths and do not have such a floor. Usually sandy,

7600-557: The proglacial rivers' shifting and redepositing gravels. Beneath the surface, they had profound and lasting influence on geothermal heat and the patterns of deep groundwater flow. The Pinedale (central Rocky Mountains) or Fraser (Cordilleran ice sheet) glaciation was the last of the major glaciations to appear in the Rocky Mountains in the United States. The Pinedale lasted from around 30,000 to 10,000 years ago, and

7695-529: The receding ice. When the enormous mass of the continental ice sheet retreated, the Great Lakes began gradually moving south due to isostatic rebound of the north shore. Niagara Falls is also a product of the glaciation, as is the course of the Ohio River, which largely supplanted the prior Teays River . With the assistance of several very broad glacial lakes, it released floods through the gorge of

7790-399: The same fate. During the last glacial period, Antarctica was blanketed by a massive ice sheet, much as it is today. The ice covered all land areas and extended into the ocean onto the middle and outer continental shelf. Counterintuitively though, according to ice modeling done in 2002, ice over central East Antarctica was generally thinner than it is today. British geologists refer to

7885-412: The sea bed begins to drop steeply. The surface of the fringe reef generally remains at the same height: just below the waterline. In older fringing reefs, whose outer regions pushed far out into the sea, the inner part is deepened by erosion and eventually forms a lagoon . Fringing reef lagoons can become over 100 metres wide and several metres deep. Like the fringing reef itself, they run parallel to

7980-609: The shallow water tropical coral reefs are best known, there are also deeper water reef-forming corals, which live in colder water and in temperate seas. Most coral reefs were formed after the Last Glacial Period when melting ice caused sea level to rise and flood continental shelves . Most coral reefs are less than 10,000 years old. As communities established themselves, the reefs grew upwards, pacing rising sea levels . Reefs that rose too slowly could become drowned, without sufficient light. Coral reefs are also found in

8075-472: The sill about 8000 BP, forming a marine Littorina Sea , which was followed by another freshwater phase before the present brackish marine system was established. "At its present state of development, the marine life of the Baltic Sea is less than about 4000 years old", Drs. Thulin and Andrushaitis remarked when reviewing these sequences in 2003. Overlying ice had exerted pressure on the Earth's surface. As

8170-480: The surrounding ice sheets. According to the sediment composition retrieved from deep-sea cores , even times of seasonally open waters must have occurred. Outside the main ice sheets, widespread glaciation occurred on the highest mountains of the Alpide belt . In contrast to the earlier glacial stages, the Würm glaciation was composed of smaller ice caps and mostly confined to valley glaciers, sending glacial lobes into

8265-424: The survival of their coral hosts. Each photosynthetic organism has a specific level of sensitivity to photodamage to compounds needed for survival, such as proteins. Rates of regeneration and replication determine the organism's ability to survive. Phylotype A is found more in the shallow waters. It is able to produce mycosporine-like amino acids that are UV resistant , using a derivative of glycerin to absorb

8360-447: The total glaciated area was in the Sierra Nevada de Mérida , and of that amount, the largest concentration, 50 km (19 sq mi), was in the areas of Pico Bolívar , Pico Humboldt [4,942 m (16,214 ft)], and Pico Bonpland [4,983 m (16,348 ft)]. Radiocarbon dating indicates that the moraines are older than 10,000 BP, and probably older than 13,000 BP. The lower moraine level probably corresponds to

8455-410: The undergrowth are analogous to clades B, C, and D. Since clades B through D are found at deeper depths, they require an elevated light absorption rate to be able to synthesize as much energy. With elevated absorption rates at UV wavelengths, these phylotypes are more prone to coral bleaching versus the shallow clade A. Last Glacial Period The Last Glacial Period ( LGP ), also known as

8550-604: The western parts of Jutland were ice-free, and a large part of what is today the North Sea was dry land connecting Jutland with Britain (see Doggerland ). The Baltic Sea , with its unique brackish water , is a result of meltwater from the Weichsel glaciation combining with saltwater from the North Sea when the straits between Sweden and Denmark opened. Initially, when the ice began melting about 10,300 BP, seawater filled

8645-728: The whole western Swiss plateau, reaching today's regions of Solothurn and Aargau. In the region of Bern, it merged with the Aar glacier. The Rhine Glacier is currently the subject of the most detailed studies. Glaciers of the Reuss and the Limmat advanced sometimes as far as the Jura. Montane and piedmont glaciers formed the land by grinding away virtually all traces of the older Günz and Mindel glaciation, by depositing base moraines and terminal moraines of different retraction phases and loess deposits, and by

8740-464: Was at its greatest extent between 23,500 and 21,000 years ago. This glaciation was somewhat distinct from the main Wisconsin glaciation, as it was only loosely related to the giant ice sheets and was instead composed of mountain glaciers, merging into the Cordilleran ice sheet. The Cordilleran ice sheet produced features such as glacial Lake Missoula , which broke free from its ice dam, causing

8835-658: Was more severe than the Last Glacial Period in some areas such as Britain, but less severe in others. The last glacial period saw alternating episodes of glacier advance and retreat with the Last Glacial Maximum occurring between 26,000 and 20,000 years ago. While the general pattern of cooling and glacier advance around the globe was similar, local differences make it difficult to compare the details from continent to continent (see picture of ice core data below for differences). The most recent cooling,

8930-468: Was more widespread. An ice sheet formed in New Zealand, covering all of the Southern Alps, where at least three glacial advances can be distinguished. Local ice caps existed in the highest mountains of the island of New Guinea , where temperatures were 5 to 6 °C colder than at present. The main areas of Papua New Guinea where glaciers developed during the LGP were the Central Cordillera ,

9025-512: Was not covered by a continental-scale ice sheet. Instead, large, but restricted, icefield complexes covered mountain ranges within northeast Siberia, including the Kamchatka-Koryak Mountains. The Arctic Ocean between the huge ice sheets of America and Eurasia was not frozen throughout, but like today, probably was covered only by relatively shallow ice, subject to seasonal changes and riddled with icebergs calving from

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