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126-602: The following phylogeny is based on the YCC 2008 tree and subsequent published research as summarized by ISOGG. Within E-M35, there are striking parallels between two haplogroups, E-V68 and E-V257. Both contain a lineage which has been frequently observed in North Africa (E-M78 and E-M81, respectively) and a group of undifferentiated chromosomes that are mostly found in southern Europe. An expansion of E-M35 carriers, possibly from

252-506: A binary tree ), and an unrooted bifurcating tree takes the form of an unrooted binary tree , a free tree with exactly three neighbors at each internal node. In contrast, a rooted multifurcating tree may have more than two children at some nodes and an unrooted multifurcating tree may have more than three neighbors at some nodes. Both rooted and unrooted trees can be either labeled or unlabeled. A labeled tree has specific values assigned to its leaves, while an unlabeled tree, sometimes called

378-547: A case system with at least two cases. Proto-Afroasiatic may have had marked nominative or ergative-absolutive alignment. A deverbal derivational prefix *mV- is also widely reconstructed. While there is disagreement about the forms of the PAA personal pronouns, there is agreement that there were independent and "bound" (unstressed, clitic ) forms. There is also agreement that a widespread demonstrative pattern of n = masculine and plural, t= feminine goes back to PAA, as well as about

504-599: A dental consonant but does co-occur with other pharyngeal consonants , it must itself have originally been a dental *d in Proto-Afroasiatic, which later became *ʕ in Egyptian. Rössler's ideas have come to dominate the field of Egyptology without, however, achieving general acceptance. Orin Gensler argues that Rössler's sound change is typologically extremely unlikely, though still possible, while many of

630-410: A terminative case in -iš . Scholars debate whether these are vestigial cases or adverbial postpositions . The ending -iš has often been connected to the Egyptian postposition js and is sometimes used to reconstruct a Proto-Afroasiatic locative case. Diakonoff also believed he could reconstruct a comitative - dative case in *-dV or *-Vd , an ablative - comparative case in *-kV ,

756-557: A "directive" case in *-l , and an ablative case in *-p . A prefix mV- is the most widely attested affix in AA that is used to derive nouns. For PAA, its shape has variously been reconstructed as *ma- , *ma(i)- , *mV- , and *-m- . In the daughter languages, it is attested with a wide variety of meanings and functions, such as forming deverbal agent nouns , place nouns, instrument nouns, as well as participles. Erin Shay argues that *mV-

882-409: A clear outgroup. Another method is midpoint rooting, or a tree can also be rooted by using a non-stationary substitution model . Unrooted trees illustrate the relatedness of the leaf nodes without making assumptions about ancestry. They do not require the ancestral root to be known or inferred. Unrooted trees can always be generated from rooted ones by simply omitting the root. By contrast, inferring

1008-814: A combination of genes that come from different genomic sources (e.g., from mitochondrial or plastid vs. nuclear genomes), or genes that would be expected to evolve under different selective regimes, so that homoplasy (false homology ) would be unlikely to result from natural selection. When extinct species are included as terminal nodes in an analysis (rather than, for example, to constrain internal nodes), they are considered not to represent direct ancestors of any extant species. Extinct species do not typically contain high-quality DNA . The range of useful DNA materials has expanded with advances in extraction and sequencing technologies. Development of technologies able to infer sequences from smaller fragments, or from spatial patterns of DNA degradation products, would further expand

1134-554: A consonant; consonants included in the pattern often involve gemination . If root-and-pattern morphology originated in Proto-Afroasiatic, then an explanation must be found for why it has mostly disappeared in the Omotic and Chadic branches; if it was not present in PAA, then an explanation must be found for why it developed independently in the Semitic, Egyptian, and Cushitic branches. Hans-Jürgen Sasse proposed that Proto-Afroasiatic

1260-464: A demonstrative *h- ('this/that') or *ha- ('this/that one'). The most common Afroasiatic interrogative pronoun is *mV , which Ehret reconstructs as *ma , *mi 'what?'. Diakonoff argued that *mV ultimately derived from a demonstrative stem *m- . Only the Semitic reflexes of this root have separate forms for animate ('"who?") and inanimate ("what?") referents. The Old Egyptian and Berber descendants both appear to be used regardless of whether

1386-405: A fact which has not yet been explained. Additionally, it is not always clear which words are cognates, as some proposed cognates may be chance resemblances. Moreover, at least some cognates are likely to have been altered irregularly due to analogical change , making them harder to recognize. As words change meaning over time, the question of which words might have originally meant the same thing

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1512-432: A few branches, making them difficult to reconstruct. In addition to a singular and plural, Egyptian and Semitic attest a dual , the endings of which can be reconstructed respectively as Ancient Egyptian : * -a(y) and Semitic * -ā (nominative) and * -ay (oblique). These endings are very similar to each other, and due to the dual's attestation in the two earliest attested branches of Afroasiatic it

1638-528: A form of the copula 'to be' or a particle meaning 'self'. Afroasiatic languages attest a variety of determiners , only some of which are likely to derive from Proto-Afroasiatic. As first noticed by Joseph Greenberg , Afroasiatic languages in all branches but Omotic attest a series of third person agreement markers in the form n- (masculine), t- (feminine), and n- (plural), which probably derive from Proto-Afroasiatic determiners; Omotic attests t- (feminine) alone of this set. Additionally, Omotic attests

1764-427: A function of the number of tips. For 10 tips, there are more than 34 × 10 6 {\displaystyle 34\times 10^{6}} possible bifurcating trees, and the number of multifurcating trees rises faster, with ca. 7 times as many of the latter as of the former. A dendrogram is a general name for a tree, whether phylogenetic or not, and hence also for the diagrammatic representation of

1890-663: A grammatical rather than a lexical function, and argue that there is thus no basis to reconstruct it as a lexical feature in PAA, as Diakonoff does; they find Ehret's reasoning more sound. Igor Diakonoff argues that Proto-Afroasiatic required a consonant at both the beginning of a syllable and the end of a word, and that only one consonant was possible at the beginning or end of a syllable. Zygmont Frajzyngier and Erin Shay note that these rules appear to be based on Semitic structures, whereas Chadic includes syllables beginning with vowels as well as initial and final consonant clusters. Christopher Ehret argues that all word stems in PAA took

2016-404: A marked nominative language. However, Abdelaziz Allati notes that, if PAA was originally ergative-aligned, it is unclear why both the attested ancient languages and modern AA languages predominantly have nominative-accusative alignment . Proto-Afroasiatic word order has not yet been established. Igor Diakonoff proposed that PAA had verb-subject-object word order (VSO word order), meaning that

2142-438: A masculine agreement form k- , while Chadic and Cushitic show a gender- and number-neutral form k- : both likely go back to a Proto-Afroasiatic determiner *k- , reconstructed by Ehret as *kaa 'this'. Diakonoff argues that in Proto-Afroasiatic these forms were originally demonstrative pronouns that later developed into third person personal pronouns in some branches and into genitive markers in others. Ehret also reconstructs

2268-536: A more reticulate evolutionary history of the organisms sampled. Proto-Afroasiatic Proto-Afroasiatic ( PAA ), also known as Proto-Hamito-Semitic , Proto-Semito-Hamitic , and Proto-Afrasian , is the reconstructed proto-language from which all modern Afroasiatic languages are descended. Though estimations vary widely, it is believed by scholars to have been spoken as a single language around 12,000 to 18,000 years ago (12 to 18 kya ), that is, between 16,000 and 10,000 BC . Although no consensus exists as to

2394-609: A more suitable metaphor than the tree . Indeed, phylogenetic corals are useful for portraying past and present life, and they have some advantages over trees ( anastomoses allowed, etc.). Phylogenetic trees composed with a nontrivial number of input sequences are constructed using computational phylogenetics methods. Distance-matrix methods such as neighbor-joining or UPGMA , which calculate genetic distance from multiple sequence alignments , are simplest to implement, but do not invoke an evolutionary model. Many sequence alignment methods such as ClustalW also create trees by using

2520-536: A number of different formats, all of which must represent the nested structure of a tree. They may or may not encode branch lengths and other features. Standardized formats are critical for distributing and sharing trees without relying on graphics output that is hard to import into existing software. Commonly used formats are Although phylogenetic trees produced on the basis of sequenced genes or genomic data in different species can provide evolutionary insight, these analyses have important limitations. Most importantly,

2646-441: A phylogenetic tree. A cladogram only represents a branching pattern; i.e., its branch lengths do not represent time or relative amount of character change, and its internal nodes do not represent ancestors. A phylogram is a phylogenetic tree that has branch lengths proportional to the amount of character change. A chronogram is a phylogenetic tree that explicitly represents time through its branch lengths. A Dahlgrenogram

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2772-608: A pluralizing morpheme in which a vowel *a was inserted between the two final consonants of the root, possibly replacing another vowel via apophony . However, Paul Newman has argued that while plurals via vowel alteration are frequent in Chadic, they cannot be reconstructed back to Proto-Chadic or Proto-Afroasiatic. Andréas Stauder likewise argues that Coptic and Egyptian plurals via vowel change may have developed independently. Lameen Souag argues that while some form of vowel-changing plural likely goes back to Proto-Afroasiatic, many of

2898-825: A sample of Arabs from Zriba , 89.5% in Andalusians (Qalaat-al-Andalous), and 100% in Berbers from Chenini-Douiret , Jradou and Takrouna. It is generally found at frequencies around 45% in coastal cities of the Maghreb ( Oran , Tunis , Algiers ). It is also prevalent among other Berber populations and reaches frequency of 72.4% in Marrakesh Berbers, 80% in Mozabite , and 71% in Middle Atlas Berbers (Moyen). It also reaches high levels (77.8%) among

3024-427: A set of species or taxa during a specific time. In other words, it is a branching diagram or a tree showing the evolutionary relationships among various biological species or other entities based upon similarities and differences in their physical or genetic characteristics. In evolutionary biology, all life on Earth is theoretically part of a single phylogenetic tree, indicating common ancestry . Phylogenetics

3150-676: A substantial increasing frequency. At the eastern extreme of this core range, M81 is found in 28.6% (10 out of 35 men) in El-Hayez in the Western desert in Egypt The pattern of distribution and variance to be consistent with the hypothesis of a post Paleolithic " demic diffusion " from the Middle East. The ancestral lineage of E-M81 in their hypothesis could have been linked with the spread of Neolithic food-producing technologies from

3276-409: A suffix *-Vb- used to mark harmful animals. Vladimir Orel also attests less well-defined uses for this suffix, while Ehret takes this as a suffix to mark animals and parts of the body. Afroasiatic languages today clearly distinguish singular and plural. One of the first features of Proto-Afroasiatic proposed by Joseph Greenberg was the existence of "internal-a plurals" (a type of broken plural ):

3402-661: A third consonant. As early as the Middle Ages, however, grammarians had noticed that some triradical roots in Arabic differed in only one consonant and had related meanings. According to supporters of original triradicalism such as Gideon Goldenberg, these variations are common in language and inconclusive for the matter. He compares phonetic similarity between words with similar meanings in English such as glow , gleam , glitter , glaze , and glade . Other scholars argue that

3528-483: A tonal system of at least two tonal phonemes, falling tone, rising tone, and possibly a third tone, level tone. Other scholars argue that Proto-AA had a pitch accent and some branches subsequently developed tone. Such scholars postulate that tones developed to compensate for lost or reduced syllables, and note that certain tones are often associated with certain syllable-final consonants. Zygmunt Frajzyngier and Erin Shay note that in AA tonal languages, tone usually has

3654-445: A tree shape, defines a topology only. Some sequence-based trees built from a small genomic locus, such as Phylotree, feature internal nodes labeled with inferred ancestral haplotypes. The number of possible trees for a given number of leaf nodes depends on the specific type of tree, but there are always more labeled than unlabeled trees, more multifurcating than bifurcating trees, and more rooted than unrooted trees. The last distinction

3780-468: A two vowel system ( *a and *ə ), as supported by Berber and Chadic data, and then developing further vowels. Some scholars postulate that Proto-Afroasiatic was a tonal language, with tonality subsequently lost in some branches. Igor Diakonoff argued for the existence of tone based on his reconstruction of many otherwise homophonous words. Christopher Ehret instead takes the fact that three branches of AA have tone as his starting point; he has postulated

3906-400: Is c.  4000 BCE , after which Egyptian and the Semitic languages are firmly attested. However, in all likelihood these languages began to diverge well before this hard boundary. The estimations offered by scholars as to when Proto-Afroasiatic was spoken vary widely, ranging from 18,000   BCE to 8,000   BCE. An estimate at the youngest end of this range still makes Afroasiatic

Haplogroup E-Z827 - Misplaced Pages Continue

4032-609: Is a diagram representing a cross section of a phylogenetic tree. A phylogenetic network is not strictly speaking a tree, but rather a more general graph , or a directed acyclic graph in the case of rooted networks. They are used to overcome some of the limitations inherent to trees. A spindle diagram, or bubble diagram, is often called a romerogram, after its popularisation by the American palaeontologist Alfred Romer . It represents taxonomic diversity (horizontal width) against geological time (vertical axis) in order to reflect

4158-480: Is a later development, which he associates primarily with Semitic. Helmut Satzinger has argued that the earliest form of conjugation in Afroasiatic was the so-called "prefix conjugation," a form found in Semitic, Berber, and Cushitic that uses prefixes to conjugate verbs for person, gender, and number. Other scholars ague that, as there is no evidence for the "prefix conjugation" in Omotic, Chadic, or Egyptian,

4284-629: Is a subclade of E-V1515. It was identified by ISOGG as the second clade within E-Z830. It was discovered before E-Z830 and is associated with the spread of pastoralism from Eastern Africa by South Cushites into Southern Africa . So far high levels have been found in specific ethnic groups in Tanzania and Southern Africa. Highest were the Datog (43%), Khwe (Kxoe) (31%), Burunge (28%), and Sandawe (24%). Two Bantu-speaking Kenyan males were found with

4410-461: Is also accepted by Takács, but he reconstructs it as *ʔaw / *wa 'who?'. Diakonoff also reconstructs an interrogative adjective, *ayyV- , which he claims left traces in Semitic, Cushitic, and Omotic. Lipiński, on the other hand, holds this term to be Semitic and deriving from a particle ʔay 'where?'. Takács derives this particle from PAA *ʔay / *ya , a variant of *ʔaw / *wa 'who?'. Most morphological reconstruction for PAA has focused on

4536-470: Is also debate about whether some of the forms may have been nominal (using verbal nouns), or possibly participial or gerundival , rather than purely verbal. TAMs may have been indicated by both changes in the verb stem and the use of suffixes and prefixes. Some scholars argue that prefixes were used for "eventive" (describing things happening) aspects, as opposed to the "suffix conjugation," which described states. Abdelaziz Allati, however, argues that this

4662-409: Is also possible for forms closer to PAA to be preserved in languages recorded later, while languages recorded earlier may have forms that diverge more from PAA. In order to provide a more accurate reconstruction of Afroasiatic, it will be necessary to first reconstruct the proto-forms of the individual branches, a task which has proven difficult. As of 2023, there is only the beginning of a consensus on

4788-491: Is also usually reconstructed for the proto-language. The loss of the dual in the other branches over time is a well attested feature in languages, including within the Egyptian and Semitic branches themselves. There is widespread agreement that Proto-Afroasiatic had case inflexion . First proposed by Hans-Jürgen Sasse on the basis of his reconstruction of the Proto-Cushitic case system in 1984, Proto-Afroasiatic

4914-425: Is attested among the Semitic languages and may have been dialectal in origin. The forms of the personal pronouns are very stable throughout Afroasiatic (excluding Omotic), but there is no consensus on what the reconstructed set of Afroasiatic pronouns might have looked like. Most modern branches have an independent / absolute pronoun, an object pronoun, and a suffix /possessive pronoun. According to Igor Diakonoff,

5040-614: Is concentrated in North Africa, and is dominated by its E-M183 subclade. E-M183 is believed to have originated in the Northwest of Africa, and has an estimated age of 2284-2984 ybp. The E-M183 sub haplogroup reaches a mean frequency of 42% in North Africa. It decreases in frequency from 100% in some populations to approximately 28.6% to the east of this range in Egypt. The E-M81 subclade is predominant among North African Berber -speaking populations. In Tunisia , it reached 100% frequency among

5166-735: Is defined by V6 and has been identified a significant presence of these lineages in Ethiopia , and also some in the neighboring Somali population. Among the Ethiopian and Somali samples, the highest were 14.7% among the Ethiopian Amhara , and 16.7% among the Ethiopian Wolayta . E-V92 was discovered in two Ethiopian Amhara . Like E-V6 and E-V42 it possibly only exists in the area of Ethiopia. Prior to 2002, there were in academic literature at least seven naming systems for

Haplogroup E-Z827 - Misplaced Pages Continue

5292-706: Is estimated at 10600 years by yfull. Archeology unearthed the remains of a member of the Hungarian conquering elite was analyzed from branch E-FGC19010, it had been discovered in Sandorfalva in Hungary and is dated to the second half of the tenth century. A skeleton was discovered at the Monastery of San Pietro, Villa Magna in Italy, whose DNA belongs to the same branch and lived around 1180CE. Scientists have examined

5418-437: Is evidence for natural gender in all branches, including Omotic, perhaps marked originally by an opposition of PAA *-u (masculine) and *-i (feminine), as also found in the second person singular pronouns . In addition to grammatical gender, Igor Diakonoff argues that Afroasiatic languages show traces of a nominal classification system , which was already unproductive in the Proto-Afroasiatic stage. In particular, he noted

5544-436: Is found in the two oldest attested branches of the family. In the Semitic languages, the "nisba" is used to form adjectives, derive nouns for people associated with a place or profession, and to form hypercoristic names . In Egyptian, it forms adjectives and nouns from nouns and prepositions. The "nisba" is often assumed to be connected to the genitive case ending in Semitic and possibly Cushitic. Igor Diakonoff argued that

5670-429: Is likely that this is inherited from proto-Afroasiatic. Vladimir Orel and Olga Stolbova (1995) reconstruct 32 consonant phonemes, while Christopher Ehret reconstructs 42. Of these, twelve in both reconstructions rely on the same sound correspondences, while an additional eighteen rely on more or less the same sound correspondences. Both reconstructions also include a number of other consonants. While some of these are

5796-506: Is most true of genetic material that is subject to lateral gene transfer and recombination , where different haplotype blocks can have different histories. In these types of analysis, the output tree of a phylogenetic analysis of a single gene is an estimate of the gene's phylogeny (i.e. a gene tree) and not the phylogeny of the taxa (i.e. species tree) from which these characters were sampled, though ideally, both should be very close. For this reason, serious phylogenetic studies generally use

5922-476: Is no agreement about PAA's vowels, the existence of tone , or its syllable structure. At the same time, scholars disagree to whether and to what extent the classical Semitic languages are a conservative, faithful representation of PAA morphology. This is particularly important for the question of whether the lexical roots in the language were originally mostly biradical or triradical , that is, whether they originally had two or three consonants. It also plays into

6048-494: Is no evidence for this in Ancient Egyptian, Cushitic, or Chadic, perhaps indicating that there was no gender distinction in the plural in Proto-Afroasiatic. Chadic has both an inclusive and exclusive form of "we", which Igor Diakonoff and Václav Blažek reconstruct also for Proto-Afroasiatic. Helmut Satzinger has argued that Proto-Afroasiatic only distinguished between the "object" and "possessive" pronouns, deriving

6174-499: Is often difficult to answer. As a result, Robert Ratcliffe suggests that Proto-Afroasiatic may never be reconstructed in the same way that Proto-Indo-European has been. The current state of reconstruction is also hindered by the fact that the Egyptian and Semitic branches of Afroasiatic are attested as early as 3000 BCE, while the languages of the Berber, Chadic, Cushitic, and Omotic branches are only attested much later, sometimes in

6300-494: Is only found in sub-Saharan Africa, indicating a separate phylogenetic history for M35.1 * (former) samples further north". E-P72 . This is a subclade of E-M293. E-V42 was discovered in two Ethiopian Jews . It was suggested that it may be restricted to the region around Ethiopia. However, further testing by commercial DNA testing companies confirmed positive results for this subclade in Arabia as well. The E-V6 subclade of E-V1515

6426-539: Is the most biologically relevant; it arises because there are many places on an unrooted tree to put the root. For bifurcating labeled trees, the total number of rooted trees is: For bifurcating labeled trees, the total number of unrooted trees is: Among labeled bifurcating trees, the number of unrooted trees with n {\displaystyle n} leaves is equal to the number of rooted trees with n − 1 {\displaystyle n-1} leaves. The number of rooted trees grows quickly as

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6552-412: Is the only prefix in the AA phylum that clearly goes back to the proto-language rather than possibly being an areal feature . The precise meaning and origin of this prefix in PAA are debated. There is a long tradition of comparing the prefix to the interrogative pronoun *mā 'who'. Carsten Peust has suggested a common PAA origin for the prefix in forming nouns of place and instrument, but proposes that

6678-443: Is the study of phylogenetic trees. The main challenge is to find a phylogenetic tree representing optimal evolutionary ancestry between a set of species or taxa. Computational phylogenetics (also phylogeny inference) focuses on the algorithms involved in finding optimal phylogenetic tree in the phylogenetic landscape. Phylogenetic trees may be rooted or unrooted. In a rooted phylogenetic tree, each node with descendants represents

6804-500: Is the use of the prefix *ʔan-/*ʔin- , which appears in the Semitic and Old Egyptian first person independent pronouns, the Old Egyptian, Cushitic, and Semitic second person singular and plural pronouns, and the Old Egyptian and Berber third person singular and plural independent pronouns. While Ehret reconstructs this as the original form of the first person singular pronoun, other scholars argue that this element either represents

6930-518: Is usually reconstructed with a case system similar to Proto-Semitic. This gives a nominative ending *-u , accusative or absolutive *-a , and genitive *-i . Besides Proto-Semitic, evidence for these endings is derived from the Cushitic languages and has been argued to exist in Berber as well. The Egyptian nominal ending -w , found on some masculine nouns, may also be evidence of this system. Some evidence for nominative -u may also exist from

7056-456: Is widely agreed to have been present in Proto-Afroasiatic. However, Russell Schuh argues that there was no gender distinction in the plural, as this feature is found only in Semitic and Berber (see also personal pronouns ). Christopher Ehret argues against the consensus that grammatical gender existed in Proto-Afroasiatic, arguing that its development is an isogloss separating all other Afroasiatic languages from Omotic, which alone preserves

7182-409: The *mV- prefix used in agent nouns and participles is actually a post-PAA development, derived from the interrogative pronoun *mā 'who'. Christopher Ehret, meanwhile, proposes that the prefix did not exist in PAA at all, but is a later development from the interrogative pronoun. Gábor Takács and Andrzej Zaborski both reject a connection to *mā entirely; Takács instead suggests that a connection to

7308-680: The Fertile Crescent via the Nile , although pastoralism rather than agriculture . E-M81 and possibly proto-Afroasiatic language may have been carried either all the way from Asia , or they may represent a "local contribution to the North African Neolithic transition". The E-M81 subclade has been found in ancient Guanche (Bimbapes) fossils excavated in Punta Azul, El Hierro , Canary Islands , which are dated to

7434-474: The Moscow School of Comparative Linguistics including Igor Diakonoff and Alexander Militarev includes also *pʼ, *tɬ, *ʃ, *kx⁽ʷ⁾, *gɣ⁽ʷ⁾, *kxʼ⁽ʷ⁾, *x⁽ʷ⁾. Taking Ehret's labialized velars as equivalent to Orel and Stolbova's non-labialized set, and taking Ehret's extra nasals as equivalent to Orel and Stolbova's <n>, the two reconstructions mostly agree on the following correspondences between

7560-743: The Pasiegos from Cantabria , ranging from 18% (8/45) to 41% (23/56). An average frequency of 8.28% (54/652) has also been reported in the Spanish Canary Islands with frequencies over 10% in the three largest islands of Tenerife (10.68%), Gran Canaria (11.54%) and Fuerteventura (13.33%). E-M81 is also found in other parts of Europe, such as Britain – especially Wales and Scotland – and France , where it has an overall incidence of 2.7% (15/555), with frequencies surpassing 5.0% in Auvergne (5/89) and Île-de-France (5/91). E-M81

7686-688: The Tuareg population inhabiting the Sahara in Burkina Faso , near Gor it reaches a much lower frequency of 11.1% in the vicinity of Tanut in the Republic of Niger . In this key area from Egypt to the Atlantic Ocean , report a pattern of decreasing STR haplotype variation (implying decreasing lineage age in those areas) from East to West (but reports West to East for M183), accompanied by

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7812-410: The "nisba" was an "expanded" form of the genitive suffix: he reconstructs the "nisba" suffix as *-iya or -*ī ; he also suggests the existence of a variant *-uwa . Lipiński suggests that the "nisba" originated as a postposition, which was also used to create the genitive case. Christopher Ehret argues that the original form of the suffix was -*iy and also reconstructs a form -*ay . This latter form

7938-929: The 10th century (~44%). Also found in ifri n'ammar that makes the Northwest African origin the likely origin of where it expanded, and not the Middle East. In Europe, E-M81 has a widespread distribution at very low frequencies but is common mostly in the Iberian Peninsula , where unlike in the rest of Europe, shows an average frequency of 4.3% (49/1140) in the Iberian Peninsula with frequencies reaching 4% and 9% in two separate surveys of Galicia , 10% in Western Andalusia and Northwest Castile . However this study also includes 153 individuals from Majorca, Minorca and Ibiza islands as well as 24 individuals from Gascony which are not in

8064-526: The 20th century. The long history of scholarship of the Semitic languages compared to other branches is another obstacle in reconstructing Proto-Afroasiatic; typical features of Semitic have often been projected back to the proto-language, despite their cross-linguistic rarity and lack of correspondences in other branches. Like cognates, shared morphological features tend to disappear over time, as can be demonstrated within Afroasiatic by comparing Old Egyptian (2600–2000 BCE) with Coptic (after 200 CE). Yet it

8190-901: The Americas, this sub-clade is found throughout Latin America , for example 6.1% in Cuba , (8 out of 132), 5.4% in Brazil (Rio de Janeiro) (6 out of 112), "The presence of chromosomes of North African origin (E3b1b-M81; can also be explained by a Portuguese-mediated influx, since this haplogroup reaches a frequency of 5.6% in Portugal , quite similar to the frequency found in Rio de Janeiro (5.4%) among European contributors." and among Hispanic men from California and Hawaii 2.4% (7 out of 295), In smaller numbers, E-M81 men can be found in areas in contact with

8316-577: The DNA of a mass grave of victims of the bubonic plague in Ellwangen in Germany, this one dates from the 16th century and belongs to another branch E-FGC18981. E-V257's dominant sub-clade E-M81 is thought to have originated in the area of the northwest of Africa 7,000 years ago, but all Yfull members are M183 and have a TMRCA just 2700 years ago. E-M81 is the most common subclade of haplogroup E-L19/V257. It

8442-482: The Egyptian preposition m needs further consideration, while Zaborski argues for a connection to a verb *VmV- 'to be'. The term "nisba" refers to a suffix found in the Semitic ( -iy ) and Egyptian ( -j ) branches, with possible relict traces in Berber. A related suffix -āwi occurs in Arabic and possibly Egyptian, as suggested by e.g. ḥmww 'craftsman', from ḥmt 'craft'. Carsten Peust argues that this suffix descends from Proto-Afroasiatic, as it

8568-731: The Iberian Peninsula. Without these 177 individuals, real average for Iberian Peninsula is 4.9% (47/963) it is more common than E-M78, with an average frequency around 5%. Its frequencies are higher in the western half of the peninsula with frequencies reaching 8% in Extremadura and South Portugal, 4% in one study and 9% in another in Galicia , 10% in Western Andalusia and Northwest Castile and 9% to 17% in Cantabria . The highest frequencies of this clade found so far in Europe were observed in

8694-653: The M293 mutation. Other E-M215 subclades are rare in Southern Africa. The authors state... Without information about M293 in the Maasai, Hema, and other populations in Kenya, Sudan, and Ethiopia, we cannot pinpoint the precise geographic source of M293 with greater confidence. However, the available evidence points to present-day Tanzania as an early and important geographic locus of M293 evolution. They also say that "M293

8820-532: The Maghreb, both around the Sahara, in places like Sudan , and around the Mediterranean in places like Lebanon , Turkey , and amongst Sephardic Jews . The following gives a summary of most of the studies which specifically tested for E-M81, showing where its distribution is greater than 1% in Europe, North Africa, the Middle East and Latin America. A recently confirmed sub-clade of E-Z827, Z830, includes

8946-570: The Middle East as proposed by other authors, and split into two branches separated by the geographic barrier of the Mediterranean Sea, would explain this geographic pattern. However, the absence of E-V68* and E-V257* in the Middle East makes a maritime spread between northern Africa and southern Europe a more plausible hypothesis. PF2431 is the sister branch of M81 which was discovered in Paolo Francalacci (2011). Previously, it

9072-413: The Omotic branch. By the evidence of Semitic, in the dual and plural , only the nominative and an oblique were distinguished. David Wilson, on the other hand, argues that the case endings are often not cognate in the individual branches of Afroasiatic and that this precludes their reconstruction for the proto-language. Old Akkadian and Palaeosyrian have two additional cases, a locative in -um and

9198-555: The PAA root may have originally been mostly biradical, to which a third radical was then added. Christopher Ehret argues that the third consonants were derivational affixes, proposing as many as thirty-seven separate verbal extensions that subsequently became fossilized as third consonants. This theory has been criticized by some, such as Andrzej Zaborski and Alan Kaye, as being too many extensions to be realistic, though Zygmont Frajzyngier and Erin Shay note that some Chadic languages have as many as twelve extensions. An alternative model

9324-420: The PAA root was originally biradical but saw the biradical roots outside of Semitic as largely the result of losing a third consonant. Afroasiatic languages feature a "root-and-pattern" ( nonconcatenative ) system of morphology, in which the root consists of consonants alone and vowels are inserted via apophony according to "templates" to create words. A "template" consists of one or more vowels and sometimes

9450-642: The Y-Chromosome Phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed the Y-Chromosome Consortium (YCC). They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being above all timely. The table below brings together all of these works at

9576-458: The addition of a consonant. Not all triradical roots can be convincingly explained as coming from biradicals, and there are cases in which triradical roots with similar meanings appear to differ in one consonant due to root-internal changes or derivation via rhyme. Andréas Stauder argues that the evidence from Ancient Egyptian shows that both tri- and biradical verbs were probably present in Proto-Afroasiatic. Igor Diakonoff, in contrast, argued that

9702-533: The book Elementary Geology , by Edward Hitchcock (first edition: 1840). Charles Darwin featured a diagrammatic evolutionary "tree" in his 1859 book On the Origin of Species . Over a century later, evolutionary biologists still use tree diagrams to depict evolution because such diagrams effectively convey the concept that speciation occurs through the adaptive and semirandom splitting of lineages. The term phylogenetic , or phylogeny , derives from

9828-475: The central vowels *e and *o could not occur together in the same root. Taking a different approach, Ronny Meyer and H. Ekkehard Wolff propose that Proto-Afroasiatic may have had no vowels as such, instead employing various syllabic consonants (*l, *m, *n, *r) and semivowels or semivowel-like consonants (*w, *y, *ʔ, *ḥ, *ʕ, *h, *ʔʷ, *ḥʷ, *ʕʷ, *hʷ) to form syllables; vowels would have later been inserted into these syllables ("vocalogenesis"), developing first into

9954-473: The confirmed sub-clades of E-M123 , E-M293, and E-V42, and is a sibling clade to E-L19. Currently, the E-M35 phylogeny project recognizes four distinct clusters of Z830* carriers, two of which are exclusively Jewish in origin . The remaining two are significantly smaller, and include scattered individuals in Germany, Spain, Latin America, Egypt, and Ethiopia. E-M123 is mostly known for its major subclade E-M34, which dominates this clade. A new clade (E-V1515)

10080-473: The different branches of Afroasiatic: Additionally, there is another proposal for the sound correspondences between – and phonetic values of – Egyptian and Semitic consonants. This second theory is known as neuere Komparatistik and was first proposed by Semiticist Otto Rössler on the basis of consonant incompatibilities . In particular, Rössler argued that, since the hieroglyph conventionally transcribed as <ʿ> and described as *ʕ never co-occurs with

10206-427: The difficulty in reconstruction is likely related to the use of vowel changes known as apophony (or "ablaut") in the "root-and-pattern" system found in various Afroasiatic languages. In addition to apophony, some modern AA languages display vowel changes referred to as umlaut . Igor Diakonoff, Viktor Porkhomovksy and Olga Stolbova proposed in 1987 that Proto-Afroasiatic had a two vowel system of *a and *ə , with

10332-442: The etymologies proposed in support of the theory have been attacked by Gábor Takács. The most important sound correspondences in the neuere Komparatistik that differ from the traditional understanding are: Attempts to reconstruct the vocalic system of Proto-Afroasiatic vary considerably. While there is no consensus, many scholars prefer to reconstruct a simple three vowel system with long and short *a , *i , and *u . Some of

10458-444: The existence of an interrogative pronoun *mV , which may not have distinguished animacy . There is some agreement that the PAA verb had two or possibly three basic forms, though there is disagreement about what those forms were and what tenses, aspects, or moods they expressed. There is also widespread agreement that there were possibly two sets of conjugational affixes (prefixes and suffixes) used for different purposes. Additionally,

10584-450: The forms of the pronouns in the other branches show evidence of marked nominative alignment. Igor Diakonoff instead argued that Proto-Afroasiatic was an ergative-absolutive language, in which the ergative case marks the subject of transitive verbs and the absolutive case marks both the object of transitive verbs and the subject of intransitive verbs. Satzinger suggests that Proto-Afroasiatic may have developed from ergative-absolutive to

10710-458: The great amount of time since Afroasiatic split into branches, there are limits to what scholars can reconstruct. Cognates tend to disappear from related languages over time. There are currently not many widely accepted Afroasiatic cognates, and it is difficult to derive sound correspondence rules from a small number of examples. The most convincing cognates in Afroasiatic often have the same or very similar consonants but very different vowels,

10836-427: The importance of verbal gemination and reduplication and the existence of three derivational affixes, especially of a causative -*s-, are commonly reconstructed. A numeral system cannot be reconstructed, although numerous PAA numerals and cognate sets from 1 to 9 have been proposed. There is no consensus as to when Proto-Afroasiatic was spoken. The absolute latest date for when Proto-Afroasiatic could have been extant

10962-450: The independent pronouns via various processes in the branches. He argues that the independent pronouns derive from various strategies combining pronominal elements with different nominal or pronominal bases. Václav Blažek reconstructs an original set of independent pronouns but argues that the ones found in most current Afroasiatic languages arose by a process of suppletion similar to that argued by Satzinger. An example of one such process

11088-561: The individual daughter languages. Most reconstructions agree that PAA had three series of obstruents ( plosives , fricatives , and affricates ) and that the continuants were all voiceless. There is also general agreement that obstruents were organized in triads of voiceless, voiced, and "emphatic" (possibly glottalized ) consonants, and that PAA included pharyngeal and laryngeal consonants . Disagreement exists about whether there were labialized velar consonants. Several Afroasiatic languages have large consonant inventories, and it

11214-412: The inferred most recent common ancestor of those descendants, and the edge lengths in some trees may be interpreted as time estimates. Each node is called a taxonomic unit. Internal nodes are generally called hypothetical taxonomic units, as they cannot be directly observed. Trees are useful in fields of biology such as bioinformatics , systematics , and phylogenetics . Unrooted trees illustrate only

11340-433: The later realized as [i] or [u] depending on its contact with labial or labialized consonants . Christopher Ehret has proposed a five vowel system with long and short *a , *e , *o , *i , and *u , arguing that his reconstruction is supported by the Chadic and Cushitic vowels. Vladimir Orel and Olga Stolbova instead proposed a six vowel system with *a , *e , *o , *i , *ü ([ y ]), and *u ; they further argued that

11466-528: The location of the Afroasiatic homeland , the putative homeland of Proto-Afroasiatic speakers, the majority of scholars agree that it was located within a region of Northeast Africa . The reconstruction of Proto-Afroasiatic is problematic and has not progressed to the degree found in Indo-European linguistics . The immense amount of time over which the branches have been separated, coupled with

11592-452: The majority of the deepest E-V1515 subhaplogroups and paragroups were found. In the southern part of the Horn (southern Ethiopia, Somalia and northern Kenya), haplogroup E-V1515 is almost exclusively represented by the recent (3.5 ka; 95% CI: 1.7–5.9 ka) subhaplogroup E-V1486. Further south, in southern Kenya and southern Africa, a single E-V1486 terminal clade, known as E-M293 (Henn et al. 2008),

11718-485: The oldest proven language family. Contrasting proposals of an early emergence, Tom Güldemann has argued that less time may have been required for the divergence than is usually assumed, as it is possible for a language to rapidly restructure due to areal contact , with the evolution of Chadic (and likely also Omotic) serving as pertinent examples. At present, there is no commonly accepted reconstruction of Afroasiatic morphology, grammar, syntax, or phonology. Because of

11844-439: The optimal tree using many of these techniques is NP-hard , so heuristic search and optimization methods are used in combination with tree-scoring functions to identify a reasonably good tree that fits the data. Tree-building methods can be assessed on the basis of several criteria: Tree-building techniques have also gained the attention of mathematicians. Trees can also be built using T-theory . Trees can be encoded in

11970-531: The original gender system of Afroasiatic had masculine endings *-y/*-w (later *-Vy / *-Vw ) and feminine endings *-H/*-y (later *-āʔ / *-āy ), the later of which was later ousted by feminine *-(a)t on nouns. Marijn van Putten has reconstructed a feminine ending *-ay/*-āy from Semitic and Berber evidence: he argues that this ending comes down from the last common ancestor of Berber and Semitic, which may be Proto-Afroasiatic. Despite arguing that Proto-Afroasiatic had no grammatical gender, Ehret argues that there

12096-465: The original, genderless grammar of the proto-language. Other scholars such as Lionel Bender argue that Omotic has lost grammatical gender despite originally having had it. A feminine morpheme -Vt is found widely in Afroasiatic languages. Lameen Souag argues that this feminine ending -t is probably a case of a grammaticalized demonstrative , as this feature has also independently developed in some Chadic and Cushitic languages. Diakonoff argued that

12222-403: The parent of all other nodes in the tree. The root is therefore a node of degree 2, while other internal nodes have a minimum degree of 3 (where "degree" here refers to the total number of incoming and outgoing edges). The most common method for rooting trees is the use of an uncontroversial outgroup —close enough to allow inference from trait data or molecular sequencing, but far enough to be

12348-458: The point of the landmark 2002 YCC Tree. This allows a researcher reviewing older published literature to quickly move between nomenclatures. The following research teams per their publications were represented in the creation of the YCC Tree. Phylogeny A phylogenetic tree , phylogeny or evolutionary tree is a graphical representation which shows the evolutionary history between

12474-406: The prefix conjugation may be a shared innovation in Semitic, Berber, and Cushitic. In those languages where it appears, the "prefix conjugation" is used with two stems, with Igor Diakonoff identifying one as perfective/punctual as well as jussive, and the other with the imperfective. These stems may also be known as "short form" (=perfective) and "long form" (=imperfective). Assuming a PAA origin,

12600-399: The prefixes can be reconstructed as agreeing with the forms of the "bound" personal pronouns in having *n- for first person plural, *t- for second person plural and singular and feminine third person singular, and *y/*i- for third person masculine and third person plural; the form of the first person singular is unclear, but may be *ʔ- . The prefixes may have originally developed from

12726-478: The question of the degree to which Proto-Afroasiatic had root-and-pattern morphology , as most fully displayed in the Semitic , Egyptian , and Cushitic branches. There are nonetheless some items of agreement and reconstructed vocabulary. Most scholars agree that Proto-Afroasiatic nouns had grammatical gender , at least two and possibly three grammatical numbers (singular, plural, and possibly dual ), as well as

12852-405: The range of DNA considered useful. Phylogenetic trees can also be inferred from a range of other data types, including morphology, the presence or absence of particular types of genes, insertion and deletion events – and any other observation thought to contain an evolutionary signal. Phylogenetic networks are used when bifurcating trees are not suitable, due to these complications which suggest

12978-420: The reconstruction of Proto-Semitic , and no widely accepted reconstruction of any of the other branches' proto-forms. Current attempts at reconstructing Afroasiatic often rely on comparing individual words or features in the daughter languages, which leads to results that are not convincing to many scholars. There is currently no consensus on the consonant phonemes of Afroasiatic or on their correspondences in

13104-443: The referent is a person or thing. It is therefore not clear if this pronoun differentiated animacy in Proto-Afroasiatic. Lack of differentiation between "who?" and "what?" is also sporadically attested in Semitic and Cushitic, but appears to be absent in Chadic; most modern AA languages use different lexical roots to make the distinction. Ehret also reconstructs a second interrogative *wa-/*wi- 'what?'. The PAA origin of this form

13230-479: The relatedness of the leaf nodes and do not require the ancestral root to be known or inferred. The idea of a tree of life arose from ancient notions of a ladder-like progression from lower into higher forms of life (such as in the Great Chain of Being ). Early representations of "branching" phylogenetic trees include a "paleontological chart" showing the geological relationships among plants and animals in

13356-574: The root of an unrooted tree requires some means of identifying ancestry. This is normally done by including an outgroup in the input data so that the root is necessarily between the outgroup and the rest of the taxa in the tree, or by introducing additional assumptions about the relative rates of evolution on each branch, such as an application of the molecular clock hypothesis . Both rooted and unrooted trees can be either bifurcating or multifurcating. A rooted bifurcating tree has exactly two descendants arising from each interior node (that is, it forms

13482-768: The same, they rely on correspondences in the daughter languages which cannot be reconciled. For instance, although both Ehret and Orel and Stolbova reconstruct *tʼ , Ehret gives its Egyptian correspondence as s , while Orel and Stolbova give it as d and t ; and though both reconstruct PAA *tlʼ , Ehret gives its Arabic correspondence as ṣ , while Orel and Stolbova give it as ḍ . Additionally, Ehret has reconstructed 11 consonants not found in Orel and Stolbova, while Orel and Stolbova have reconstructed 2 not found in Ehret. The additional consonants are: An earlier, larger reconstruction from 1992 by Orel, Stolbova and other collaborators from

13608-477: The shape CV (with a possible alternate form VC) and CVC, with suffixes often giving the syllabic shape CVCC. David Wilson agrees with Diakonoff that the root syllable could only begin with a single consonant, but adds a requirement that syllables have two mora weight and argues for the possibility of an extra-syllabic consonant at the end of a root (CVC-C or CV:C). The degree to which the Proto-AA verbal root

13734-417: The simpler algorithms (i.e. those based on distance) of tree construction. Maximum parsimony is another simple method of estimating phylogenetic trees, but implies an implicit model of evolution (i.e. parsimony). More advanced methods use the optimality criterion of maximum likelihood , often within a Bayesian framework , and apply an explicit model of evolution to phylogenetic tree estimation. Identifying

13860-620: The suffix/possessive pronoun was originally used as the object of verbs and to show a possessive relationship, the "independent" pronoun served to show emphasis, and the "object" pronoun was used to mark the subject of intransitive verbs and the direct object of transitive verbs. All Afroasiatic branches differentiate between masculine and feminine third person singular pronouns, and all except for Cushitic and Omotic also differentiate between second person singular masculine and feminine pronouns. Semitic and Berber also differentiate between masculine and feminine second and third person plural, but there

13986-663: The templates found in the branches likely do not. Several Afroasiatic languages of the Semitic, Chadic, and Cushitic branches attest pluralization via reduplication , a feature which has often been assumed to go back to Proto-Afroasiatic. Robert Ratcliffe has instead argued that this reduplicating pattern originated after PAA, as a way to allow biradical nouns to insert "internal-a," a process which then became generalized to other roots in some languages; as an alternative hypothesis, they may have developed from forms with plural suffixes. Afroasiatic languages also use several pluralizing affixes – few of these, however, are present in more than

14112-411: The tree before hybridisation takes place, and conserved sequences . Also, there are problems in basing an analysis on a single type of character, such as a single gene or protein or only on morphological analysis, because such trees constructed from another unrelated data source often differ from the first, and therefore great care is needed in inferring phylogenetic relationships among species. This

14238-529: The trees that they generate are not necessarily correct – they do not necessarily accurately represent the evolutionary history of the included taxa. As with any scientific result, they are subject to falsification by further study (e.g., gathering of additional data, analyzing the existing data with improved methods). The data on which they are based may be noisy ; the analysis can be confounded by genetic recombination , horizontal gene transfer , hybridisation between species that were not nearest neighbors on

14364-422: The two ancient greek words φῦλον ( phûlon ), meaning "race, lineage", and γένεσις ( génesis ), meaning "origin, source". A rooted phylogenetic tree (see two graphics at top) is a directed tree with a unique node — the root — corresponding to the (usually imputed ) most recent common ancestor of all the entities at the leaves of the tree. The root node does not have a parent node, but serves as

14490-407: The typical Maghrebin core haplotype 13-14-30-24-9-11-13 has been found in only two out of the five E-M81 individuals. These results, along with the negligible contribution from North-African populations revealed by the admixture-like plot analysis, suggest only a marginal impact of trans-Mediterranean gene flows on the current SSI genetic pool. As a result of Spanish and Portuguese colonization of

14616-404: The variation of abundance of various taxa through time. A spindle diagram is not an evolutionary tree: the taxonomic spindles obscure the actual relationships of the parent taxon to the daughter taxon and have the disadvantage of involving the paraphyly of the parental group. This type of diagram is no longer used in the form originally proposed. Darwin also mentioned that the coral may be

14742-745: The verb would come first in most sentences. Carsten Peust likewise supports VSO word order, as this is found in the two oldest attested branches, Egyptian and Semitic. However, Ronny Meyer and H. Ekkehard Wolff argue that this proposal does not concord with Diakonoff's suggestion that PAA was an ergative-absolutive language, in which subject and object are not valid categories. Zygmont Frajzyngier and Erin Shay further note that, if Proto-Afroasiatic had VSO word order, then an explanation must be found for why two of its branches, Omotic and Cushitic, show subject–object–verb word order (SOV word order). Both sets of scholars argue that this area needs more research. A system of sex-based male and female grammatical gender

14868-404: The verb, with categories found in Semitic languages such as aspect , voice , and person . There is little agreement about which tenses, aspects, or moods ( TAMs ) Proto-Afroasiatic might have had: it may have had two basic forms (indicative vs. subjunctive, state vs. action, transitive vs. intransitive, or perfective vs. imperfective) or three (unmarked vs. perfective vs. imperfective). There

14994-417: The wide gap between the attestations of the original branches (3rd millennium BC for Egyptian and Semitic, 19th and 20th centuries for many Chadic , Cushitic , and Omotic languages ) mean that determining sound correspondences has not yet been possible. In addition to more traditional proposed consonant correspondences, there is also a divergent proposal that has become popular among Egyptologists ; there

15120-400: Was a marked nominative language, in which the nominative case is only used to mark the subject of a verb, whereas an absolutive case is the citation form of the noun and also marks the object. Evidence for marked nominative alignment comes primarily from the use of cases in Cushitic and the so-called "states" of the noun in Berber languages; additionally, Helmut Satzinger has argued that

15246-729: Was also observed in Italy with frequencies of 0,7% to 5,8% in Sardinia , approximately 2.12% overall in Sicily (but up to 7.14% in Piazza Armerina ), and in very much lower frequency near Lucera (1.7%), in continental Italy , possibly due to ancient migrations during the Islamic , Roman , and Carthaginian empires. In a 2014 study by Stefania Sarno et al. with 326 samples from Cosenza , Reggio Calabria , Lecce and five Sicilian provinces, E-M81 shows an average frequency of 1.53%, but

15372-471: Was defined by Trombetta et al. 2015, which originated about 12 kya (95% CI 8.6-16.4) in eastern Africa where it is currently mainly distributed. This clade includes all the sub-Saharan haplogroups (E-V42, E-M293, E-V92, E-V6) reported as E-M35 basal clades in a previous phylogeny. We observed the highest frequency and diversity of this haplogroup in the northern part of the Horn of Africa (present day Eritrea and northern Ethiopia ), where

15498-618: Was designated L19*/V257*. This mutation has been discovered in North Africa (in Souss in Morocco, in central and eastern Algeria, West Nile in Egypt), the Sahel (Chad, Gambia), Western Europe (United Kingdom (Derbyshire), Germany, Switzerland, Spain, Italy) and Near Eastern (Turkey, Karabakh and Urmia). It would have formed 13800 years ago and is thought to originate from the "green" Sahara . Its TMRCA

15624-495: Was found (fig. 3). This phylogeographic pattern is strongly suggestive of human movements from the northern part of the Horn to the Ethiopian/Kenyan borders between 12 ka (the coalescence of E-V1515) and 3.5 ka (the coalescence of E-V1486), and from here toward southern Africa across the equatorial belt in more recent times. Multiple instances of commercially observed E-V1515 have also been detected in Arabia. E-M293

15750-404: Was originally triradical (having three consonants) or biradical (having two consonants) is debated. Among the modern branches, most Semitic roots are triradical, whereas most Chadic, Omotic, and Cushitic roots are biradical. The "traditional theory" argues for original triradicalism in the family, as is the case in Semitic. In this theory, almost all biradical roots are the result of the loss of

15876-536: Was proposed by Georges Bohas , who argued that the third consonants were added to differentiate roots of similar meaning but without the third consonant having a particular meaning itself. Biradical verbs may also have been made triradical on the model of so-called "weak verbs," which have a final radical y or w . Many scholars do not argue for the original nature of either biradical or triradical roots, instead arguing that there are original triradical roots, original biradical roots, and triradical roots resulting from

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