173-407: A forest is an ecosystem characterized by a dense community of trees . Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' Food and Agriculture Organization (FAO) defines a forest as, "Land spanning more than 0.5 hectares with trees higher than 5 meters and
346-423: A canopy cover of more than 10 percent, or trees able to reach these thresholds in situ . It does not include land that is predominantly under agricultural or urban use." Using this definition, Global Forest Resources Assessment 2020 found that forests covered 4.06 billion hectares (10.0 billion acres; 40.6 million square kilometres; 15.7 million square miles), or approximately 31 percent of
519-409: A cell wall . Newly dead animals may be covered by an exoskeleton . Fragmentation processes, which break through these protective layers, accelerate the rate of microbial decomposition. Animals fragment detritus as they hunt for food, as does passage through the gut. Freeze-thaw cycles and cycles of wetting and drying also fragment dead material. The chemical alteration of the dead organic matter
692-421: A farm can stabilize and enhance the provision of pollination services. The presence of such ecosystem elements functions almost like an insurance policy for farmers. Coastal and estuarine ecosystems act as buffer zones against natural hazards and environmental disturbances, such as floods, cyclones, tidal surges and storms. The role they play is to "[absorb] a portion of the impact and thus [lessen] its effect on
865-495: A food chain . Real systems are much more complex than this—organisms will generally feed on more than one form of food, and may feed at more than one trophic level. Carnivores may capture some prey that is part of a plant-based trophic system and others that are part of a detritus-based trophic system (a bird that feeds both on herbivorous grasshoppers and earthworms, which consume detritus). Real systems, with all these complexities, form food webs rather than food chains which present
1038-619: A habitat . Ecosystem ecology is the "study of the interactions between organisms and their environment as an integrated system". The size of ecosystems can range up to ten orders of magnitude , from the surface layers of rocks to the surface of the planet. The Hubbard Brook Ecosystem Study started in 1963 to study the White Mountains in New Hampshire . It was the first successful attempt to study an entire watershed as an ecosystem. The study used stream chemistry as
1211-495: A Bayesian decision support system to both model the uncertainty in the scientific information Bayes Nets and to assist collecting and fusing the input from stakeholders. This study was about siting wave energy devices off the Oregon Coast, but presents a general method for managing uncertain spatial science and stakeholder information in a decision making environment. Remote sensing data and analyses can be used to assess
1384-609: A central role over a wide range, for example, in the slow development of soil from bare rock and the faster recovery of a community from disturbance . Disturbance also plays an important role in ecological processes. F. Stuart Chapin and coauthors define disturbance as "a relatively discrete event in time that removes plant biomass". This can range from herbivore outbreaks, treefalls, fires, hurricanes, floods, glacial advances , to volcanic eruptions . Such disturbances can cause large changes in plant, animal and microbe populations, as well as soil organic matter content. Disturbance
1557-462: A combination of measures aimed at increasing forest carbon stocks, andsustainable timber offtake will generate the largest carbon sequestration benefit. The term forest-dependent people is used to describe any of a wide variety of livelihoods that are dependent on access to forests, products harvested from forests, or ecosystem services provided by forests, including those of Indigenous peoples dependent on forests. In India , approximately 22 percent of
1730-507: A critical role in global nutrient cycling and ecosystem function. Phosphorus enters ecosystems through weathering . As ecosystems age this supply diminishes, making phosphorus-limitation more common in older landscapes (especially in the tropics). Calcium and sulfur are also produced by weathering, but acid deposition is an important source of sulfur in many ecosystems. Although magnesium and manganese are produced by weathering, exchanges between soil organic matter and living cells account for
1903-405: A different classification of forest vegetation is often used: tree, shrub, herb, and moss layers (see stratification (vegetation) ). Forests are classified differently and to different degrees of specificity. One such classification is in terms of the biomes in which they exist, combined with leaf longevity of the dominant species (whether they are evergreen or deciduous ). Another distinction
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#17327810614162076-517: A faster recovery. More severe and more frequent disturbance result in longer recovery times. From one year to another, ecosystems experience variation in their biotic and abiotic environments. A drought , a colder than usual winter, and a pest outbreak all are short-term variability in environmental conditions. Animal populations vary from year to year, building up during resource-rich periods and crashing as they overshoot their food supply. Longer-term changes also shape ecosystem processes. For example,
2249-436: A few main pathways, including increase in commercial tree plantations, adoption of agroforestry techniques by small farmers, or spontaneous regeneration when former agricultural land is abandoned. It can be motivated by the economic benefits of forests, the ecosystem services forests provide, or cultural changes where people increasingly appreciate forests for their spiritual, aesthetic, or otherwise intrinsic value. According to
2422-409: A forest may be of a great variety of species (as in tropical rainforests and temperate deciduous forests ), or relatively few species over large areas (e.g., taiga and arid montane coniferous forests). The biodiversity of forests also encompasses shrubs , herbaceous plants, mosses , ferns , lichens , fungi , and a variety of animals . Trees rising up to 35 meters (115 ft) in height add
2595-549: A forested area by cutting or burning, either to harvest timber or to make way for farming. Most deforestation today occurs in tropical forests. The vast majority of this deforestation is because of the production of four commodities: wood , beef , soy , and palm oil . Over the past 2,000 years, the area of land covered by forest in Europe has been reduced from 80% to 34%. Large areas of forest have also been cleared in China and in
2768-454: A form that can be readily used by plants and microbes. Ecosystems provide a variety of goods and services upon which people depend, and may be part of. Ecosystem goods include the "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like
2941-427: A function-based typology has been proposed to leverage the strengths of these different approaches into a unified system. Human activities are important in almost all ecosystems. Although humans exist and operate within ecosystems, their cumulative effects are large enough to influence external factors like climate. Ecosystems provide a variety of goods and services upon which people depend. Ecosystem goods include
3114-698: A general level, for example, tropical forests , temperate grasslands , and arctic tundra . There can be any degree of subcategories among ecosystem types that comprise a biome, e.g., needle-leafed boreal forests or wet tropical forests. Although ecosystems are most commonly categorized by their structure and geography, there are also other ways to categorize and classify ecosystems such as by their level of human impact (see anthropogenic biome ), or by their integration with social processes or technological processes or their novelty (e.g. novel ecosystem ). Each of these taxonomies of ecosystems tends to emphasize different structural or functional properties. None of these
3287-492: A legal term, as seen in Latin texts such as Magna Carta , to denote uncultivated land that was legally designated for hunting by feudal nobility (see royal forest ). These hunting forests did not necessarily contain any trees. Because that often included significant areas of woodland, "forest" eventually came to connote woodland in general, regardless of tree density. By the beginning of the fourteenth century, English texts used
3460-665: A local ecosystem. The mixture of fresh water and salt water ( brackish water ) in estuaries provides many nutrients for marine life . Salt marshes , mangroves and beaches also support a diversity of plants, animals and insects crucial to the food chain . The high level of biodiversity creates a high level of biological activity, which has attracted human activity for thousands of years. Coasts also create essential material for organisms to live by, including estuaries, wetland , seagrass , coral reefs , and mangroves. Coasts provide habitats for migratory birds , sea turtles, marine mammals, and coral reefs. There are questions regarding
3633-533: A lot of tourists also travel to resorts close to the sea or rivers or lakes to be able to experience these activities, and relax near the water. The United Nations Sustainable Development Goal 14 also has targets aimed at enhancing the use of ecosystem services for sustainable tourism especially in Small Island Developing States . Estuarine and marine coastal ecosystems are both marine ecosystems . Together, these ecosystems perform
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#17327810614163806-672: A means of monitoring ecosystem properties, and developed a detailed biogeochemical model of the ecosystem. Long-term research at the site led to the discovery of acid rain in North America in 1972. Researchers documented the depletion of soil cations (especially calcium) over the next several decades. Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Studies can be carried out at
3979-528: A more important role in moving nutrients around. This can be especially important as the soil thaws in the spring, creating a pulse of nutrients that become available. Decomposition rates are low under very wet or very dry conditions. Decomposition rates are highest in wet, moist conditions with adequate levels of oxygen. Wet soils tend to become deficient in oxygen (this is especially true in wetlands ), which slows microbial growth. In dry soils, decomposition slows as well, but bacteria continue to grow (albeit at
4152-406: A number of common, non random properties in the topology of their network. The carbon and nutrients in dead organic matter are broken down by a group of processes known as decomposition. This releases nutrients that can then be re-used for plant and microbial production and returns carbon dioxide to the atmosphere (or water) where it can be used for photosynthesis. In the absence of decomposition,
4325-491: A nutritional resource is not limited to low- and middle-income countries; more than 100 million people in the European Union (EU) regularly consume wild food. Some 2.4 billion people – in both urban and rural settings – use wood-based energy for cooking. Regulating services are the "benefits obtained from the regulation of ecosystem processes". These include: An example for water purification as an ecosystem service
4498-421: A process known as denitrification . Mycorrhizal fungi which are symbiotic with plant roots, use carbohydrates supplied by the plants and in return transfer phosphorus and nitrogen compounds back to the plant roots. This is an important pathway of organic nitrogen transfer from dead organic matter to plants. This mechanism may contribute to more than 70 Tg of annually assimilated plant nitrogen, thereby playing
4671-580: A region, releasing water from their leaves in anticipation of seasonal rains to trigger the wet season early. Because of this, seasonal rainfall in the Amazon begins two to three months earlier than the climate would otherwise allow. Deforestation in the Amazon and anthropogenic climate change hold the potential to interfere with this process, causing the forest to pass a threshold where it transitions into savanna. Deforestation threatens many forest ecosystems. Deforestation occurs when humans remove trees from
4844-696: A significant portion of ecosystem fluxes. Potassium is primarily cycled between living cells and soil organic matter. Biodiversity plays an important role in ecosystem functioning. Ecosystem processes are driven by the species in an ecosystem, the nature of the individual species, and the relative abundance of organisms among these species. Ecosystem processes are the net effect of the actions of individual organisms as they interact with their environment. Ecological theory suggests that in order to coexist, species must have some level of limiting similarity —they must be different from one another in some fundamental way, otherwise, one species would competitively exclude
5017-408: A slower rate) even after soils become too dry to support plant growth. Ecosystems are dynamic entities. They are subject to periodic disturbances and are always in the process of recovering from past disturbances. When a perturbation occurs, an ecosystem responds by moving away from its initial state. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance,
5190-464: A small effect on ecosystem function. Ecologically distinct species, on the other hand, have a much larger effect. Similarly, dominant species have a large effect on ecosystem function, while rare species tend to have a small effect. Keystone species tend to have an effect on ecosystem function that is disproportionate to their abundance in an ecosystem. An ecosystem engineer is any organism that creates, significantly modifies, maintains or destroys
5363-526: A system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Biomes are general classes or categories of ecosystems. However, there
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5536-522: A variety of scales, ranging from whole-ecosystem studies to studying microcosms or mesocosms (simplified representations of ecosystems). American ecologist Stephen R. Carpenter has argued that microcosm experiments can be "irrelevant and diversionary" if they are not carried out in conjunction with field studies done at the ecosystem scale. In such cases, microcosm experiments may fail to accurately predict ecosystem-level dynamics. Biomes are general classes or categories of ecosystems. However, there
5709-719: A vertical dimension to the area of land that can support plant and animal species, opening up numerous ecological niches for arboreal animal species, epiphytes , and various species that thrive under the regulated microclimate created under the canopy. Forests have intricate three-dimensional structures that increase in complexity with lower levels of disturbance and greater variety of tree species. The biodiversity of forests varies considerably according to factors such as forest type, geography, climate, and soils – in addition to human use. Most forest habitats in temperate regions support relatively few animal and plant species, and species that tend to have large geographical distributions, while
5882-410: A wide range of characteristics. Generally, richer households derive more cash value from forest resources, whereas among poorer households, forest resources are more important for home consumption and increase community resilience. Forests are fundamental to the culture and livelihood of indigenous people groups that live in and depend on forests, many of which have been removed from and denied access to
6055-527: Is a peculiar English spelling of the Latin silva , denoting a "woodland", and has precedent in English, including its plural forms. While its use as a synonym of forest , and as a Latinate word denoting a woodland, may be admitted; in a specific technical sense it is restricted to denoting the species of trees that comprise the woodlands of a region, as in its sense in the subject of silviculture . The resorting to sylva in English indicates more precisely
6228-458: Is an international synthesis by over 1000 of the world's leading biological scientists that analyzes the state of the Earth's ecosystems and provides summaries and guidelines for decision-makers. The report identified four major categories of ecosystem services: provisioning, regulating, cultural and supporting services. It concludes that human activity is having a significant and escalating impact on
6401-661: Is as follows: In New York City , where the quality of drinking water had fallen below standards required by the U.S. Environmental Protection Agency (EPA) , authorities opted to restore the polluted Catskill Watershed that had previously provided the city with the ecosystem service of water purification. Once the input of sewage and pesticides to the watershed area was reduced, natural abiotic processes such as soil absorption and filtration of chemicals, together with biotic recycling via root systems and soil microorganisms , water quality improved to levels that met government standards. The cost of this investment in natural capital
6574-400: Is based on tree densities measured on over 400,000 plots. It remains subject to a wide margin of error, not least because the samples are mainly from Europe and North America. Forests can also be classified according to the amount of human alteration. Old-growth forest contains mainly natural patterns of biodiversity in established seral patterns, and they contain mainly species native to
6747-442: Is consumed by animals while still alive and enters the plant-based trophic system. After plants and animals die, the organic matter contained in them enters the detritus-based trophic system. Ecosystem respiration is the sum of respiration by all living organisms (plants, animals, and decomposers) in the ecosystem. Net ecosystem production is the difference between gross primary production (GPP) and ecosystem respiration. In
6920-423: Is controlled by internal factors. Therefore, internal factors not only control ecosystem processes but are also controlled by them. Ecosystems are dynamic entities—they are subject to periodic disturbances and are always in the process of recovering from some past disturbance. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance, is termed its resistance . The capacity of
7093-538: Is expected to cause a 9% decline in ecosystem services on average at global scale by 2100 Ecosystem-based adaptation (EBA or EbA) encompasses a broad set of approaches to adapt to climate change . They all involve the management of ecosystems and their services to reduce the vulnerability of human communities to the impacts of climate change . The Convention on Biological Diversity (CBD) defines EBA as "the use of biodiversity and ecosystem services as part of an overall adaptation strategy to help people to adapt to
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7266-569: Is followed by succession, a "directional change in ecosystem structure and functioning resulting from biotically driven changes in resource supply." The frequency and severity of disturbance determine the way it affects ecosystem function. A major disturbance like a volcanic eruption or glacial advance and retreat leave behind soils that lack plants, animals or organic matter. Ecosystems that experience such disturbances undergo primary succession . A less severe disturbance like forest fires, hurricanes or cultivation result in secondary succession and
7439-554: Is governed by three sets of factors—the physical environment (temperature, moisture, and soil properties), the quantity and quality of the dead material available to decomposers, and the nature of the microbial community itself. Temperature controls the rate of microbial respiration; the higher the temperature, the faster the microbial decomposition occurs. Temperature also affects soil moisture, which affects decomposition. Freeze-thaw cycles also affect decomposition—freezing temperatures kill soil microorganisms, which allows leaching to play
7612-541: Is infinite, so having debate about what is the total value of nature is actually pointless because we can't live without it'. As of 2012, many companies were not fully aware of the extent of their dependence and impact on ecosystems and the possible ramifications. Likewise, environmental management systems and environmental due diligence tools are more suited to handle "traditional" issues of pollution and natural resource consumption . Most focus on environmental impacts , not dependence. Several tools and methodologies can help
7785-657: Is legally protected from resource development. Much more forest land—about 40 percent of the total forest land base—is subject to varying degrees of protection through processes such as integrated land use planning or defined management areas, such as certified forests. Ecosystem An ecosystem (or ecological system ) is a system that environments and their organisms form through their interaction. The biotic and abiotic components are linked together through nutrient cycles and energy flows. Ecosystems are controlled by external and internal factors . External factors such as climate , parent material which forms
7958-405: Is no clear distinction between biomes and ecosystems. Biomes are always defined at a very general level. Ecosystems can be described at levels that range from very general (in which case the names are sometimes the same as those of biomes) to very specific, such as "wet coastal needle-leafed forests". Biomes vary due to global variations in climate . Biomes are often defined by their structure: at
8131-516: Is no clear distinction between biomes and ecosystems. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of the definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Biotic factors of the ecosystem are living things; such as plants, animals, and bacteria, while abiotic are non-living components; such as water, soil and atmosphere. Plants allow energy to enter
8304-542: Is not on track to meet the target of the United Nations Strategic Plan for Forests to increase forest area by 3 percent by 2030. While deforestation is taking place in some areas, new forests are being established through natural expansion or deliberate efforts in other areas. As a result, the net loss of forest area is less than the rate of deforestation; and it, too, is decreasing: from 7.8 million hectares (19 million acres) per year in
8477-531: Is primarily achieved through bacterial and fungal action. Fungal hyphae produce enzymes that can break through the tough outer structures surrounding dead plant material. They also produce enzymes that break down lignin , which allows them access to both cell contents and the nitrogen in the lignin. Fungi can transfer carbon and nitrogen through their hyphal networks and thus, unlike bacteria, are not dependent solely on locally available resources. Decomposition rates vary among ecosystems. The rate of decomposition
8650-486: Is said to provide substantial ecosystem services to local communities, including benefits to carbon storage, resiliency to climate, and endangered species habitat. As of 2020, the Eglin Air Force Base is said to provide about $ 110 million in ecosystem services per year, $ 40 million more than if no base was present. Although monetary pricing continues with respect to the valuation of ecosystem services,
8823-485: Is termed its resistance . The capacity of a system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Resilience thinking also includes humanity as an integral part of the biosphere where we are dependent on ecosystem services for our survival and must build and maintain their natural capacities to withstand shocks and disturbances. Time plays
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#17327810614168996-423: Is that forests can turn from a carbon sink to a carbon source if plant diversity, density or forest area decreases, as has been observed in different tropical forests The typical tropical forest may become a carbon source by the 2060s. An assessment of European forests found early signs of carbon sink saturation, after decades of increasing strength. The Intergovernmental Panel on Climate Change (IPCC) concluded that
9169-422: Is the "best" classification. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of the definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Different approaches to ecological classifications have been developed in terrestrial, freshwater and marine disciplines, and
9342-722: Is usually assumed that humans benefit from a combination of these services. The services offered by diverse types of ecosystems (forests, seas, coral reefs, mangroves, etc.) differ in nature and in consequence. In fact, some services directly affect the livelihood of neighboring human populations (such as fresh water, food or aesthetic value, etc.) while other services affect general environmental conditions by which humans are indirectly impacted (such as climate change , erosion regulation or natural hazard regulation, etc.). The Millennium Ecosystem Assessment report 2005 defined ecosystem services as benefits people obtain from ecosystems and distinguishes four categories of ecosystem services, where
9515-416: Is whether the forests are composed predominantly of broadleaf trees, coniferous (needle-leaved) trees, or mixed. The number of trees in the world, according to a 2015 estimate, is 3 trillion, of which 1.4 trillion are in the tropics or sub-tropics, 0.6 trillion in the temperate zones, and 0.7 trillion in the coniferous boreal forests. The 2015 estimate is about eight times higher than previous estimates, and
9688-553: The Equator , and temperate forests at the middle latitudes . Forests form in areas of the Earth with high rainfall, while drier conditions produce a transition to savanna . However, in areas with intermediate rainfall levels, forest transitions to savanna rapidly when the percentage of land that is covered by trees drops below 40 to 45 percent. Research conducted in the Amazon rainforest shows that trees can alter rainfall rates across
9861-486: The European Union , just 60% of the yearly forest growth is harvested. Forests also provide non-wood forest products, including fodder, aromatic and medicinal plants, and wild foods. Worldwide, around 1 billion people depend to some extent on wild foods such as wild meat, edible insects, edible plant products, mushrooms and fish, which often contain high levels of key micronutrients. The value of forest foods as
10034-651: The Medieval Latin foresta , denoting "open wood", Carolingian scribes first used foresta in the capitularies of Charlemagne , specifically to denote the royal hunting grounds of the king. The word was not endemic to the Romance languages , e.g., native words for forest in the Romance languages derived from the Latin silva , which denoted "forest" and " wood(land) " ( cf. the English sylva and sylvan ;
10207-590: The Proto-Germanic * furhísa- , * furhíþija- , denoting "a fir-wood , coniferous forest", from the Proto-Indo-European * perku- , denoting "a coniferous or mountain forest , wooded height" all attest to the Frankish * forhist . Uses of forest in English to denote any uninhabited and unenclosed area are presently considered archaic. The Norman rulers of England introduced the word as
10380-553: The Special Report on Global Warming of 1.5 °C of the Intergovernmental Panel on Climate Change , to avoid temperature rise by more than 1.5 degrees above pre-industrial levels, there will need to be an increase in global forest cover equal to the land area of Canada (10 million square kilometres (3.9 million square miles)) by 2050. China instituted a ban on logging, beginning in 1998, due to
10553-591: The World Resources Institute recorded that only 20% of the world's original forests remained in large intact tracts of undisturbed forest. More than 75% of these intact forests lie in three countries: the boreal forests of Russia and Canada, and the rainforest of Brazil. According to Food and Agriculture Organization 's (FAO) Global Forest Resources Assessment 2020 , an estimated 420 million hectares (1.0 billion acres) of forest have been lost worldwide through deforestation since 1990, but
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#173278106141610726-418: The boreal region and in the seasonally dry tropics. At high latitudes, north of the main zone of boreal forestland, growing conditions are not adequate to maintain a continuously closed forest cover , so tree cover is both sparse and discontinuous. This vegetation is variously called open taiga , open lichen woodland, and forest tundra. A savanna is a mixed woodland – grassland ecosystem characterized by
10899-407: The equator are mostly covered in tropical rainforest , and the latitudes between 53°N and 67°N have boreal forest . As a general rule, forests dominated by angiosperms ( broadleaf forests ) are more species-rich than those dominated by gymnosperms ( conifer , montane , or needleleaf forests ), although exceptions exist. The trees that form the principal structural and defining component of
11072-760: The gross primary productivity of the Earth's biosphere , and contain 80% of the Earth's plant biomass. Biomass per unit area is high compared to other vegetation communities. Much of this biomass occurs below ground in the root systems and as partially decomposed plant detritus . The woody component of a forest contains lignin , which is relatively slow to decompose compared with other organic materials such as cellulose or carbohydrate. The world's forests contain about 606 gigatonnes of living biomass (above- and below-ground) and 59 gigatonnes of dead wood. The total biomass has decreased slightly since 1990, but biomass per unit area has increased. Forest ecosystems broadly differ based on climate ; latitudes 10° north and south of
11245-555: The resource inputs are generally controlled by external processes like climate and parent material, the availability of these resources within the ecosystem is controlled by internal factors like decomposition, root competition or shading. Other factors like disturbance, succession or the types of species present are also internal factors. Primary production is the production of organic matter from inorganic carbon sources. This mainly occurs through photosynthesis . The energy incorporated through this process supports life on earth, while
11418-408: The slash and burn practices of swidden agriculture or shifting cultivation . The loss and re-growth of forests lead to a distinction between two broad types of forest: primary or old-growth forest and secondary forest . There are also many natural factors that can cause changes in forests over time, including forest fires , insects , diseases , weather, competition between species, etc. In 1997,
11591-435: The "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . They also include less tangible items like tourism and recreation, and genes from wild plants and animals that can be used to improve domestic species. Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like
11764-1112: The 1990s is the marketing of ecosystem services protection. Payment and trading of services is an emerging worldwide small-scale solution where one can acquire credits for activities such as sponsoring the protection of carbon sequestration sources or the restoration of ecosystem service providers. In some cases, banks for handling such credits have been established and conservation companies have even gone public on stock exchanges, defining an evermore parallel link with economic endeavors and opportunities for tying into social perceptions. However, crucial for implementation are clearly defined land rights , which are often lacking in many developing countries . In particular, many forest-rich developing countries suffering deforestation experience conflict between different forest stakeholders. In addition, concerns for such global transactions include inconsistent compensation for services or resources sacrificed elsewhere and misconceived warrants for irresponsible use. As of 2001, another approach focused on protecting ecosystem service biodiversity hotspots . Recognition that
11937-427: The 1990s to 4.7 million hectares (12 million acres) per year during 2010–2020. In absolute terms, the global forest area decreased by 178 million hectares (440 million acres; 1,780,000 square kilometres; 690,000 square miles) between 1990 and 2020, which is an area about the size of Libya. Forests provide a diversity of ecosystem services including: The main ecosystem services can be summarized in
12110-520: The 1990s. In 2015, a study for Nature Climate Change showed that the trend has recently been reversed, leading to an "overall gain" in global biomass and forests. This gain is due especially to reforestation in China and Russia. New forests are not equivalent to old growth forests in terms of species diversity, resilience, and carbon capture. On 7 September 2015, the FAO released a new study stating that over
12283-617: The 1999 cyclone that hit India. Villages that were surrounded with mangrove forests encountered less damages than other villages that were not protected by mangroves. Supporting services are the services that allow for the other ecosystem services to be present. They have indirect impacts on humans that last over a long period of time. Several services can be considered as being both supporting services and regulating/cultural/provisioning services. Supporting services include for example nutrient cycling , primary production , soil formation , habitat provision. These services make it possible for
12456-457: The Earth's plant biomass . Net primary production is estimated at 21.9 gigatonnes of biomass per year for tropical forests , 8.1 for temperate forests , and 2.6 for boreal forests . Forests form distinctly different biomes at different latitudes and elevations, and with different precipitation and evapotranspiration rates. These biomes include boreal forests in subarctic climates, tropical moist forests and tropical dry forests around
12629-725: The Italian, Spanish, and Portuguese selva ; the Romanian silvă ; the Old French selve ). Cognates of forest in Romance languages—e.g., the Italian foresta , Spanish and Portuguese floresta , etc.—are all ultimately derivations of the French word. The precise origin of Medieval Latin foresta is obscure. Some authorities claim the word derives from the Late Latin phrase forestam silvam , denoting "the outer wood"; others claim
12802-584: The Mediterranean. It was not until the late 1940s that three key authors— Henry Fairfield Osborn, Jr , William Vogt , and Aldo Leopold —promoted recognition of human dependence on the environment. In 1956, Paul Sears drew attention to the critical role of the ecosystem in processing wastes and recycling nutrients. In 1970, Paul Ehrlich and Rosa Weigert called attention to "ecological systems" in their environmental science textbook and "the most subtle and dangerous threat to man's existence ...
12975-469: The Middle Rio Grande basin of New Mexico. This study focused on modeling the stakeholder inputs across a spatial decision, but ignored uncertainty. Another study used Monte Carlo methods to exercise econometric models of landowner decisions in a study of the effects of land-use change . Here the stakeholder inputs were modeled as random effects to reflect the uncertainty. A third study used
13148-493: The Millennium Ecosystem Assessment lumped all of these together as ecosystem services . Four different types of ecosystem services have been distinguished by the scientific body: regulating services, provisioning services, cultural services and supporting services. An ecosystem does not necessarily offer all four types of services simultaneously; but given the intricate nature of any ecosystem, it
13321-451: The U.S. state of Wisconsin , forests managed by indigenous people have more plant diversity, fewer invasive species, higher tree regeneration rates, and higher volume of trees. Forest management has changed considerably over the last few centuries, with rapid changes from the 1980s onward, culminating in a practice now referred to as sustainable forest management . Forest ecologists concentrate on forest patterns and processes, usually with
13494-481: The absence of disturbance, net ecosystem production is equivalent to the net carbon accumulation in the ecosystem. Energy can also be released from an ecosystem through disturbances such as wildfire or transferred to other ecosystems (e.g., from a forest to a stream to a lake) by erosion . In aquatic systems , the proportion of plant biomass that gets consumed by herbivores is much higher than in terrestrial systems. In trophic systems, photosynthetic organisms are
13667-432: The adverse effects of climate change ". Ecosystem services decisions require making complex choices at the intersection of ecology , technology , society , and the economy . The process of making ecosystem services decisions must consider the interaction of many types of information, honor all stakeholder viewpoints, including regulatory agencies , proposal proponents, decision makers, residents, NGOs , and measure
13840-460: The aim of elucidating cause-and-effect relationships. Foresters who practice sustainable forest management focus on the integration of ecological, social, and economic values, often in consultation with local communities and other stakeholders . Humans have generally decreased the amount of forest worldwide. Anthropogenic factors that can affect forests include logging, urban sprawl , human-caused forest fires , acid rain , invasive species , and
14013-402: The amount of energy available to the ecosystem. Parent material determines the nature of the soil in an ecosystem, and influences the supply of mineral nutrients. Topography also controls ecosystem processes by affecting things like microclimate , soil development and the movement of water through a system. For example, ecosystems can be quite different if situated in a small depression on
14186-504: The amount of light available, the amount of leaf area a plant has to capture light (shading by other plants is a major limitation of photosynthesis), the rate at which carbon dioxide can be supplied to the chloroplasts to support photosynthesis, the availability of water, and the availability of suitable temperatures for carrying out photosynthesis. Energy and carbon enter ecosystems through photosynthesis, are incorporated into living tissue, transferred to other organisms that feed on
14359-481: The basis of all food webs. Further, it generates oxygen (O2), a molecule necessary to sustain animals and humans. On average, a human consumes about 550 liter of oxygen per day, whereas plants produce 1,5 liter of oxygen per 10 grams of growth. Cultural services relate to the non-material world, as they benefit the benefit recreational, aesthetic, cognitive and spiritual activities, which are not easily quantifiable in monetary terms. They include: As of 2012, there
14532-419: The basis of the variation of physiognomy corresponding to changes in altitude. Tropical dry forests are characteristic of areas in the tropics affected by seasonal drought. The seasonality of rainfall is usually reflected in the deciduousness of the forest canopy, with most trees being leafless for several months of the year. Under some conditions, such as less fertile soils or less predictable drought regimes,
14705-411: The best grassland management solution for concrete grassland. It will look holistically at the processes in the countryside and help to find best grassland management solutions by taking into account both natural and socioeconomic factors of the particular site. While the notion of human dependence on Earth's ecosystems reaches to the start of Homo sapiens ' existence, the term 'natural capital'
14878-453: The biodiversity of the world ecosystems, reducing both their resilience and biocapacity . The report refers to natural systems as humanity's "life-support system", providing essential ecosystem services. The assessment measures 24 ecosystem services and concludes that only four have shown improvement over the last 50 years, 15 are in serious decline, and five are in a precarious condition. Ecosystem services Ecosystem services are
15051-589: The canopy; but other taxa are also important. In the Southern Hemisphere , most coniferous trees (members of Araucariaceae and Podocarpaceae ) occur mixed with broadleaf species, and are classed as broadleaf-and-mixed forests. Temperate broadleaf and mixed forests include a substantial component of trees of the Anthophyta group. They are generally characteristic of the warmer temperate latitudes, but extend to cool temperate ones, particularly in
15224-408: The carbon makes up much of the organic matter in living and dead biomass, soil carbon and fossil fuels . It also drives the carbon cycle , which influences global climate via the greenhouse effect . Through the process of photosynthesis, plants capture energy from light and use it to combine carbon dioxide and water to produce carbohydrates and oxygen . The photosynthesis carried out by all
15397-688: The challenges in policy implementation and management are significant and considerable. The administration of common pool resources has been a subject of extensive academic pursuit. From defining the problems to finding solutions that can be applied in practical and sustainable ways, there is much to overcome. Considering options must balance present and future human needs, and decision-makers must frequently work from valid but incomplete information. Existing legal policies are often considered insufficient since they typically pertain to human health-based standards that are mismatched with necessary means to protect ecosystem health and services. In 2000, to improve
15570-423: The children live. Canada has about 4 million square kilometres (1.5 million square miles) of forest land. More than 90% of forest land is publicly owned and about 50% of the total forest area is allocated for harvesting. These allocated areas are managed using the principles of sustainable forest management, which include extensive consultation with local stakeholders. About eight percent of Canada's forest
15743-432: The combustion of fossil fuels, ammonia gas which evaporates from agricultural fields which have had fertilizers applied to them, and dust. Anthropogenic nitrogen inputs account for about 80% of all nitrogen fluxes in ecosystems. When plant tissues are shed or are eaten, the nitrogen in those tissues becomes available to animals and microbes. Microbial decomposition releases nitrogen compounds from dead organic matter in
15916-477: The concept of cultural ecosystem services that builds on three arguments: The Common International Classification of Ecosystem Services (CICES) is a classification scheme developed to accounting systems (like National counts etc.), in order to avoid double-counting of Supporting Services with others Provisioning and Regulating Services. Sea sports are very popular among coastal populations: surfing, snorkeling, whale watching, kayaking, recreational fishing ...
16089-418: The concept to draw attention to the importance of transfers of materials between organisms and their environment. He later refined the term, describing it as "The whole system, ... including not only the organism-complex, but also the whole complex of physical factors forming what we call the environment". Tansley regarded ecosystems not simply as natural units, but as "mental isolates". Tansley later defined
16262-481: The conservation of many ecosystem services aligns with more traditional conservation goals (i.e. biodiversity ) has led to the suggested merging of objectives for maximizing their mutual success. This may be particularly strategic when employing networks that permit the flow of services across landscapes , and might also facilitate securing the financial means to protect services through a diversification of investors. For example, as of 2013, there had been interest in
16435-854: The control of climate and disease. Supporting services , such as nutrient cycles and oxygen production. And finally there are cultural services , such as spiritual and recreational benefits. Evaluations of ecosystem services may include assigning an economic value to them. For example, estuarine and coastal ecosystems are marine ecosystems that perform the four categories of ecosystem services in several ways. Firstly, their provisioning services include marine resources and genetic resources . Secondly, their supporting services include nutrient cycling and primary production . Thirdly, their regulating services include carbon sequestration (which helps with climate change mitigation ) and flood control. Lastly, their cultural services include recreation and tourism . The Millennium Ecosystem Assessment (MA) in
16608-515: The cutoff points are between a forest, woodland , and savanna . Under some definitions, to be considered a forest requires very high levels of tree canopy cover, from 60% to 100%, which excludes woodlands and savannas, which have a lower canopy cover . Other definitions consider savannas to be a type of forest, and include all areas with tree canopies over 10%. Some areas covered with trees are legally defined as agricultural areas, for example Norway spruce plantations, under Austrian forest law, when
16781-403: The dead organic matter would accumulate in an ecosystem, and nutrients and atmospheric carbon dioxide would be depleted. Decomposition processes can be separated into three categories— leaching , fragmentation and chemical alteration of dead material. As water moves through dead organic matter, it dissolves and carries with it the water-soluble components. These are then taken up by organisms in
16954-540: The denotation that the use of forest intends. The first known forests on Earth arose in the Middle Devonian (approximately 390 million years ago ), with the evolution of cladoxylopsid plants like Calamophyton . Appeared in the Late Devonian , Archaeopteris was both tree-like and fern -like plant, growing to 20 metres (66 ft) in height or more. It quickly spread throughout the world, from
17127-416: The density of trees, area of tree canopy cover, or area of the land occupied by the cross-section of tree trunks ( basal area ) meeting a particular threshold. This type of definition depends upon the presence of trees sufficient to meet the threshold, or at least of immature trees that are expected to meet the threshold once they mature. Under land-cover definitions, there is considerable variation on where
17300-419: The dynamics of ecological processes relative to ecosystem services is essential in aiding economic decisions. Weighting factors such as a service's irreplaceability or bundled services can also allocate economic value such that goal attainment becomes more efficient. The economic valuation of ecosystem services also involves social communication and information, areas that remain particularly challenging and are
17473-511: The early 2000s has made this concept better known. Ecosystem services or eco-services are defined as the goods and services provided by ecosystems to humans. Per the 2006 Millennium Ecosystem Assessment (MA), ecosystem services are "the benefits people obtain from ecosystems". The MA also delineated the four categories of ecosystem services into provisioning, regulating, supporting, and cultural. By 2010, there had evolved various working definitions and descriptions of ecosystem services in
17646-409: The eastern United States , in which only 0.1% of land was left undisturbed. Almost half of Earth's forest area (49 percent) is relatively intact, while 9 percent is found in fragments with little or no connectivity. Tropical rainforests and boreal coniferous forests are the least fragmented, whereas subtropical dry forests and temperate oceanic forests are among the most fragmented. Roughly 80 percent of
17819-655: The ecosystem or to gradual disruption of biotic processes and degradation of abiotic conditions of the ecosystem. Once the original ecosystem has lost its defining features, it is considered "collapsed ". Ecosystem restoration can contribute to achieving the Sustainable Development Goals . An ecosystem (or ecological system) consists of all the organisms and the abiotic pools (or physical environment) with which they interact. The biotic and abiotic components are linked together through nutrient cycles and energy flows. "Ecosystem processes" are
17992-425: The ecosystems to continue providing services such as food supply, flood regulation, and water purification. Nutrient cycling is the movement of nutrients through an ecosystem by biotic and abiotic processes. The ocean is a vast storage pool for these nutrients, such as carbon, nitrogen and phosphorus. The nutrients are absorbed by the basic organisms of the marine food web and are thus transferred from one organism to
18165-400: The environmental and economic values of ecosystem services. Some people may be unaware of the environment in general and humanity's interrelatedness with the natural environment, which may cause misconceptions. Although environmental awareness is rapidly improving in our contemporary world, ecosystem capital and its flow are still poorly understood, threats continue to impose, and we suffer from
18338-450: The equator to subpolar latitudes. It is the first species known to cast shade due to its fronds and forming soil from its roots. Archaeopteris was deciduous , dropping its fronds onto the forest floor, the shade, soil, and forest duff from the dropped fronds creating the early forest. The shed organic matter altered the freshwater environment, slowing its flow and providing food. This promoted freshwater fish. Forests account for 75% of
18511-584: The erosion and flooding that it caused. In addition, ambitious tree-planting programmes in countries such as China, India, the United States, and Vietnam – combined with natural expansion of forests in some regions – have added more than 7 million hectares (17 million acres) of new forests annually. As a result, the net loss of forest area was reduced to 5.2 million hectares (13 million acres) per year between 2000 and 2010, down from 8.3 million hectares (21 million acres) annually in
18684-421: The flow of energy through a lake was the primary driver of the ecosystem. Hutchinson's students, brothers Howard T. Odum and Eugene P. Odum , further developed a "systems approach" to the study of ecosystems. This allowed them to study the flow of energy and material through ecological systems. Ecosystems are controlled by both external and internal factors. External factors, also called state factors, control
18857-602: The focus of many researchers. In general, the idea is that although individuals make decisions for any variety of reasons, trends reveal the aggregated preferences of a society, from which the economic value of services can be inferred and assigned. The six major methods for valuing ecosystem services in monetary terms are: A peer-reviewed study published in 1997 estimated the value of the world's ecosystem services and natural capital to be between US$ 16 and $ 54 trillion per year, with an average of US$ 33 trillion per year. However, Salles (2011) indicated 'The total value of biodiversity
19030-414: The forest ecosystem. Since 2002, the amount of land that is legally owned by or designated for indigenous peoples has broadly increased, but land acquisition in lower-income countries by multinational corporations, often with little or no consultation of indigenous peoples, has also increased. Research in the Amazon rainforest suggests that indigenous methods of agroforestry form reservoirs of biodiversity. In
19203-458: The forests of eastern North America still show legacies of cultivation which ceased in 1850 when large areas were reverted to forests. Another example is the methane production in eastern Siberian lakes that is controlled by organic matter which accumulated during the Pleistocene . Ecosystems continually exchange energy and carbon with the wider environment . Mineral nutrients, on
19376-696: The four categories of ecosystem services in a variety of ways: The provisioning services include forest products, marine products, fresh water , raw materials, biochemical and genetic resources. Regulating services include carbon sequestration (contributing to climate change mitigation ) as well as waste treatment and disease regulation and buffer zones. Supporting services of coastal ecosystems include nutrient cycling , biologically mediated habitats and primary production . Cultural services of coastal ecosystems include inspirational aspects, recreation and tourism , science and education. Coasts and their adjacent areas on and offshore are an important part of
19549-502: The health and extent of land cover classes that provide ecosystem services, which aids in planning, management, monitoring of stakeholders' actions, and communication between stakeholders. In Baltic countries scientists, nature conservationists and local authorities are implementing integrated planning approach for grassland ecosystems. They are developing an integrated planning tool based on GIS (geographic information system) technology and put online that will help for planners to choose
19722-542: The higher latitudes of the Northern Hemisphere , as well as some warm temperate areas, especially on nutrient-poor or otherwise unfavourable soils. These forests are composed entirely, or nearly so, of coniferous species ( Coniferophyta ). In the Northern Hemisphere, pines Pinus , spruces Picea , larches Larix , firs Abies , Douglas firs Pseudotsuga , and hemlocks Tsuga make up
19895-411: The impacts on all four parts of the intersection. These decisions are usually spatial , always multi-objective , and based on uncertain data, models, and estimates. Often it is the combination of the best science combined with the stakeholder values, estimates and opinions that drive the process. One analytical study modeled the stakeholders as agents to support water resource management decisions in
20068-543: The information available, the implementation of an Ecosystem Services Framework has been suggested (ESF ), which integrates the biophysical and socio-economic dimensions of protecting the environment and is designed to guide institutions through multidisciplinary information and jargon, helping to direct strategic choices. As of 2005 Local to regional collective management efforts were considered appropriate for services like crop pollination or resources like water. Another approach that has become increasingly popular during
20241-407: The land". Wetlands (which include saltwater swamps , salt marshes , ...) and the vegetation it supports – trees, root mats, etc. – retain large amounts of water (surface water, snowmelt, rain, groundwater) and then slowly releases them back, decreasing the likeliness of floods. Mangrove forests protect coastal shorelines from tidal erosion or erosion by currents; a process that was studied after
20414-464: The lands on which they lived as part of global colonialism . Indigenous lands contain 36% or more of intact forest worldwide, host more biodiversity, and experience less deforestation. Indigenous activists have argued that degradation of forests and indigenous peoples' marginalization and land dispossession are interconnected. Other concerns among indigenous peoples include lack of Indigenous involvement in forest management and loss of knowledge related for
20587-691: The landscape, versus one present on an adjacent steep hillside. Other external factors that play an important role in ecosystem functioning include time and potential biota , the organisms that are present in a region and could potentially occupy a particular site. Ecosystems in similar environments that are located in different parts of the world can end up doing things very differently simply because they have different pools of species present. The introduction of non-native species can cause substantial shifts in ecosystem function. Unlike external factors, internal factors in ecosystems not only control ecosystem processes but are also controlled by them. While
20760-491: The last 25 years the global deforestation rate has decreased by 50% due to improved management of forests and greater government protection. There is an estimated 726 million hectares (1.79 billion acres) of forest in protected areas worldwide. Of the six major world regions, South America has the highest share of forests in protected areas, at 31 percent. The area of such areas globally has increased by 191 million hectares (470 million acres) since 1990, but
20933-477: The list of "ecosystem engineers"—organisms that physically, biologically or chemically modify the environment around them in ways that influence the health of other organisms. Many of the ecological functions and processes performed or affected by shellfish contribute to human well-being by providing a stream of valuable ecosystem services over time by filtering out particulate materials and potentially mitigating water quality issues by controlling excess nutrients in
21106-611: The literature. To prevent double-counting in ecosystem services audits, for instance, The Economics of Ecosystems and Biodiversity (TEEB) replaced "Supporting Services" in the MA with "Habitat Services" and "ecosystem functions", defined as "a subset of the interactions between ecosystem structure and processes that underpin the capacity of an ecosystem to provide goods and services". While Gretchen Daily 's original definition distinguished between ecosystem goods and ecosystem services , Robert Costanza and colleagues' later work and that of
21279-423: The living and dead plant matter, and eventually released through respiration. The carbon and energy incorporated into plant tissues (net primary production) is either consumed by animals while the plant is alive, or it remains uneaten when the plant tissue dies and becomes detritus . In terrestrial ecosystems , the vast majority of the net primary production ends up being broken down by decomposers . The remainder
21452-475: The main wood-based product groups ranged from 1 percent (woodbased panels) to 5 percent (industrial roundwood). The fastest growth occurred in the Asia-Pacific, Northern American and European regions, likely due to positive economic growth in these areas. Over 40% of the territory in the European Union is covered by forests. This region has grown via afforestation by roughly 0.4% year in recent decades. In
21625-471: The maintenance of hydrological cycles , cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. Many ecosystems become degraded through human impacts, such as soil loss , air and water pollution , habitat fragmentation , water diversion , fire suppression , and introduced species and invasive species . These threats can lead to abrupt transformation of
21798-408: The maintenance of hydrological cycles, cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. While material from the ecosystem had traditionally been recognized as being the basis for things of economic value, ecosystem services tend to be taken for granted. The Millennium Ecosystem Assessment
21971-988: The montane forests of Africa, South America, Southeast Asia, and lowland forests of Australia, coastal Brazil, the Caribbean islands, Central America, and insular Southeast Asia have many species with small geographical distributions. Areas with dense human populations and intense agricultural land use, such as Europe, parts of Bangladesh, China, India, and North America, are less intact in terms of their biodiversity. Northern Africa, southern Australia, coastal Brazil, Madagascar, and South Africa are also identified as areas with striking losses in biodiversity intactness. A forest consists of many components that can be broadly divided into two categories: biotic (living) and abiotic (non-living). The living parts include trees , shrubs , vines , grasses and other herbaceous (non-woody) plants, mosses , algae , fungi , insects , mammals , birds , reptiles , amphibians , and microorganisms living on
22144-514: The next table: Some researchers state that forests do not only provide benefits, but can in certain cases also incur costs to humans. Forests may impose an economic burden, diminish the enjoyment of natural areas, reduce the food-producing capacity of grazing land and cultivated land, reduce biodiversity, reduce available water for humans and wildlife, harbour dangerous or destructive wildlife, and act as reservoirs of human and livestock disease. An important consideration regarding carbon sequestration
22317-557: The other and from one ecosystem to the other. Nutrients are recycled through the life cycle of organisms as they die and decompose, releasing the nutrients into the neighboring environment. "The service of nutrient cycling eventually impacts all other ecosystem services as all living things require a constant supply of nutrients to survive". Primary production refers to the production of organic matter, i.e., chemically bound energy, through processes such as photosynthesis and chemosynthesis. The organic matter produced by primary producers forms
22490-537: The other hand, are mostly cycled back and forth between plants, animals, microbes and the soil. Most nitrogen enters ecosystems through biological nitrogen fixation , is deposited through precipitation, dust, gases or is applied as fertilizer . Most terrestrial ecosystems are nitrogen-limited in the short term making nitrogen cycling an important control on ecosystem production. Over the long term, phosphorus availability can also be critical. Macronutrients which are required by all plants in large quantities include
22663-509: The other. Despite this, the cumulative effect of additional species in an ecosystem is not linear: additional species may enhance nitrogen retention, for example. However, beyond some level of species richness, additional species may have little additive effect unless they differ substantially from species already present. This is the case for example for exotic species . The addition (or loss) of species that are ecologically similar to those already present in an ecosystem tends to only have
22836-403: The overall structure of an ecosystem and the way things work within it, but are not themselves influenced by the ecosystem. On broad geographic scales, climate is the factor that "most strongly determines ecosystem processes and structure". Climate determines the biome in which the ecosystem is embedded. Rainfall patterns and seasonal temperatures influence photosynthesis and thereby determine
23009-410: The plants and animals and in the soil, connected by mycorrhizal networks . The main layers of all forest types are the forest floor , the understory, and the canopy. The emergent layer, above the canopy, exists in tropical rainforests. Each layer has a different set of plants and animals, depending upon the availability of sunlight, moisture, and food. In botany and countries like Germany and Poland,
23182-405: The plants in an ecosystem is called the gross primary production (GPP). About half of the gross GPP is respired by plants in order to provide the energy that supports their growth and maintenance. The remainder, that portion of GPP that is not used up by respiration, is known as the net primary production (NPP). Total photosynthesis is limited by a range of environmental factors. These include
23355-510: The population belongs to forest-dependent communities, which live in close proximity to forests and practice agroforestry as a principal part of their livelihood. People of Ghana who rely on timber and bushmeat harvested from forests and Indigenous peoples of the Amazon rainforest are also examples of forest-dependent people. Though forest-dependence by more common definitions is statistically associated with poverty and rural livelihoods, elements of forest-dependence exist in communities with
23528-452: The potential destruction, by man's own activities, of those ecological systems upon which the very existence of the human species depends". The term environmental services was introduced in a 1970 report of the Study of Critical Environmental Problems , which listed services including insect pollination, fisheries , climate regulation and flood control. In following years, variations of
23701-585: The primary nutrients (which are most limiting as they are used in largest amounts): Nitrogen, phosphorus, potassium. Secondary major nutrients (less often limiting) include: Calcium, magnesium, sulfur. Micronutrients required by all plants in small quantities include boron, chloride, copper, iron, manganese, molybdenum, zinc. Finally, there are also beneficial nutrients which may be required by certain plants or by plants under specific environmental conditions: aluminum, cobalt, iodine, nickel, selenium, silicon, sodium, vanadium. Until modern times, nitrogen fixation
23874-413: The primary producers. The organisms that consume their tissues are called primary consumers or secondary producers — herbivores . Organisms which feed on microbes ( bacteria and fungi ) are termed microbivores . Animals that feed on primary consumers— carnivores —are secondary consumers. Each of these constitutes a trophic level. The sequence of consumption—from plant to herbivore, to carnivore—forms
24047-665: The private sector value and assess ecosystem services, including Our Ecosystem, the 2008 Corporate Ecosystem Services Review, the Artificial Intelligence for Environment & Sustainability (ARIES) project from 2007, the Natural Value Initiative (2012) and InVEST (Integrated Valuation of Ecosystem Services & Tradeoffs, 2012) To provide an example of a cost comparison: The land of the United States Department of Defense
24220-549: The production of timber, forestry activities may also result in products that undergo little processing, such as fire wood, charcoal, wood chips and roundwood used in an unprocessed form. Global production and trade of all major wood-based products recorded their highest ever values in 2018. Production, imports and exports of roundwood, sawnwood, wood-based panels, wood pulp, wood charcoal and pellets reached their maximum quantities since 1947 when FAO started reporting global forest product statistics. In 2018, growth in production of
24393-614: The proportion of evergreen species increases and the forests are characterised as " sclerophyllous ". Thorn forest , a dense forest of low stature with a high frequency of thorny or spiny species, is found where drought is prolonged, and especially where grazing animals are plentiful. On very poor soils, and especially where fire or herbivory are recurrent phenomena, savannas develop. Sparse trees and savanna are forests with sparse tree- canopy cover. They occur principally in areas of transition from forested to non-forested landscapes. The two major zones in which these ecosystems occur are in
24566-427: The rate of forest loss has declined substantially. In the most recent five-year period (2015–2020), the annual rate of deforestation was estimated at 10 million hectares (25 million acres), down from 12 million hectares (30 million acres) annually in 2010–2015. The transition of a region from forest loss to net gain in forested land is referred to as the forest transition. This change occurs through
24739-464: The rate of annual increase slowed in 2010–2020. Smaller areas of woodland in cities may be managed as urban forestry , sometimes within public parks. These are often created for human benefits; Attention Restoration Theory argues that spending time in nature reduces stress and improves health, while forest schools and kindergartens help young people to develop social as well as scientific skills in forests. These typically need to be close to where
24912-458: The region and habitat. In contrast, secondary forest is forest regrowing following timber harvest and may contain species originally from other regions or habitats. Different global forest classification systems have been proposed, but none has gained universal acceptance. UNEP - WCMC 's forest category classification system is a simplification of other, more complex systems (e.g. UNESCO 's forest and woodland 'subformations'). This system divides
25085-507: The so-called ' tragedy of the commons '. Many efforts to inform decision-makers of current versus future costs and benefits now involve organizing and translating scientific knowledge to economics , which articulate the consequences of our choices in comparable units of impact on human well-being. An especially challenging aspect of this process is that interpreting ecological information collected from one spatial-temporal scale does not necessarily mean it can be applied at another; understanding
25258-462: The so-called supporting services are regarded as the basis for the services of the other three categories. Provisioning services consist of all "the products obtained from ecosystems". The following services are also known as ecosystem goods : Forests and forest management produce a large type and variety of timber products, including roundwood, sawnwood, panels, and engineered wood, e.g., cross-laminated timber, as well as pulp and paper. Besides
25431-409: The soil and topography , control the overall structure of an ecosystem but are not themselves influenced by the ecosystem. Internal factors are controlled, for example, by decomposition , root competition, shading, disturbance, succession, and the types of species present. While the resource inputs are generally controlled by external processes, the availability of these resources within the ecosystem
25604-611: The soil, react with mineral soil, or are transported beyond the confines of the ecosystem (and are considered lost to it). Newly shed leaves and newly dead animals have high concentrations of water-soluble components and include sugars , amino acids and mineral nutrients. Leaching is more important in wet environments and less important in dry ones. Fragmentation processes break organic material into smaller pieces, exposing new surfaces for colonization by microbes. Freshly shed leaf litter may be inaccessible due to an outer layer of cuticle or bark , and cell contents are protected by
25777-511: The soil, where plants, fungi, and bacteria compete for it. Some soil bacteria use organic nitrogen-containing compounds as a source of carbon, and release ammonium ions into the soil. This process is known as nitrogen mineralization . Others convert ammonium to nitrite and nitrate ions, a process known as nitrification . Nitric oxide and nitrous oxide are also produced during nitrification. Under nitrogen-rich and oxygen-poor conditions, nitrates and nitrites are converted to nitrogen gas ,
25950-615: The southern hemisphere. They include such forest types as the mixed deciduous forests of the United States and their counterparts in China and Japan; the broadleaf evergreen rainforests of Japan, Chile , and Tasmania ; the sclerophyllous forests of Australia, central Chile, the Mediterranean , and California; and the southern beech Nothofagus forests of Chile and New Zealand. There are many different types of tropical moist forests , with lowland evergreen broad-leaf tropical rainforests : for example várzea and igapó forests and
26123-483: The spatial extent of ecosystems using the term " ecotope ". G. Evelyn Hutchinson , a limnologist who was a contemporary of Tansley's, combined Charles Elton 's ideas about trophic ecology with those of Russian geochemist Vladimir Vernadsky . As a result, he suggested that mineral nutrient availability in a lake limited algal production . This would, in turn, limit the abundance of animals that feed on algae. Raymond Lindeman took these ideas further to suggest that
26296-454: The system through photosynthesis , building up plant tissue. Animals play an important role in the movement of matter and energy through the system, by feeding on plants and on one another. They also influence the quantity of plant and microbial biomass present. By breaking down dead organic matter , decomposers release carbon back to the atmosphere and facilitate nutrient cycling by converting nutrients stored in dead biomass back to
26469-414: The term were used, but eventually 'ecosystem services' became the standard in scientific literature. The ecosystem services concept has continued to expand and includes socio-economic and conservation objectives. [REDACTED] This article incorporates text from a free content work. Licensed under CC BY-SA 3.0 IGO ( license statement/permission ). Text taken from The State of
26642-690: The terra firme forests of the Amazon Basin ; the peat swamp forests ; dipterocarp forests of Southeast Asia ; and the high forests of the Congo Basin . Seasonal tropical forests , perhaps the best description for the colloquial term " jungle ", typically range from the rainforest zone 10 degrees north or south of the equator , to the Tropic of Cancer and Tropic of Capricorn . Forests located on mountains are also included in this category, divided largely into upper and lower montane formations, on
26815-421: The transfers of energy and materials from one pool to another. Ecosystem processes are known to "take place at a wide range of scales". Therefore, the correct scale of study depends on the question asked. The term "ecosystem" was first used in 1935 in a publication by British ecologist Arthur Tansley . The term was coined by Arthur Roy Clapham , who came up with the word at Tansley's request. Tansley devised
26988-474: The trees are being grown as Christmas trees and are below a certain height. The word forest derives from the Old French forest (also forès ), denoting "forest, vast expanse covered by trees"; forest was first introduced into English as the word denoting wild land set aside for hunting without necessarily having trees on the land. Possibly a borrowing, probably via Frankish or Old High German , of
27161-424: The trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to support an unbroken herbaceous layer that consists primarily of grasses. Savannas maintain an open canopy despite a high tree density. Forest plantations are generally intended for the production of timber and pulpwood . Commonly mono-specific, planted with even spacing between
27334-401: The trees, and intensively managed, these forests are generally important as habitat for native biodiversity . Some are managed in ways that enhance their biodiversity protection functions and can provide ecosystem services such as nutrient capital maintenance, watershed and soil structure protection and carbon storage. The annual net loss of forest area has decreased since 1990, but the world
27507-504: The type of vegetation that grows upon the land; an area can be legally designated "forest" even if no trees grow on it. Land-use definitions are based on the primary purpose the land is used for. Under a land-use definition, any area used primarily for harvesting timber, including areas that have been cleared by harvesting, disease, fire, or for the construction of roads and infrastructure, are still defined as forests, even if they contain no trees. Land-cover definitions define forests based upon
27680-453: The valuation of ecosystem services provided by shellfish production and restoration. A keystone species, low in the food chain, bivalve shellfish such as oysters support a complex community of species by performing a number of functions essential to the diverse array of species that surround them. There is also increasing recognition that some shellfish species may impact or control many ecological processes; so much so that they are included on
27853-426: The various benefits that humans derive from healthy ecosystems . These ecosystems, when functioning well, offer such things as provision of food, natural pollination of crops, clean air and water, decomposition of wastes, or flood control . Ecosystem services are grouped into four broad categories of services. There are provisioning services , such as the production of food and water. Regulating services , such as
28026-653: The water. As of 2018, the concept of ecosystem services had not been properly implemented into international and regional legislation yet. Notwithstanding, the United Nations Sustainable Development Goal 15 has a target to ensure the conservation, restoration, and sustainable use of ecosystem services. An estimated $ 125 trillion to $ 140 trillion is added to the economy each year by all ecosystem services. However, many of these services are at risk due to climate and other anthropogenic impacts. Climate-driven shifts in biome ranges
28199-621: The word forest is commonly used, there is no universally recognised precise definition, with more than 800 definitions of forest used around the world. Although a forest is usually defined by the presence of trees, under many definitions an area completely lacking trees may still be considered a forest if it grew trees in the past, will grow trees in the future, or was legally designated as a forest regardless of vegetation type. There are three broad categories of definitions of forest in use: administrative, land use , and land cover . Administrative definitions are legal designations, and may not reflect
28372-591: The word in all three of its senses: common, legal, and archaic. Other English words used to denote "an area with a high density of trees" are firth , frith , holt , weald , wold , wood , and woodland . Unlike forest , these are all derived from Old English and were not borrowed from another language. Some present classifications reserve woodland for denoting a locale with more open space between trees, and distinguish kinds of woodlands as open forests and closed forests , premised on their crown covers . Finally, sylva (plural sylvae or, less classically, sylvas )
28545-461: The word is a Latinisation of the Frankish * forhist , denoting "forest, wooded country", and was assimilated to forestam silvam , pursuant to the common practice of Frankish scribes. The Old High German forst denoting "forest"; Middle Low German vorst denoting "forest"; Old English fyrhþ denoting "forest, woodland, game preserve, hunting ground" (English frith ); and Old Norse fýri , denoting " coniferous forest "; all of which derive from
28718-576: The world's forest area is found in patches larger than 1 million hectares (2.5 million acres). The remaining 20 percent is located in more than 34 million patches around the world – the vast majority less than 1,000 hectares (2,500 acres) in size. Human society and forests can affect one another positively or negatively. Forests provide ecosystem services to humans and serve as tourist attractions. Forests can also affect people's health. Human activities, including unsustainable use of forest resources, can negatively affect forest ecosystems. Although
28891-415: The world's forests into 26 major types, which reflect climatic zones as well as the principal types of trees. These 26 major types can be reclassified into 6 broader categories: temperate needleleaf, temperate broadleaf and mixed, tropical moist, tropical dry, sparse trees and parkland, and forest plantations. Each category is described in a separate section below. Temperate needleleaf forests mostly occupy
29064-417: The world's land area in 2020. Forests are the largest terrestrial ecosystems of Earth by area, and are found around the globe. 45 percent of forest land is in the tropical latitudes . The next largest share of forests are found in subarctic climates , followed by temperate , and subtropical zones. Forests account for 75% of the gross primary production of the Earth's biosphere , and contain 80% of
29237-407: Was a discussion as to how the concept of cultural ecosystem services could be operationalized, how landscape aesthetics, cultural heritage, outdoor recreation, and spiritual significance to define can fit into the ecosystem services approach. who vote for models that explicitly link ecological structures and functions with cultural values and benefits. Likewise, there has been a fundamental critique of
29410-476: Was estimated at $ 1–1.5 billion, which contrasted dramatically with the estimated $ 6–8 billion cost of constructing a water filtration plant plus the $ 300 million annual running costs. Pollination of crops by bees is required for 15–30% of U.S. food production ; most large-scale farmers import non-native honey bees to provide this service. A 2005 study reported that in California's agricultural region, it
29583-476: Was first coined by E. F. Schumacher in 1973 in his book Small is Beautiful . Recognition of how ecosystems could provide complex services to humankind date back to at least Plato (c. 400 BC) who understood that deforestation could lead to soil erosion and the drying of springs. Modern ideas of ecosystem services probably began when Marsh challenged in 1864 the idea that Earth's natural resources are unbounded by pointing out changes in soil fertility in
29756-480: Was found that wild bees alone could provide partial or complete pollination services or enhance the services provided by honey bees through behavioral interactions. However, intensified agricultural practices can quickly erode pollination services through the loss of species. The remaining species are unable to compensate this. The results of this study also indicate that the proportion of chaparral and oak-woodland habitat available for wild bees within 1–2 km of
29929-679: Was the major source of nitrogen for ecosystems. Nitrogen-fixing bacteria either live symbiotically with plants or live freely in the soil. The energetic cost is high for plants that support nitrogen-fixing symbionts—as much as 25% of gross primary production when measured in controlled conditions. Many members of the legume plant family support nitrogen-fixing symbionts. Some cyanobacteria are also capable of nitrogen fixation. These are phototrophs , which carry out photosynthesis. Like other nitrogen-fixing bacteria, they can either be free-living or have symbiotic relationships with plants. Other sources of nitrogen include acid deposition produced through
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