Approximately 251.9 million years ago, the Permian–Triassic ( P–T , P–Tr ) extinction event ( PTME ; also known as the Late Permian extinction event , the Latest Permian extinction event , the End-Permian extinction event , and colloquially as the Great Dying ) forms the boundary between the Permian and Triassic geologic periods , and with them the Paleozoic and Mesozoic eras. It is Earth 's most severe known extinction event , with the extinction of 57% of biological families , 83% of genera, 81% of marine species and 70% of terrestrial vertebrate species. It is also the greatest known mass extinction of insects . It is the greatest of the "Big Five" mass extinctions of the Phanerozoic . There is evidence for one to three distinct pulses, or phases, of extinction.
192-617: Fergana Valley is a valley in Central Asia , lying mostly in eastern Uzbekistan , extending into southern Kyrgyzstan and northern Tajikistan . Encompassing three former Soviet Republics , the valley is ethnically diverse and relations among the countries are tense. Ethnic enclaves of Uzbeks in Kyrgyz sovereign territory, restricted right of movement, lack of agreement about border demarcation and disputes over access to family members and places of economic activity contribute to tension at
384-458: A lithological sequence as being on or very close to the Permian–Triassic boundary in rocks that are unsuitable for radiometric dating or have a lack of suitable index fossils . However, even the proposers of the fungal spike hypothesis pointed out that "fungal spikes" may have been a repeating phenomenon created by the post-extinction ecosystem during the earliest Triassic. The very idea of
576-727: A stringed instrument —in Kyrgyzstan, a three-stringed komuz , and in Kazakhstan, a similar two-stringed instrument, the dombra . Photography in Central Asia began to develop after 1882, when a Russian Mennonite photographer named Wilhelm Penner moved to the Khanate of Khiva during the Mennonite migration to Central Asia led by Claas Epp, Jr. Upon his arrival to Khanate of Khiva , Penner shared his photography skills with
768-620: A brief period of domination in the early Spathian, probably related to a transient oxygenation of deep waters. Neospathodid conodonts survived the crisis but underwent proteromorphosis. In the PTME's aftermath, disaster taxa of benthic foraminifera filled many of their vacant niches. The recovery of benthic foraminifera was very slow and frequently interrupted until the Spathian. In the Tethys, foraminiferal communities remained low in diversity into
960-632: A ceiling limiting the maximum ecological complexity of marine ecosystems until the Spathian. Recovery biotas appear to have been ecologically uneven and unstable into the Anisian , making them vulnerable to environmental stresses. Whereas most marine communities were fully recovered by the Middle Triassic, global marine diversity reached pre-extinction values no earlier than the Middle Jurassic, approximately 75 million years after
1152-588: A considerable degree with the local Turkic and Turkic-speaking population, so that at the time of Timur's reign the Barlas had become thoroughly Turkicized in terms of language and habits. Additionally, by adopting Islam, the Central Asian Turks and Mongols also adopted the Persian literary and high culture which had dominated Central Asia since the early days of Islamic influence. Persian literature
1344-449: A duration of 60 ± 48 thousand years. A large, abrupt global decrease in δ C , the ratio of the stable isotope carbon-13 to that of carbon-12 , coincides with this extinction, and is sometimes used to identify the Permian–Triassic boundary and PTME in rocks that are unsuitable for radiometric dating . The negative carbon isotope excursion's magnitude was 4-7% and lasted for approximately 500 kyr, though estimating its exact value
1536-513: A family of large-size fusuline foraminifera . The impact of the end-Guadalupian extinction on marine organisms appears to have varied between locations and between taxonomic groups – brachiopods and corals had severe losses. Marine invertebrates suffered the greatest losses during the P–Tr extinction. Evidence of this was found in samples from south China sections at the P–Tr boundary. Here, 286 out of 329 marine invertebrate genera disappear within
1728-420: A fungal origin for Reduviasporonites , diluting these critiques. Uncertainty exists regarding the duration of the overall extinction and about the timing and duration of various groups' extinctions within the greater process. Some evidence suggests that there were multiple extinction pulses or that the extinction was long and spread out over a few million years, with a sharp peak in the last million years of
1920-491: A fungal spike has been criticized on several grounds, including: Reduviasporonites , the most common supposed fungal spore, may be a fossilized alga ; the spike did not appear worldwide; and in many places it did not fall on the Permian–Triassic boundary. The Reduviasporonites may even represent a transition to a lake-dominated Triassic world rather than an earliest Triassic zone of death and decay in some terrestrial fossil beds. Newer chemical evidence agrees better with
2112-465: A gallop, the sculpture symbolizes peak speed and power, with its V-shaped leg arrangement echoing Greek symbols of speed and victory. Detailed anatomical features, such as visible ribs and dynamic muscle contractions, showcase a profound understanding of equine biomechanics. The lifelike depiction extends to aerodynamic adaptations like flat ears, a streamlined mane, and a fanned tail, enhancing balance and stability at high speeds. Decorative swirls symbolize
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#17327654944942304-425: A good many camels are bred. Over 6,900 ha (17,000 acres) were planted with vines , and some 140,000 ha (350,000 acres) were under cotton . Nearly 400,000 ha (1,000,000 acres) were covered with forests. The government maintained a forestry farm at Marghelan , from which 120,000 to 200,000 young trees were distributed free every year amongst the inhabitants of the province. Silkworm breeding, formerly
2496-581: A great reduction in their geographic range. Following this transition, coal swamps vanished. The North Chinese floral extinction correlates with the decline of the Gigantopteris flora of South China. In South China, the subtropical Cathaysian gigantopterid dominated rainforests abruptly collapsed. The floral extinction in South China is associated with bacterial blooms in soil and nearby lacustrine ecosystems, with soil erosion resulting from
2688-424: A group are also colloquially referred to as the "-stans" as all have names ending with the Persian suffix " -stan " (meaning 'land') in both respective native languages and most other languages. In the pre-Islamic and early Islamic eras ( c. 1000 and earlier) Central Asia was inhabited predominantly by Iranian peoples , populated by Eastern Iranian -speaking Bactrians , Sogdians , Chorasmians , and
2880-513: A group of prominent local businessmen brought underlying tensions to a head in the region around Andijan and Qorasuv during the May 2005 unrest in Uzbekistan in which hundreds of protestors were killed by troops. There was violence again in 2010 in the Kyrgyz part of the valley, heated by ethnic tensions, worsening economic conditions due to the global economic crisis, and political conflict over
3072-420: A high background extinction rate (by implication, taxa with a high turnover ). The extinction rate of marine organisms was catastrophic. Bioturbators were extremely severely affected, as evidenced by the loss of the sedimentary mixed layer in many marine facies during the end-Permian extinction. Surviving marine invertebrate groups included articulate brachiopods (those with a hinge), which had undergone
3264-487: A high-resolution age model for the extinction – allowing exploration of the links between global environmental perturbation, carbon cycle disruption, mass extinction, and recovery at millennial timescales. The first appearance of the conodont Hindeodus parvus has been used to delineate the Permian-Triassic boundary. The extinction occurred between 251.941 ± 0.037 and 251.880 ± 0.031 million years ago,
3456-433: A large body of water, temperature fluctuations are often severe, excluding the hot, sunny summer months. In most areas, the climate is dry and continental, with hot summers and cool to cold winters, with occasional snowfall. Outside high-elevation areas, the climate is mostly semi-arid to arid. In lower elevations, summers are hot with blazing sunshine. Winters feature occasional rain or snow from low-pressure systems that cross
3648-567: A local student Khudaybergen Divanov, who later became the founder of Uzbek photography . Some also learn to sing the Manas , Kyrgyzstan's epic poem (those who learn the Manas exclusively but do not improvise are called manaschis ). During Soviet rule, akyn performance was co-opted by the authorities and subsequently declined in popularity. With the fall of the Soviet Union , it has enjoyed
3840-566: A massive rearrangement of ecosystems does occur, with plant abundances and distributions changing profoundly and all the forests virtually disappearing. The dominant floral groups changed, with many groups of land plants entering abrupt decline, such as Cordaites ( gymnosperms ) and Glossopteris ( seed ferns ). The severity of plant extinction has been disputed. The Glossopteris -dominated flora that characterised high-latitude Gondwana collapsed in Australia around 370,000 years before
4032-544: A more heterogeneous region with increasing East Asian ancestry through Turkic and Mongolian groups in the past thousands years, including extensive Turkic and later Mongol migrations out of Mongolia and slow assimilation of local populations. In the 8th century AD, the Islamic expansion reached the region but had no significant demographic impact. In the 13th century AD, the Mongolian invasion of Central Asia brought most of
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#17327654944944224-819: A path between Greek, Chinese, Bactrian and Parthian civilisations. It was home to Babur , founder of the Mughal dynasty , tying the region to modern Afghanistan and South Asia. The Russian Empire conquered the valley at the end of the 19th century, and it became part of the Soviet Union in the 1920s. Its three Soviet republics gained independence in 1991. The area largely remains Muslim , populated by ethnic Uzbek , Tajik and Kyrgyz people , often intermixed and not matching modern borders. Historically there have also been substantial numbers of Russian , Kashgarian , Kipchak , Bukharan Jewish and Romani minorities. The economy centers around extensive cotton cultivation,
4416-471: A practice initiated by the Soviets, complemented by a diverse array of grains, fruits, and vegetables. Additionally, the region has a rich heritage in stock breeding, leatherwork, and a progressively expanding mining industry, including deposits of coal , iron , sulfur , gypsum , rock-salt , naphtha and some small known oil reserves. The word Fergana ( / f ɜːr ˈ ɡ ɑː n ə / fur-GAHN-ə )
4608-624: A prosperous industry, had decayed, despite the encouragement of a state farm at New Marghelan. Coal, iron, sulfur, gypsum , rock-salt , and naphtha are all known to exist, but only the last two have ever been extracted in significant quantities. In the late 19th century there were a few small oil-wells in Fergana, but these no longer function. In the Tsarist period the only industrial enterprises were some seventy or eighty factories engaged in cotton cleaning. Leather, saddlery, paper and cutlery were
4800-523: A resurgence, although akyns still do use their art to campaign for political candidates. A 2005 The Washington Post article proposed a similarity between the improvisational art of akyns and modern freestyle rap performed in the West. As a consequence of Russian colonisation, European fine arts – painting, sculpture and graphics – have developed in Central Asia. The first years of the Soviet regime saw
4992-653: A revolt which was far more widespread than that of 1898, and which was not entirely suppressed by the time of the Russian Revolution . In 1924, the new boundaries separating the Uzbek SSR and Kyrgyz SSR cut off the eastern end of the Fergana Valley, as well as the slopes surrounding it. This was compounded in 1928 when the Tajik ASSR became a fully-fledged republic, and the area around Khujand
5184-462: A scarcity of research on climate impacts in Central Asia, even though it experiences faster warming than the global average and is generally considered to be one of the more climate-vulnerable regions in the world. Along with West Asia , it has already had greater increases in hot temperature extremes than the other parts of Asia, Rainfall in Central Asia had decreased, unlike elsewhere in Asia, and
5376-631: A separate geographic zone. The valley owes its fertility to two rivers, the Naryn and the Kara Darya , which unite in the valley, near Namangan , to form the Syr Darya . Numerous other tributaries of these rivers exist in the valley including the Sokh River . The streams, and their numerous mountain effluents, not only supply water for irrigation, but also bring down vast quantities of sand, which
5568-553: A severe bottleneck in diversity. Evidence from South China indicates the foraminiferal extinction had two pulses. Foraminiferal biodiversity hotspots shifted into deeper waters during the PTME. Approximately 93% of latest Permian foraminifera became extinct, with 50% of the clade Textulariina, 92% of Lagenida, 96% of Fusulinida, and 100% of Miliolida disappearing. Foraminifera that were calcaerous suffered an extinction rate of 91%. The reason why lagenides survived while fusulinoidean fusulinides went completely extinct may have been due to
5760-481: A slow decline in numbers since the P–Tr extinction; the Ceratitida order of ammonites ; and crinoids ("sea lilies"), which very nearly became extinct but later became abundant and diverse. The groups with the highest survival rates generally had active control of circulation , elaborate gas exchange mechanisms, and light calcification; more heavily calcified organisms with simpler breathing apparatuses suffered
5952-406: A specific region were more likely to go extinct than cosmopolitan taxa. There was little latitudinal difference in the survival rates of taxa. Organisms that inhabited refugia less affected by global warming experienced lesser or delayed extinctions. Among benthic organisms the extinction event multiplied background extinction rates , and therefore caused maximum species loss to taxa that had
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6144-544: A wingspan over 20 cm. Unusual small reptiles, Longisquama with long scales, and Sharovipteryx , which glided using a membrane stretched between their paws, were also found here. The climate of this valley is dry and continental, being mostly a cool arid climate ( Köppen BWk ) or a cool semi-arid climate ( BSk ) in less shielded areas. In March the temperature reaches 20 °C (68 °F), and then rapidly rises to 35 °C (95 °F) in June, July and August. During
6336-523: Is Babur , heir to Timur and famous conqueror and founder of the Mughal dynasty in Medieval India . Islamic proselytizers from the Fergana Valley such as al-Firghani الفرغاني, al-Andijani الأندجاني, al-Namangani النمنگاني, al-Khojandi الخوجندي spread Islam into parts of present-day Russia, China, and India. The Fergana valley was ruled by a series of Muslim states in the medieval period. For much of this period local and southwestern rulers divided
6528-730: Is between the Oxus and Jaxartes, and the Karakum Desert is between the Oxus and Kopet Dagh in Turkmenistan. Khorasan meant approximately northeast Persia and northern Afghanistan. Margiana was the region around Merv. The Ustyurt Plateau is between the Aral and Caspian Seas. To the southwest, across the Kopet Dagh, lies Persia. From here Persian and Islamic civilisation penetrated Central Asia and dominated its high culture until
6720-521: Is challenging due to diagenetic alteration of many sedimentary facies spanning the boundary. Further evidence for environmental change around the Permian-Triassic boundary suggests an 8 °C (14 °F) rise in temperature, and an increase in CO 2 levels to 2,500 ppm (for comparison, the concentration immediately before the Industrial Revolution was 280 ppm , and
6912-519: Is characterized by block subsidence , originally to depths estimated at 6 to 7 kilometres (3.7 to 4.3 mi), largely filled with sediments that range in age as far back as the Permian-Triassic boundary . Some of the sediments are marine carbonates and clays . The faults are upthrusts and overthrusts. Anticlines associated with these faults form traps for petroleum and natural gas , which has been discovered in 52 small fields . In
7104-612: Is cotton, although the Daewoo plant in Andizhan sends cars all over Uzbekistan. Central Asia Central Asia is a region of Asia bounded by the Caspian Sea to the southwest, European Russia to the northwest, Western China and Mongolia to the east, Afghanistan and Iran to the south, and Siberia to the north. It includes Kazakhstan , Kyrgyzstan , Tajikistan , Turkmenistan , and Uzbekistan . The countries as
7296-510: Is deposited alongside their courses, more especially alongside the Syr Darya where it cuts its way through the Khujand - Ajar ridge and forms the valley. This expanse of quicksand , covering an area of 1,900 km (750 sq mi), under the influence of south-west winds, encroaches upon the agricultural districts. The central part of the geological depression that forms the valley
7488-468: Is difficult to know whether the end-Capitanian had finished, depending on the factor considered. Many of the extinctions once dated to the Permian-Triassic boundary have more recently been redated to the end- Capitanian . Further, it is unclear whether some species who survived the prior extinction(s) had recovered well enough for their final demise in the Permian-Triassic event to be considered separate from Capitanian event. A minority point of view considers
7680-600: Is likely attributable to their ability to thrive in a wide range of environmental conditions. Conodonts saw a rapid recovery during the Induan, with anchignathodontids experiencing a diversity peak in the earliest Induan. Gondolellids diversified at the end of the Griesbachian; this diversity spike was most responsible for the overall conodont diversity peak in the Smithian. Segminiplanate conodonts again experienced
7872-451: Is likely that post-extinction microbial mats played a vital, indispensable role in the survival and recovery of various bioturbating organisms. The microbialite refuge hypothesis has been criticised as reflecting a taphonomic bias due to the greater preservation potential of microbialite deposits, however, rather than a genuine phenomenon. Ichnocoenoses show that marine ecosystems recovered to pre-extinction levels of ecological complexity by
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8064-672: Is now northern India and Pakistan, driving east through Fergana. Kushan power also consolidated long-distance trade, linking Central Asia to both Han dynasty China and the Roman Empire in Europe. The Kushans ruled the area as part of their larger empire until the 3rd century AD, when the Zoroastrian Persian Sassanid Empire invaded Kushan territory from the southwest. Fergana remained under shifting local and Transoxian rulers thereafter. For periods in
8256-542: Is of Persian origin. The valley has been set foot on by various peoples. The Fergana Valley is an intermountain depression in Central Asia, between the mountain systems of the Tien Shan in the north and the Alay in the south. The valley is approximately 300 kilometres (190 mi) long and up to 70 kilometres (43 mi) wide, forming an area covering 22,000 square kilometres (8,500 sq mi). Its position makes it
8448-476: Is played throughout the Central Asian region, the countries sometimes organise Buzkashi competition amongst each other. The First regional competition among the Central Asian countries, Russia , Chinese Xinjiang and Turkey was held in 2013. The first world title competition was played in 2017 and won by Kazakhstan . Association football is popular across Central Asia. Most countries are members of
8640-525: The Bowen Basin of Queensland indicates numerous intermittent periods of marine environmental stress from the middle to late Lopingian leading up to the end-Permian extinction proper, supporting aspects of the gradualist hypothesis. Additionally, the decline in marine species richness and the structural collapse of marine ecosystems may have been decoupled as well, with the former preceding the latter by about 61,000 years according to one study. Whether
8832-700: The Central Asian Football Association , a region of the Asian Football Confederation . However, Kazakhstan is a member of the UEFA . Wrestling is popular across Central Asia, with Kazakhstan having claimed 14 Olympic medals, Uzbekistan seven, and Kyrgyzstan three. As former Soviet states, Central Asian countries have been successful in gymnastics . Mixed Martial Arts is one of more common sports in Central Asia, Kyrgyz athlete Valentina Shevchenko holding
9024-936: The Han-Dayuan war , installing a king there. Later the Han set up the Protectorate of the Western Regions The Kushan Empire formed from the same Yuezhi who had conquered the Hellenistic Fergana. The Kushan spread out in the 1st century AD from the Yuezhi confederation in the territories of ancient Bactria on either side of the middle course of the Oxus River or Amu Darya in what is now northern Afghanistan, and southern Tajikistan and Uzbekistan. The Kushan conquered most of what
9216-688: The Hellenistic Greco-Bactrian Kingdom . The Greek city of Alexandria on the Oxus was apparently burnt to the ground by the Yuezhi around 145 BC. Pushed by these twin forces, the Greco-Bactrian Kingdom reoriented itself around lands in what is now Afghanistan, while the new invaders were partially assimilated into the Hellenistic culture left in Fergana Valley. According to the Han dynasty Records of
9408-687: The Iron Age . Certain Turkic ethnic groups, specifically the Kazakhs , display even higher East Asian ancestry. This is explained by substantial Mongolian influence on the Kazakh genome, through significant admixture between blue eyes, blonde hair, the medieval Kipchaks of Central Asia and the invading medieval Mongolians. The data suggests that the Mongol invasion of Central Asia had lasting impacts onto
9600-566: The Kashmir Valley of India may also be included. The Tibetans and Ladakhis are also included. Most of the mentioned peoples are considered the "indigenous" peoples of the vast region. Central Asia is sometimes referred to as Turkestan . Central Asia is a region of varied geography, including high passes and mountains ( Tian Shan ), vast deserts ( Kyzyl Kum , Taklamakan ), and especially treeless, grassy steppes . The vast steppe areas of Central Asia are considered together with
9792-547: The Khanate of Kokand , Kokand was capital of a territory stretching over modern eastern Uzbekistan and Tajikistan, southern Kazakhstan and all of Kyrgyzstan . Fergana was a province of Russian Turkestan , formed in 1876 out of the former khanate of Kokand . It was bounded by the provinces of Syr-darya in the North and Northwest, Samarkand in the West, and Zhetysu in the Northeast, by Chinese Turkestan ( Kashgaria ) in
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#17327654944949984-670: The Kopet Dagh near the Persian border. East of the Kopet Dagh is the important oasis of Merv and then a few places in Afghanistan like Herat and Balkh . Two projections of the Tian Shan create three "bays" along the eastern mountains. The largest, in the north, is eastern Kazakhstan, traditionally called Jetysu or Semirechye which contains Lake Balkhash . In the center is the small but densely-populated Ferghana valley . In
10176-598: The Middle Triassic ) due to the severity of the extinction. However, studies in Bear Lake County , near Paris, Idaho , and nearby sites in Idaho and Nevada showed a relatively quick rebound in a localized Early Triassic marine ecosystem ( Paris biota ), taking around 1.3 million years to recover, while an unusually diverse and complex ichnobiota is known from Italy less than a million years after
10368-404: The Roadian (middle Permian), suffered a selective extinction pulse 10 million years before the main event, at the end of the Capitanian stage. In this preliminary extinction, which greatly reduced disparity , or the range of different ecological guilds, environmental factors were apparently responsible. Diversity and disparity fell further until the P–Tr boundary; the extinction here (P–Tr)
10560-884: The UFC Flyweight Champion title. Cricket is the most popular sport in Afghanistan . The Afghanistan national cricket team , first formed in 2001, has claimed wins over Bangladesh, West Indies and Zimbabwe. Notable Kazakh competitors include cyclists Alexander Vinokourov and Andrey Kashechkin , boxer Vassiliy Jirov and Gennady Golovkin , runner Olga Shishigina , decathlete Dmitriy Karpov , gymnast Aliya Yussupova , judoka Askhat Zhitkeyev and Maxim Rakov , skier Vladimir Smirnov , weightlifter Ilya Ilyin , and figure skaters Denis Ten and Elizabet Tursynbaeva . Notable Uzbekistani competitors include cyclist Djamolidine Abdoujaparov , boxer Ruslan Chagaev , canoer Michael Kolganov , gymnast Oksana Chusovitina , tennis player Denis Istomin , chess player Rustam Kasimdzhanov , and figure skater Misha Ge . Since gaining independence in
10752-404: The Yuezhi from the east and the Sakas Scythians from the south. After 155 BC, the Yuezhi were pushed into Fergana by the alliance of the powerful Xiongnu and the neighboring Wusun from the north and east, invaded urban civilization of the Dayuan , eventually settling on the northern bank of the Oxus in the region of Transoxiana in modern-day Tajikistan and Uzbekistan , just north of
10944-406: The ocean acidification that resulted from increased atmospheric CO 2 . Organisms that relied on haemocyanin or haemoglobin for transporting oxygen were more resistant to extinction than those utilising haemerythrin or oxygen diffusion. There is also evidence that endemism was a strong risk factor influencing a taxon's likelihood of extinction. Bivalve taxa that were endemic and localised to
11136-406: The 18th century as a result of a long struggle with the Dzungars . The Russian Empire conquered the lands of the nomadic Kazakhs, Turkmens, Kyrgyz and Central Asian khanates in the 19th century. A major revolt known as the Dungan Revolt occurred in the 1860s and 1870s in the eastern part of Central Asia, and Qing rule almost collapsed in all of East Turkestan. After the Russian Revolution ,
11328-497: The 18th century was Tibetan Buddhist and would sometimes travel from Beijing to other cities for personal religious worship. Central Asia also has an indigenous form of improvisational oral poetry that is over 1000 years old. It is principally practiced in Kyrgyzstan and Kazakhstan by akyns , lyrical improvisationalists. They engage in lyrical battles , the aytysh or the alym sabak . The tradition arose out of early bardic oral historians . They are usually accompanied by
11520-534: The 19th century, a considerable trade carried on with Russia: raw cotton, raw silk , tobacco, hides, sheepskins, fruit and cotton and leather goods were exported, and manufactured wares, textiles, tea and sugar were imported and in part re-exported to Kashgaria and Bokhara. The total trade of Fergana reached an annual value of nearly £3.5 million in 1911. Nowadays it suffers from the same depression that affects all trade that either originates in or has to pass through Uzbekistan. The only significant international export
11712-431: The 2020 and 2019 IWB World Competitiveness rankings. In particular, they have been modernizing the industrial sector and fostering the development of service industries through business-friendly fiscal policies and other measures, to reduce the share of agriculture in GDP. Between 2005 and 2013, the share of agriculture dropped in all but Tajikistan, where it increased while industry decreased. The fastest growth in industry
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#173276549449411904-424: The 4th and 5th centuries, the Sassanid Empire directly controlled Transoxiana and Fergana, led by the conquests of Shapur II and Khosrau I against the Kushans and the Hephthalite Empire . Sassanid rule of Fergana was interrupted by the Hepthalites , possibly Turkic. Hepthalite rule was ended by the Gokturks in mid of 6th century. The Turkic Khaganates ruled it until the first quarter of 8th century when it
12096-490: The Anisian. Metazoan reefs became common again during the Anisian because the oceans cooled down then from their overheated state during the Early Triassic. Biodiversity amongst metazoan reefs did not recover until well into the Anisian, millions of years after non-reef ecosystems recovered their diversity. Microbially induced sedimentary structures (MISS) from the earliest Triassic have been found to be associated with abundant opportunistic bivalves and vertical burrows, and it
12288-437: The Caspian Sea in the west, the Altai mountains in the north and the Hindu Kush and Pamir mountains in the South. He did not give an eastern border for the region. His legacy is still seen: Humboldt University of Berlin , named after him, offers a course in Central Asian studies . The Russian geographer Nikolaĭ Khanykov questioned the latitudinal definition of Central Asia and preferred a physical one of all countries located in
12480-452: The Chinese as unusual in features, with a sophisticated urban civilization, similar to that of the Bactrians and Parthians : "The Son of Heaven on hearing all this reasoned thus: Fergana (Dayuan) and the possessions of Bactria and Parthia are large countries, full of rare things, with a population living in fixed abodes and given to occupations somewhat identical with those of the Chinese people, but with weak armies, and placing great value on
12672-655: The Early Triassic; and they dominated many surviving communities across the recovery from the mass extinction. Microbialite deposits appear to have declined in the early Griesbachian synchronously with a significant sea level drop that occurred then. Metazoan-built reefs reemerged during the Olenekian, mainly being composed of sponge biostrome and bivalve builups. Keratose sponges were particularly noteworthy in their integral importance to Early Triassic microbial-metazoan reef communities, and they helped to create stability in heavily damaged ecosystems during early phases of biotic recovery. " Tubiphytes "-dominated reefs appeared at
12864-487: The East, and by Bukhara and Afghanistan in the South. Its southern limits, in the Pamirs , were fixed by an Anglo-Russian commission in 1885, from Zorkul (Victoria Lake) to the Chinese frontier; and Khignan , Roshan and Wakhan were assigned to Afghanistan in exchange for part of Darvaz (on the left bank of the Panj ), which was given to Bukhara . The area amounted to some 53,000 km (20,463 sq mi), of which 17,600 km (6,795 sq mi) are in
13056-469: The Grand Historian or Shiji , based on the travels of Zhang Qian and published around 126 BC, the region of Fergana is presented as the country of the Dayuan (Ta-Yuan), possibly descendants of Greeks colonists (Dayuan may be a transliteration of "Great Ionians"). The area was renowned for its Heavenly Horses , which the Chinese tried to obtain from the Dayuan with little success until they waged war against them in 104 BC. The Dayuan were identified by
13248-553: The Great founded the city of Alexandria Eschate "The Furthest", probably renaming Cyropolis. This was in the southwestern part of the Fergana Valley, on the southern bank of the river Syr Darya (ancient Jaxartes), at the location of the modern city of Khujand , in the state of Tajikistan . Supplemented by Alexander's veterans, it was later ruled by Seleucids before secession of Bactria. The Hellenistic settlements, such as Alexandria Eschate, serve as vibrant examples of cultural and artistic exchanges that occurred following Alexander
13440-401: The Great's conquests. Among the artifacts that highlight this fusion is the Ferghana horse sculpture from the Greco-Bactrian Kingdom, dating back to around 4 to 1 BCE. The Ferghana horse sculpture, a ceremonial gilt bronze finial from the 4th-1st century BCE, exemplifies the Greco-Bactrian Kingdom's artistic synthesis of Greek and Scythian influences. Capturing a horse in the suspension phase of
13632-405: The Greco-Bactrian king Euthydemus extended his control to Sogdiana. There are indications that from Alexandria Eschate the Greco-Bactrians may have led expeditions as far as Kashgar and Ürümqi in Chinese Turkestan , leading to the first known contacts between China and the West around 220 BC. Several statuettes and representations of Greek soldiers have been found north of the Tian Shan , on
13824-804: The Karoo Basin indicates a protracted extinction lasting a million years. Other evidence from the Karoo deposits suggest it took 50,000 years or less, while a study of coprolites in the Vyazniki fossil beds in Russia suggests it took only a few thousand years. Aridification induced by global warming was the chief culprit behind terrestrial vertebrate extinctions. There is enough evidence to indicate that over two thirds of terrestrial labyrinthodont amphibians , sauropsid ("reptile") and therapsid ("proto-mammal") taxa became extinct. Large herbivores suffered
14016-484: The Late Cretaceous to recover their full diversity. Crinoids ("sea lilies") suffered a selective extinction, resulting in a decrease in the variety of their forms. Though cladistic analyses suggest the beginning of their recovery to have taken place in the Induan, the recovery of their diversity as measured by fossil evidence was far less brisk, showing up in the late Ladinian. Their adaptive radiation after
14208-514: The Middle Triassic, with the exception of a notable Ladinian fauna from the Catalonian Basin. Microbial reefs were common across shallow seas for a short time during the earliest Triassic, predominating in low latitudes while being rarer in higher latitudes, occurring both in anoxic and oxic waters. Polybessurus -like microfossils often dominated these earliest Triassic microbialites . Microbial-metazoan reefs appeared very early in
14400-420: The PTME, being the most severely affected clade among the lophophorates. Deep water sponges suffered a significant diversity loss and exhibited a decrease in spicule size over the course of the PTME. Shallow water sponges were affected much less strongly; they experienced an increase in spicule size and much lower loss of morphological diversity compared to their deep water counterparts. Foraminifera suffered
14592-666: The PTME, but some tentative evidence suggests they may have survived into the Triassic. Freshwater and euryhaline fishes, having experienced minimal diversity losses before the PTME, were unaffected during the PTME and actually appear to have increased in diversity across the Permian-Triassic boundary. However, faunal turnovers in freshwater fish communities occurred in areas like the Kuznetsk Basin. The groups that survived suffered extremely heavy losses of species and some terrestrial vertebrate groups very nearly became extinct at
14784-517: The PTME. The Cordaites flora, which dominated the Angaran floristic realm corresponding to Siberia, collapsed over the course of the extinction. In the Kuznetsk Basin , the aridity-induced extinction of the regions's humid-adapted forest flora dominated by cordaitaleans occurred approximately 252.76 Ma, around 820,000 years before the end-Permian extinction in South China, suggesting that
14976-616: The Pamirs. Not all the inhabitants of the area were happy with this state of affairs. In 1898 Muhammed Ali Khalfa proclaimed a jihad against the Russians. However, after about 20 Russians had been killed, Khalfa was captured and executed. When the 1905 Revolution spread across the Russian Empire, some Jadids were active in the Fergana Valley. When the Tsarist regime extended the military draft to include Muslims , this led to
15168-566: The Permian mass extinction event, both complex and simple marine ecosystems were equally common. After the recovery from the mass extinction, the complex communities outnumbered the simple communities by nearly three to one, and the increase in predation pressure and durophagy led to the Mesozoic Marine Revolution . Marine vertebrates recovered relatively quickly, with complex predator-prey interactions with vertebrates at
15360-447: The Permian-Triassic boundary are highly variable depending on the location and preservation quality of any given site. Plants are relatively immune to mass extinction, with the impact of all the major mass extinctions "insignificant" at a family level. Floral diversity losses were more superficial than those of marine animals. Even the reduction observed in species diversity (of 50%) may be mostly due to taphonomic processes. However,
15552-462: The Permian-Triassic boundary, with this flora's collapse being less constrained in western Gondwana but still likely occurring a few hundred thousand years before the boundary. The collapse of this flora is indirectly marked by an abrupt change in river morphology from meandering to braided river systems, signifying the widespread demise of rooted plants. Palynological or pollen studies from East Greenland of sedimentary rock strata laid down during
15744-526: The Permian-Triassic mass extinction marked a key turning point in this ecological shift that began after the Capitanian mass extinction and culminated in the Late Jurassic . Typical taxa of shelly benthic faunas were now bivalves , snails , sea urchins and Malacostraca , whereas bony fishes and marine reptiles diversified in the pelagic zone . On land, dinosaurs and mammals arose in
15936-864: The Permian. Statistical analyses of some highly fossiliferous strata in Meishan, Zhejiang Province in southeastern China, suggest that the main extinction was clustered around one peak, while a study of the Liangfengya section found evidence of two extinction waves, MEH-1 and MEH-2, which varied in their causes, and a study of the Shangsi section showed two extinction pulses with different causes too. Recent research shows that different groups became extinct at different times; for example, while difficult to date absolutely, ostracod and brachiopod extinctions were separated by around 670,000 to 1.17 million years. Palaeoenvironmental analysis of Lopingian strata in
16128-520: The Permian–Triassic boundary. The best-known record of vertebrate changes across the Permian–Triassic boundary occurs in the Karoo Supergroup of South Africa , but statistical analyses have so far not produced clear conclusions. One study of the Karoo Basin found that 69% of terrestrial vertebrates went extinct over 300,000 years leading up to the Permian-Triassic boundary, followed by a minor extinction pulse involving four taxa that survived
16320-476: The Russian conquest. In the southeast is the route to India. In early times Buddhism spread north and throughout much of history warrior kings and tribes would move southeast to establish their rule in northern India. Most nomadic conquerors entered from the northeast. After 1800 western civilisation in its Russian and Soviet form penetrated from the northwest. Because Central Asia is landlocked and not buffered by
16512-520: The Russian-Ukrainian steppe and eastward into the steppes and deserts of Dzungaria and Mongolia. Southward the land becomes increasingly dry and the nomadic population increasingly thin. The south supports areas of dense population and cities wherever irrigation is possible. The main irrigated areas are along the eastern mountains, along the Oxus and Jaxartes Rivers and along the north flank of
16704-501: The Soviet Central Asian Republics". An alternative method is to define the region based on ethnicity, and in particular, areas populated by Eastern Turkic , Eastern Iranian , or Mongolian peoples. These areas include Xinjiang Uyghur Autonomous Region, the Turkic regions of southern Siberia , the five republics, and Afghan Turkestan . Afghanistan as a whole, the northern and western areas of Pakistan and
16896-499: The Soviet Union in 1991, the leaders of the four former Soviet Central Asian Republics met in Tashkent and declared that the definition of Central Asia should include Kazakhstan as well as the original four included by the Soviets. Since then, this has become the most common definition of Central Asia. In 1978, UNESCO defined the region as "Afghanistan, north-eastern Iran , Pakistan , northern India , western China, Mongolia and
17088-495: The Soviet Union to Central Asia and Siberia . According to Touraj Atabaki and Sanjyot Mehendale, "From 1959 to 1970, about two million people from various parts of the Soviet Union migrated to Central Asia, of which about one million moved to Kazakhstan." With the collapse of the Soviet Union , five countries gained independence, that is, Kazakhstan , Kyrgyzstan , Tajikistan , Turkmenistan , and Uzbekistan . The historian and Turkologist Peter B. Golden explains that without
17280-591: The Soviet Union. Afghanistan remained relatively independent of major influence by the Soviet Union until the Saur Revolution of 1978. The Soviet areas of Central Asia saw much industrialisation and construction of infrastructure , but also the suppression of local cultures, hundreds of thousands of deaths from failed collectivisation programmes, and a lasting legacy of ethnic tensions and environmental problems. Soviet authorities deported millions of people, including entire nationalities, from western areas of
17472-432: The Spathian and Anisian. Accordingly, low levels of interspecific competition in seafloor communities that are dominated by primary consumers correspond to slow rates of diversification and high levels of interspecific competition among nektonic secondary and tertiary consumers to high diversification rates. Other explanations state that life was delayed in its recovery because grim conditions returned periodically over
17664-609: The Spathian. Despite high taxonomic turnover, the ecological life modes of Early Triassic ostracods remained rather similar to those of pre-PTME ostracods. Bryozoans in the Early Triassic were restricted to the Boreal realm. They were also not diverse, represented mainly by members of Trepostomatida . During the Middle Triassic, there was a rise in bryozoan diversity, which peaked in the Carnian. However, bryozoans took until
17856-446: The Triassic, taking over niches that were filled primarily by brachiopods before the mass extinction event. Bivalves were once thought to have outcompeted brachiopods, but this outdated hypothesis about the brachiopod-bivalve transition has been disproven by Bayesian analysis . The success of bivalves in the aftermath of the extinction event may have been a function of them possessing greater resilience to environmental stress compared to
18048-711: The Turkification of Central Asia, while extending its cultural influence. The Tang Chinese were defeated by the Abbasid Caliphate at the Battle of Talas in 751, marking the end of the Tang dynasty's western expansion and the 150 years of Chinese influence. The Tibetan Empire would take the chance to rule portions of Central Asia and South Asia. During the 13th and 14th centuries, the Mongols conquered and ruled
18240-522: The Turkmens have a large percentage from populations to the east, their main components are Central Asian. The study further suggests that both migration and linguistic assimilation helped to spread the Turkic languages in Eurasia. The Tang dynasty of China expanded westwards and controlled large parts of Central Asia, directly and indirectly through their Turkic vassals. Tang China actively supported
18432-476: The amount today is about 422 ppm ). There is also evidence of increased ultraviolet radiation reaching the earth, causing the mutation of plant spores. It has been suggested that the Permian–Triassic boundary is associated with a sharp increase in the abundance of marine and terrestrial fungi , caused by the sharp increase in the amount of dead plants and animals fed upon by the fungi. This "fungal spike" has been used by some paleontologists to identify
18624-492: The appearance of modernism, which took inspiration from the Russian avant-garde movement. Until the 1980s, Central Asian arts had developed along with general tendencies of Soviet arts. In the 90s, arts of the region underwent some significant changes. Institutionally speaking, some fields of arts were regulated by the birth of the art market, some stayed as representatives of official views, while many were sponsored by international organisations. The years of 1990–2000 were times for
18816-764: The area from the Mediterranean Sea . Average monthly precipitation is very low from July to September, rises in autumn (October and November) and is highest in March or April, followed by swift drying in May and June. Winds can be strong, producing dust storms sometimes, especially toward the end of the summer in September and October. Specific cities that exemplify Central Asian climate patterns include Tashkent and Samarkand , Uzbekistan, Ashgabat , Turkmenistan, and Dushanbe , Tajikistan. The last of these represents one of
19008-675: The area, along with most of central Asia as fiefs from the Golden Horde of the Mongol Empire. The Fergana became part of a larger Turco-Mongol empire. This Mongolian nomadic confederation known as Barlas , were remnants of the original Mongol army of Genghis Khan . After the Mongol conquest of Central Asia, the Barlas settled in Turkistan (which then became also known as Moghulistan - "Land of Mongols") and intermingled to
19200-677: The area. Modern-day Central Asia is home to a large population of European settlers , who mostly live in Kazakhstan: 7 million Russians, 500,000 Ukrainians , and about 170,000 Germans . During the Stalinist period, the forced deportation of Koreans in the Soviet Union resulted in a population of over 300,000 Koreans in the region. Central Asia has a population of about 72 million, in five countries: Kazakhstan (19 million), Kyrgyzstan (7 million), Tajikistan (10 million), Turkmenistan (6 million), and Uzbekistan (35 million). One of
19392-466: The borders. Located in an arid region, the Fergana valley owes its fertility to two rivers, the Naryn and the Kara Darya , which run from the east, joining near Namangan , forming the Syr Darya river . The valley's history stretches back over 2,300 years, when Alexander the Great founded Alexandria Eschate at its southwestern end. Chinese chroniclers date its towns to more than 2,100 years ago, as
19584-423: The brachiopods that they coexisted with, whilst other studies have emphasised the greater niche breadth of the former. The rise of bivalves to taxonomic and ecological dominance over brachiopods was not synchronous, however, and brachiopods retained an outsized ecological dominance into the Middle Triassic even as bivalves eclipsed them in taxonomic diversity. Some researchers think the brachiopod-bivalve transition
19776-412: The city by Abdallah, the governor of Khurasan. By the time of Ahmad's death in 864 or 865, he was the ruler of most of Transoxiana , Bukhara and Khwarazm . Samarkand and Fergana went to his son, Nasr I of Samanid , leading to a series of Samanid dynasty Muslim rulers of the valley. During demise of Samanids in 10th century, Fergana Valley was conquered by Karakhanids . Eastern part of Fergana later
19968-480: The course of the Triassic . The profound change in the taxonomic composition was partly a result of the selectivity of the extinction event, which affected some taxa (e.g., brachiopods ) more severely than others (e.g., bivalves ). However, recovery was also differential between taxa. Some survivors became extinct some million years after the extinction event without having rediversified ( dead clade walking , e.g.
20160-635: The course of the Early Triassic, causing further extinction events, such as the Smithian-Spathian boundary extinction . Continual episodes of extremely hot climatic conditions during the Early Triassic have been held responsible for the delayed recovery of oceanic life, in particular skeletonised taxa that are most vulnerable to high carbon dioxide concentrations. The relative delay in the recovery of benthic organisms has been attributed to widespread anoxia, but high abundances of benthic species contradict this explanation. A 2019 study attributed
20352-465: The die-off of plants being their likely cause. Wildfires too likely played a role in the fall of Gigantopteris . A conifer flora in what is now Jordan, known from fossils near the Dead Sea , showed unusual stability over the Permian-Triassic transition, and appears to have been only minimally affected by the crisis. The tempo of the terrestrial vertebrate extinction is disputed. Some evidence from
20544-423: The dissimilarity of recovery times between different ecological communities to differences in local environmental stress during the biotic recovery interval, with regions experiencing persistent environmental stress post-extinction recovering more slowly, supporting the view that recurrent environmental calamities were culpable for retarded biotic recovery. Recurrent Early Triassic environmental stresses also acted as
20736-514: The diversion of water from rivers that feed them for irrigation and industrial purposes. Water is an extremely valuable resource in arid Central Asia and can lead to rather significant international disputes. Central Asia is bounded on the north by the forests of Siberia. The northern half of Central Asia (Kazakhstan) is the middle part of the Eurasian steppe . Westward the Kazakh steppe merges into
20928-859: The doorstep to China, and are today on display in the Xinjiang museum at Urumqi (Boardman). Of the Greco-Bactrians, the Greek historian Strabo too writes that: they extended their empire even as far as the Seres (Chinese) and the Phryni . The Fergana area, called Dayuan by the Chinese, remained an integral part of the Greco-Bactrian Kingdom until after the time of Demetrius I of Bactria (c. 120 BC), when confronted with invasions by
21120-401: The early 1990s, the Central Asian republics have gradually been moving from a state-controlled economy to a market economy. However, reform has been deliberately gradual and selective, as governments strive to limit the social cost and ameliorate living standards. All five countries are implementing structural reforms to improve competitiveness. Kazakhstan is the only CIS country to be included in
21312-492: The early 2000s, the Chinese government engaged in a series of human rights abuses against Uyghurs and other ethnic and religious minorities in Xinjiang . At the crossroads of Asia, shamanistic practices live alongside Buddhism . Thus, Yama , Lord of Death, was revered in Tibet as a spiritual guardian and judge. Mongolian Buddhism , in particular, was influenced by Tibetan Buddhism . The Qianlong Emperor of Qing China in
21504-549: The end of the Olenekian, representing the earliest platform-margin reefs of the Triassic, though they did not become abundant until the late Anisian, when reefs' species richness increased. The first scleractinian corals appear in the late Anisian as well, although they would not become the dominant reef builders until the end of the Triassic period. Bryozoans, after sponges, were the most numerous organisms in Tethyan reefs during
21696-440: The end of the Permian. Some of the surviving groups did not persist for long past this period, but others that barely survived went on to produce diverse and long-lasting lineages. However, it took 30 million years for the terrestrial vertebrate fauna to fully recover both numerically and ecologically. It is difficult to analyze extinction and survival rates of land organisms in detail because few terrestrial fossil beds span
21888-504: The end-Permian biotic catastrophe may have started earlier on land and that the ecological crisis may have been more gradual and asynchronous on land compared to its more abrupt onset in the marine realm. In North China, the transition between the Upper Shihhotse and Sunjiagou Formations and their lateral equivalents marked a very large extinction of plants in the region. Those plant genera that did not go extinct still experienced
22080-655: The end-Permian extinction. Additionally, the complex Guiyang biota found near Guiyang , China also indicates life thrived in some places just a million years after the mass extinction, as does a fossil assemblage known as the Shanggan fauna found in Shanggan, China, the Wangmo biota from the Luolou Formation of Guizhou, and a gastropod fauna from the Al Jil Formation of Oman. Regional differences in
22272-631: The establishment of contemporary arts. In the region, many important international exhibitions are taking place, Central Asian art is represented in European and American museums, and the Central Asian Pavilion at the Venice Biennale has been organised since 2005. Equestrian sports are traditional in Central Asia, with disciplines like endurance riding , buzkashi , dzhigit and kyz kuu . The traditional game of Buzkashi
22464-531: The establishment of independent republics, borders have been strongly enforced, though the impact of the new international borders was minor until 1998–2000. Uzbekistan regularly closes its borders with Tajikistan and Kyrgyzstan, strangling trade and causing immense difficulties for those who live in the region. People in the Tajikistan city of Khujand traveling to the Tajik capital of Dushanbe , unable to take
22656-553: The expansion of more habitable climatic zones. Brachiopod taxa during the Anisian recovery interval were only phylogenetically related to Late Permian brachiopods at a familial taxonomic level or higher; the ecology of brachiopods had radically changed from before in the mass extinction's aftermath. Ostracods were extremely rare during the basalmost Early Triassic. Taxa associated with microbialites were disproportionately represented among ostracod survivors. Ostracod recovery began in
22848-415: The extinction event resulted in forms possessing flexible arms becoming widespread; motility , predominantly a response to predation pressure, also became far more prevalent. Though their taxonomic diversity remained relatively low, crinoids regained much of their ecological dominance by the Middle Triassic epoch. Stem-group echinoids survived the PTME. The survival of miocidarid echinoids such as Eotiaris
23040-552: The extinction event. Prior to the extinction, about two-thirds of marine animals were sessile and attached to the seafloor. During the Mesozoic, only about half of the marine animals were sessile while the rest were free-living. Analysis of marine fossils from the period indicated a decrease in the abundance of sessile epifaunal suspension feeders such as brachiopods and sea lilies and an increase in more complex mobile species such as snails , sea urchins and crabs . Before
23232-409: The extinction period indicate dense gymnosperm woodlands before the event. At the same time that marine invertebrate macrofauna declined, these large woodlands died out and were followed by a rise in diversity of smaller herbaceous plants including Lycopodiophyta , both Selaginellales and Isoetales . Data from Kap Stosch suggest that floral species richness was not significantly affected during
23424-543: The fertile abundance of Fergana, as well as the possibility of purchasing further high-quality silk manufactured in Margilan . The most famous export from the region were the 'blood-sweating' Heavenly Horses which captured the imagination of the Chinese during the Han dynasty, but in fact these were almost certainly bred on the Steppe, either west of Bukhara or north of Tashkent , and merely brought to Fergana for sale. In
23616-588: The final extinction killed off only about 80% of marine species alive at that time, whereas the other losses occurred during the first pulse or the interval between pulses. According to this theory, one of these extinction pulses occurred at the end of the Guadalupian epoch of the Permian. For example, all dinocephalian genera died out at the end of the Guadalupian, as did the Verbeekinidae ,
23808-409: The final two sedimentary zones containing conodonts from the Permian. The decrease in diversity was probably caused by a sharp increase in extinctions, rather than a decrease in speciation . The extinction primarily affected organisms with calcium carbonate skeletons, especially those reliant on stable CO 2 levels to produce their skeletons. These organisms were susceptible to the effects of
24000-487: The first geographers to mention Central Asia as a distinct region of the world was Alexander von Humboldt . The borders of Central Asia are subject to multiple definitions. Historically, political geography and culture have been two significant parameters widely used in scholarly definitions of Central Asia. Humboldt's definition comprised every country between 5° North and 5° South of the latitude 44.5°N. Humboldt mentions some geographic features of this region, which include
24192-580: The five months following April precipitation is rare, but increases in frequency starting in October. Snow and frost, down to −20 °C (−4 °F), occurs in December and January. Fergana, on the route to Tarim Basin from the west, remained at the boundaries of a number of classical era empires. As early as 500 BC, the western sections of the Fergana Valley formed part of the Sogdiana region, which
24384-463: The frequency and intensity of dust storms had grown (partly due to poor land use practices). Droughts have already become more likely, and their likelihood is expected to continue increasing with greater climate change. By 2050, people in the Amu Darya basin may be faced with severe water scarcity due to both climate and socioeconomic reasons. Although, during the golden age of Orientalism
24576-497: The full genome of Uzbeks and other Central Asian populations found that about ~27-60% of the Uzbek ancestry is derived from East Asian sources, with the remainder ancestry (~40–73%) being made up by European and Middle Eastern components. According to a recent study, the Kyrgyz, Kazakhs, Uzbeks, and Turkmens share more of their gene pool with various East Asian and Siberian populations than with West Asian or European populations, though
24768-619: The full impact of the event. Many sedimentary sequences from South China show synchronous terrestrial and marine extinctions. Research in the Sydney Basin of the PTME's duration and course also supports a synchronous occurrence of the terrestrial and marine biotic collapses. Other scientists believe the terrestrial mass extinction began between 60,000 and 370,000 years before the onset of the marine mass extinction. Chemostratigraphic analysis from sections in Finnmark and Trøndelag shows
24960-469: The gasification of methane clathrates ; emissions of methane by novel methanogenic microorganisms nourished by minerals dispersed in the eruptions; longer and more intense El Niño events; and an extraterrestrial impact which created the Araguainha crater and caused seismic release of methane and the destruction of the ozone layer with increased exposure to solar radiation. Previously, it
25152-616: The genetic makeup of Kazakhs. According to recent genetic genealogy testing, the genetic admixture of the Uzbeks clusters somewhere between the Iranian peoples and the Mongols . Another study shows that the Uzbeks are closely related to other Turkic peoples of Central Asia and rather distant from Iranian people. The study also analysed the maternal and paternal DNA haplogroups and shows that Turkic speaking groups are more homogenous than Iranian speaking groups. Genetic studies analyzing
25344-730: The great trading cities of Bokhara and Samarkand . The other great commercial city was Tashkent northwest of the mouth of the Ferghana valley. The land immediately north of the Oxus was called Transoxiana and also Sogdia , especially when referring to the Sogdian merchants who dominated the silk road trade. To the east, Dzungaria and the Tarim Basin were united into the Manchu-Chinese province of Xinjiang (Sinkiang; Hsin-kiang) about 1759. Caravans from China usually went along
25536-441: The greater range of environmental tolerance and greater geographic distribution of the former compared to the latter. Cladodontomorph sharks likely survived the extinction by surviving in refugia in the deep oceans, a hypothesis based on the discovery of Early Cretaceous cladodontomorphs in deep, outer shelf environments. Ichthyosaurs , which evolved immediately before the PTME, were also PTME survivors. The Lilliput effect ,
25728-421: The greatest loss of species diversity. In the case of the brachiopods, at least, surviving taxa were generally small, rare members of a formerly diverse community. Conodonts were severely affected both in terms of taxonomic and morphological diversity, although not as severely as during the Capitanian mass extinction. The ammonoids , which had been in a long-term decline for the 30 million years since
25920-628: The heaviest losses. All Permian anapsid reptiles died out except the procolophonids (although testudines have morphologically -anapsid skulls, they are now thought to have separately evolved from diapsid ancestors). Pelycosaurs died out before the end of the Permian. Too few Permian diapsid fossils have been found to support any conclusion about the effect of the Permian extinction on diapsids (the "reptile" group from which lizards, snakes, crocodilians, and dinosaurs (including birds) evolved). Tangasaurids were largely unaffected. Gorgonopsians are traditionally thought to have gone extinct during
26112-439: The horse's power and celestial nature, reflecting the cultural reverence for horses and the era's craftsmanship. Integrating elements of movement and triumph, this sculpture connects deeply to ancient narratives of success and achievement, offering insights into the historical and artistic context of its time. After 250 BC, the city probably remained in contact with the Greco-Bactrian Kingdom centered on Bactria , especially when
26304-588: The imperial manipulations of the Russian Empire but above all the Soviet Union, the creation of said republics would have been impossible. In nearly all the new states, former Communist Party officials retained power as local strongmen. None of the new republics could be considered functional democracies in the early days of independence, although in recent years Kyrgyzstan, Kazakhstan and Mongolia have made further progress towards more open societies, unlike Uzbekistan, Tajikistan, and Turkmenistan, which have maintained many Soviet-style repressive tactics. Beginning in
26496-423: The inarticulate brachiopod Lingularia , and the foraminifera Earlandia and Rectocornuspira kalhori , the latter of which is sometimes classified under the genus Ammodiscus . Their guild diversity was also low. Post-PTME faunas had a flat, insignificant latitudinal diversity gradient. The speed of recovery from the extinction is disputed. Some scientists estimate that it took 10 million years (until
26688-505: The largest contiguous empire in recorded history. Most of Central Asia fell under the control of the Chagatai Khanate . The dominance of the nomads ended in the 16th century, as firearms allowed settled peoples to gain control of the region. Russia , China , and other powers expanded into the region and had captured the bulk of Central Asia by the end of the 19th century. The Qing dynasty gained control of East Turkestan in
26880-724: The mass extinction, exemplifying the Lilliput effect's opposite, which has been dubbed the Brobdingnag effect. The Permian had great diversity in insect and other invertebrate species, including the largest insects ever to have existed. The end-Permian is the largest known mass extinction of insects; according to some sources, it may well be the only mass extinction to significantly affect insect diversity. Eight or nine insect orders became extinct and ten more were greatly reduced in diversity. Palaeodictyopteroids (insects with piercing and sucking mouthparts) began to decline during
27072-512: The mid-Permian; these extinctions have been linked to a change in flora. The greatest decline occurred in the Late Permian and was probably not directly caused by weather-related floral transitions. However, some observed entomofaunal declines in the PTME were biogeographic changes rather than outright extinctions. The geological record of terrestrial plants is sparse and based mostly on pollen and spore studies. Floral changes across
27264-454: The more direct route through Uzbekistan, have to cross a high mountain pass between the two cities instead, along a terrible road. Communications between the Kyrgyzstan cities of Bishkek and Osh pass through difficult mountainous country. Ethnic tensions also flared into riots in 1990 , most notably in the town of Uzgen , near Osh. There has been no further ethnic violence, and things appeared to have quieted down for several years. However,
27456-575: The north or south side of the Tarim basin and joined at Kashgar before crossing the mountains northwest to Ferghana or southwest to Bactria. A minor branch of the silk road went north of the Tian Shan through Dzungaria and Zhetysu before turning southwest near Tashkent. Nomadic migrations usually moved from Mongolia through Dzungaria before turning southwest to conquer the settled lands or continuing west toward Europe. The Kyzyl Kum Desert or semi-desert
27648-411: The oceans . The level of atmospheric carbon dioxide rose from around 400 ppm to 2,500 ppm with approximately 3,900 to 12,000 gigatonnes of carbon being added to the ocean-atmosphere system during this period. Several other contributing factors have been proposed, including the emission of carbon dioxide from the burning of oil and coal deposits ignited by the eruptions; emissions of methane from
27840-547: The ouster of Kyrgyz President Kurmanbek Bakiyev in April 2010. In June 2010, about 200 people have been reported to be killed during clashes in Osh and Jalal-Abad , and 2000 more were injured. Between 100,000 and 300,000 refugees, predominantly of Uzbek ethnic origin, attempted to flee to Uzbekistan, causing a major humanitarian crisis. The area has also been subject to informal radicalization. In Tsarist times, out of some 1,200,000 ha (3,000,000 acres) of cultivated land, about two thirds were under constant irrigation and
28032-475: The pace of biotic recovery existed, which suggests that the impact of the extinction may have been felt less severely in some areas than others, with differential environmental stress and instability being the source of the variance. In addition, it has been proposed that although overall taxonomic diversity rebounded rapidly, functional ecological diversity took much longer to return to its pre-extinction levels; one study concluded that marine ecological recovery
28224-415: The parameters were now shared differently among clades . Ostracods experienced prolonged diversity perturbations during the Changhsingian before the PTME proper, when immense proportions of them abruptly vanished. At least 74% of ostracods died out during the PTME itself. Bryozoans had been on a long-term decline throughout the Late Permian epoch before they suffered even more catastrophic losses during
28416-516: The phenomenon of dwarfing of species during and immediately following a mass extinction event, has been observed across the Permian-Triassic boundary, notably occurring in foraminifera, brachiopods, bivalves, and ostracods. Though gastropods that survived the cataclysm were smaller in size than those that did not, it remains debated whether the Lilliput effect truly took hold among gastropods. Some gastropod taxa, termed "Gulliver gastropods", ballooned in size during and immediately following
28608-405: The place of Central Asia in the world history was marginalised, contemporary historiography has rediscovered the "centrality" of the Central Asia. The history of Central Asia is defined by the area's climate and geography. The aridness of the region made agriculture difficult, and its distance from the sea cut it off from much trade. Thus, few major cities developed in the region; instead, the area
28800-405: The previous extinction interval. Another study of latest Permian vertebrates in the Karoo Basin found that 54% of them went extinct due to the PTME. In the wake of the extinction event, the ecological structure of present-day biosphere evolved from the stock of surviving taxa. In the sea, the "Palaeozoic evolutionary fauna" declined while the "modern evolutionary fauna" achieved greater dominance;
28992-445: The principal products of the domestic or cottage industries. This was not greatly added to in Soviet times, when industrialisation was concentrated in the cities of Samarkand and Bukhara . Historically the Fergana Valley was an important staging-post on the Silk Road for goods and people traveling from China to the Middle East and Europe. After crossing the passes from Kashgar in Xinjiang, traders would have found welcome relief in
29184-456: The quick recovery seen in nektonic organisms such as ammonoids , which exceeded pre-extinction diversities already two million years after the crisis, and conodonts, which diversified considerably over the first two million years of the Early Triassic. Recent work suggests that the pace of recovery was intrinsically driven by the intensity of competition among species, which drives rates of niche differentiation and speciation . That recovery
29376-513: The region landlocked from water, including Afghanistan , Khorasan (Northeast Iran), Kyrgyzstan , Tajikistan , Turkmenistan , Uyghuristan (Xinjiang), Mongolia , and Uzbekistan . Russian culture has two distinct terms: Средняя Азия ( Srednyaya Aziya or "Middle Asia", the narrower definition, which includes only those traditionally non-Slavic, Central Asian lands that were incorporated within those borders of historical Russia) and Центральная Азия ( Tsentralnaya Aziya or "Central Asia",
29568-712: The region under Mongolian influence, which had "enormous demographic success", but did not impact the cultural or linguistic landscape. Once populated by Iranian tribes and other Indo-European speaking people , Central Asia experienced numerous invasions emanating out of Southern Siberia and Mongolia that would drastically affect the region. Genetic data shows that the different Central Asian Turkic-speaking peoples have between ~22% and ~70% East Asian ancestry (represented by "Baikal hunter-gatherer ancestry" shared with other Northeast Asians and Eastern Siberians), in contrast to Iranian-speaking Central Asians, specifically Tajiks , which display genetic continuity to Indo-Iranians of
29760-414: The region was known to the Greeks as Cyropolis , named after Cyrus the Great. The Sogdian Rock or Rock of Ariamazes, a fortress in Sogdiana, was captured in 327 BC by the forces of Alexander the Great ; after an extended campaign putting down Sogdian resistance and founding military outposts manned by his Greek veterans, Alexander united Sogdiana with Bactria into one satrapy . In 329 BC, Alexander
29952-402: The region's cities. Major rivers of the region include the Amu Darya , the Syr Darya , Irtysh , the Hari River and the Murghab River . Major bodies of water include the Aral Sea and Lake Balkhash , both of which are part of the huge west-central Asian endorheic basin that also includes the Caspian Sea . Both of these bodies of water have shrunk significantly in recent decades due to
30144-431: The remaining third under partial irrigation. The soil was considered by the authors of the article in the Encyclopædia Britannica Eleventh Edition to be admirably cultivated, the principal crops having been cotton, wheat , rice , barley , maize , millet , lucerne , tobacco , vegetables and fruit. Gardening was conducted with a high degree of skill and success. Large numbers of horses, cattle and sheep were kept, and
30336-454: The rich produce of China" ( Book of the Later Han ). Agricultural activities of the Dayuan reported by Zhang Qian included cultivation of grain and grapes for wine-making. The area of Fergana was thus the theater of the first major interaction between an urbanized culture speaking Indo-European languages and the Chinese civilization, which led to the opening up the Silk Road from the 1st century BC onwards. The Han later captured Dayuan in
30528-469: The rise of prosperous trade cities. acting as a crossroads for the movement of people, goods, and ideas between Europe and the Far East . Most countries in Central Asia are still integral to parts of the world economy. From the mid-19th century until near the end of the 20th century, Central Asia was colonised by the Russians , and incorporated into the Russian Empire , and later the Soviet Union , which led to Russians and other Slavs migrating into
30720-411: The semi-nomadic Scythians and Dahae . As the result of Turkic migration , Central Asia also became the homeland for the Kazakhs , Kyrgyzs , Tatars , Turkmens , Uyghurs , and Uzbeks ; Turkic languages largely replaced the Iranian languages spoken in the area, with the exception of Tajikistan and areas where Tajik is spoken. The Silk Road trade routes crossed through Central Asia, leading to
30912-412: The sequence of environmental disasters to have effectively constituted a single, prolonged extinction event, perhaps depending on which species is considered. This older theory, still supported in some recent papers, proposes that there were two major extinction pulses 9.4 million years apart, separated by a period of extinctions that were less extensive, but still well above the background level, and that
31104-417: The snail family Bellerophontidae ), whereas others rose to dominance over geologic times (e.g., bivalves). A cosmopolitanism event began immediately after the end-Permian extinction event. Marine post-extinction faunas were mostly species-poor and were dominated by few disaster taxa such as the bivalves Claraia , Unionites , Eumorphotis , and Promyalina , the conodonts Clarkina and Hindeodus ,
31296-466: The south is Bactria , later called Tocharistan, which is bounded on the south by the Hindu Kush mountains of Afghanistan. The Syr Darya (Jaxartes) rises in the Ferghana valley and the Amu Darya (Oxus) rises in Bactria. Both flow northwest into the Aral Sea . Where the Oxus meets the Aral Sea it forms a large delta called Khwarazm and later the Khanate of Khiva . North of the Oxus is the less-famous but equally important Zarafshan River which waters
31488-413: The south of the valley, in the spurs of the Turkestan ridge, there is the Madygen fossil locality , in which more than 15 thousand fossils of Triassic plants ( Ginkgoales , cycads , ferns , Neocalamitaceae ) and animals ( insects , crustaceans , fishes , amphibians ) were discovered. The most numerous are insect fossils which include the large Gigatitan from the extinct order Titanoptera with
31680-453: The steppes of Eastern Europe as a homogeneous geographical zone known as the Eurasian Steppe . Much of the land of Central Asia is too dry or too rugged for farming. The Gobi Desert extends from the foot of the Pamirs , 77° E, to the Great Khingan (Da Hinggan) Mountains, 116°–118° E. Central Asia has the following geographic extremes: A majority of the people earn a living by herding livestock. Industrial activity centers in
31872-427: The terrestrial and marine extinctions were synchronous or asynchronous is another point of controversy. Evidence from a well-preserved sequence in east Greenland suggests that the terrestrial and marine extinctions began simultaneously. In this sequence, the decline of animal life is concentrated in a period approximately 10,000 to 60,000 years long, with plants taking an additional several hundred thousand years to show
32064-415: The terrestrial extinction occurred after the marine extinction in the tropics. Studies of the timing and causes of the Permian-Triassic extinction are complicated by the often-overlooked Capitanian extinction (also called the Guadalupian extinction), just one of perhaps two mass extinctions in the late Permian that closely preceded the Permian-Triassic event. In short, when the Permian-Triassic starts it
32256-446: The terrestrial floral turnover occurred before the large negative δ C shift during the marine extinction. Dating of the boundary between the Dicynodon and Lystrosaurus assemblage zones in the Karoo Basin indicates that the terrestrial extinction occurred earlier than the marine extinction. The Sunjiagou Formation of South China also records a terrestrial ecosystem demise predating the marine crisis. Other research still has found that
32448-426: The throne. During the 8th century, Fergana was the location of fierce rivalry between Tang dynasty China and the expansion of Muslim power. The Umayyads waged several wars against the Sogdian and Turkic population. They were defeated by the Turgesh who came dominated the Ferghana Valley until their defeat by Tang in 750. At the same time, the Abbasids defeated the Umayyads and sent their forces to Central Asia. This
32640-420: The top of the food web being known from coprolites five million years after the PTME. Post-PTME hybodonts exhibited extremely rapid tooth replacement. Ichthyopterygians appear to have ballooned in size extremely rapidly following the PTME. Bivalves rapidly recolonised many marine environments in the wake of the catastrophe. Bivalves were fairly rare before the P–Tr extinction but became numerous and diverse in
32832-426: The valley into a series of small states. From the 16th century, the Shaybanid dynasty of the Khanate of Bukhara ruled Fergana, replaced by the Janid dynasty of Bukhara in 1599. In 1709 Shaybanid emir Shahrukh of the Minglar Uzbeks declared independence from the Khanate of Bukhara , establishing a state in the eastern part of the Fergana Valley. He built a citadel to be his capital in the small town of Kokand . As
33024-405: The valley is a religiously conservative region which was particularly hard-hit by President Karimov 's secularization legislation in Uzbekistan, together with his decision to close the borders with Kyrgyzstan in 2003. This devastated the local economy by preventing the importation of cheap Chinese consumer goods. The deposition of Askar Akayev in Kyrgyzstan in April 2005, coupled with the arrest of
33216-405: The western Central Asian regions were incorporated into the Soviet Union . The eastern part of Central Asia, known as Xinjiang , was incorporated into the People's Republic of China , having been previously ruled by the Qing dynasty and the Republic of China . Mongolia gained its independence from China and has remained independent but became a Soviet satellite state until the dissolution of
33408-447: The wettest climates in Central Asia, with an average annual precipitation of over 560 mm (22 inches). Biogeographically, Central Asia is part of the Palearctic realm . The largest biome in Central Asia is the temperate grasslands, savannas, and shrublands biome. Central Asia also contains the montane grasslands and shrublands , deserts and xeric shrublands and temperate coniferous forests biomes. As of 2022, there has been
33600-404: The wider definition, which includes Central Asian lands that have never been part of historical Russia). The latter definition includes Afghanistan and ' East Turkestan '. The most limited definition was the official one of the Soviet Union , which defined Middle Asia as consisting solely of Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan, omitting Kazakhstan . Soon after the dissolution of
33792-452: Was achieved was most probably due to the influence of the Silk Road , which traveled along Central Asia. Periodically, great leaders or changing conditions would organise several tribes into one force and create an almost unstoppable power. These included the Hun invasion of Europe, the Five Barbarians rebellions in China and most notably the Mongol conquest of much of Eurasia . During pre-Islamic and early Islamic times, Central Asia
33984-588: Was attributable not only to the end-Permian extinction but also the ecological restructuring that began as a result of the Capitanian extinction. Infaunal habits in bivalves became more common after the PTME. Linguliform brachiopods were commonplace immediately after the extinction event, their abundance having been essentially unaffected by the crisis. Adaptations for oxygen-poor and warm environments, such as increased lophophoral cavity surface, shell width/length ratio, and shell miniaturisation, are observed in post-extinction linguliforms. The surviving brachiopod fauna
34176-408: Was for millennia dominated by the nomadic horse peoples of the steppe . Relations between the steppe nomads and the settled people in and around Central Asia were long marked by conflict. The nomadic lifestyle was well suited to warfare , and the steppe horse riders became some of the most militarily potent people in the world, limited only by their lack of internal unity. Any internal unity that
34368-546: Was inhabited predominantly by speakers of Iranian languages . Among the ancient sedentary Iranian peoples , the Sogdians and Chorasmians played an important role, while Iranian peoples such as Scythians and the later on Alans lived a nomadic or semi-nomadic lifestyle. The main migration of Turkic peoples occurred between the 6th and 11th centuries, when they spread across most of Central Asia. The Eurasian Steppe slowly transitioned from Indo European and Iranian -speaking groups with dominant West-Eurasian ancestry to
34560-434: Was instrumental in the assimilation of the Timurid elite to the Perso-Islamic courtly culture. Heir to one of these confederations, Timur , founder of the Timurid dynasty , added the valley to a newly consolidated empire in the late 14th century, ruling the area from Samarkand . Located on the Northern Silk Road , the Fergana played a significant part in the flowering of medieval Central Asian Islam. Its most famous son
34752-400: Was leading to the Battle of Talas in 751, which resulted in a victory for the Abbasids and the disengagement of China from Central Asia. Two antecedent battles in 715 and 717 had seen the Chinese prevail over Arab forces. A series of Arab, Persian, and later Turkic Muslim rulers reigned over the Fergana. The Samanid Empire , rising from the Arab Muslim conquest of Persia , pushed into what
34944-426: Was made a part of it. This blocked the valley's natural outlet and the routes to Samarkand and Bukhara, but none of these borders was of any great significance so long as Soviet rule lasted. The whole region was part of a single economy geared to cotton production on a massive scale, and the overarching political structures meant that crossing borders was not a problem. With the breakup of the Soviet Union in 1991 and
35136-410: Was non-selective, consistent with a catastrophic initiator. During the Triassic, diversity rose rapidly, but disparity remained low. The range of morphospace occupied by the ammonoids, that is, their range of possible forms, shapes or structures, became more restricted as the Permian progressed. A few million years into the Triassic, the original range of ammonoid structures was once again reoccupied, but
35328-465: Was observed in Turkmenistan, whereas the services sector progressed most in the other four countries. Permian%E2%80%93Triassic extinction event The scientific consensus is that the main cause of the extinction was the flood basalt volcanic eruptions that created the Siberian Traps , which released sulfur dioxide and carbon dioxide , resulting in euxinia (oxygen-starved, sulfurous oceans), elevating global temperatures, and acidifying
35520-438: Was ruled from further west and owed fealty to the Achaemenid Empire at the time of Darius the Great . The independent and warlike Sogdiana formed a border region insulating the Achaemenid Persians from the nomadic Scythians to the north and east. It was forcibly settled by exiled Greeks from the Anatolian coast, who had rebelled or otherwise given Persia trouble. Eventually, it had a significant Greek community. The capital of
35712-425: Was slow in the Early Triassic can be explained by low levels of biological competition due to the paucity of taxonomic diversity, and that biotic recovery explosively accelerated in the Anisian can be explained by niche crowding, a phenomenon that would have drastically increased competition, becoming prevalent by the Anisian. Biodiversity rise thus behaved as a positive feedback loop enhancing itself as it took off in
35904-519: Was still ongoing 50 million years after the extinction, during the latest Triassic, even though taxonomic diversity had rebounded in a tenth of that time. The pace and timing of recovery also differed based on clade and mode of life. Seafloor communities maintained a comparatively low diversity until the end of the Early Triassic, approximately 4 million years after the extinction event. Epifaunal benthos took longer to recover than infaunal benthos. This slow recovery stands in remarkable contrast with
36096-454: Was subjugated by the Tang dynasty The Kingdom of Ferghana was ruled by the Ikhshids , who submitted as vassal to the Chinese Tang between 659 and 790. It was attacked by the Tibetan Empire in 715. The Umayyad Caliphate in 715 deposed the ruler, and installed a new king Alutar on the throne. The Chinese sent 10,000 troops under Zhang Xiaosong to Ferghana. He defeated Alutar and the Arab occupation force at Namangan and reinstalled Ikhshid on
36288-447: Was then called Greater Khorasan , including Transoxiana and the Fergana Valley from the West. In 819, Ahmad ibn Asad —son of Asad ibn Saman —was granted authority over the city of Fergana by Caliph Al-Ma'mun 's governor of Khorasan , Ghassan ibn 'Abbad, as a reward for his support against the rebel Rafi' ibn Laith . Following the death of his brother Nuh , who ruled in Samarkand, Ahmad and another brother Yahya were given rule over
36480-404: Was thought that rock sequences spanning the Permian–Triassic boundary were too few and contained too many gaps for scientists to reliably determine its details. However, it is now possible to date the extinction with millennial precision. U–Pb zircon dates from five volcanic ash beds from the Global Stratotype Section and Point for the Permian–Triassic boundary at Meishan , China , establish
36672-445: Was under suzerenaity of Karakhitays . Karakhanid rule lasted till 1212, when Khwarezmshahs conquered the western part of the valley. Genghis Khan invaded Transoxiana and Fergana in 1219 during his conquest of Khwarazm . Before his death in 1227, he assigned the lands of Western Central Asia to his second son Chagatai , and this region became known as the Chagatai Khanate . But it was not long before Transoxian Turkic leaders ruled
36864-435: Was very low in diversity and exhibited no provincialism whatsoever. Brachiopods began their recovery around 250.1 ± 0.3 Ma, as marked by the appearance of the genus Meishanorhynchia , believed to be the first of the progenitor brachiopods that evolved after the mass extinction. Major brachiopod rediversification only began in the late Spathian and Anisian in conjunction with the decline of widespread anoxia and extreme heat and
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