122-552: Hluhluwe–Imfolozi Park , formerly Hluhluwe–Umfolozi Game Reserve , is the oldest proclaimed nature reserve in Africa. It consists of 960 km² (96,000 ha) of hilly topography 280 kilometres (170 mi) north of Durban in central KwaZulu-Natal , South Africa and is known for its rich wildlife and conservation efforts. Operated by Ezemvelo KZN Wildlife , the park is the only state-run park in KwaZulu-Natal where each of
244-600: A wildlife refuge , wildlife sanctuary , biosphere reserve or bioreserve , natural or nature preserve , or nature conservation area ) is a protected area of importance for flora , fauna , funga , or features of geological or other special interest, which is reserved and managed for purposes of conservation and to provide special opportunities for study or research . They may be designated by government institutions in some countries, or by private landowners, such as charities and research institutions . Nature reserves fall into different IUCN categories depending on
366-467: A UNESCO World Heritage Site since 1 December 1999. In Kosovo, there are 2 national parks , 11 nature reserves, 99 natural monuments and 3 protected landscapes . The national parks are Bjeshkët e Nemuna National Park and Sharr Mountains National Park . As of 2011, Poland has 1469 nature reserves. Nature reserves are one of the 11 types of protected areas in Portugal. As of 2012, Portugal had
488-417: A buildup of scent attracting the attention of predators, the lioness moves her cubs to a new den site several times a month, carrying them one-by-one by the nape of the neck. Usually, the mother does not integrate herself and her cubs back into the pride until the cubs are six to eight weeks old. Sometimes the introduction to pride life occurs earlier, particularly if other lionesses have given birth at about
610-549: A closer relationship with P. leo melanochaita . Lion samples from some parts of the Ethiopian Highlands cluster genetically with those from Cameroon and Chad, while lions from other areas of Ethiopia cluster with samples from East Africa. Researchers, therefore, assume Ethiopia is a contact zone between the two subspecies. Genome -wide data of a wild-born historical lion sample from Sudan showed that it clustered with P. l. leo in mtDNA-based phylogenies, but with
732-471: A constant lookout for circling vultures, which indicate the death or distress of an animal. Most carrion on which both hyenas and lions feed upon are killed by hyenas rather than lions. Carrion is thought to provide a large part of lion diet. Lions and spotted hyenas occupy a similar ecological niche and compete for prey and carrion; a review of data across several studies indicates a dietary overlap of 58.6%. Lions typically ignore hyenas unless they are on
854-470: A diverse floral community. The park is the birthplace of rhino preservation, having been responsible for breeding the southern white rhinoceros back from near extinction in the first half of the 20th century. There are reportedly 1,600 white rhino in the reserve. The rhino population remains severely threatened by poaching, with 222 rhinos poached in the province in 2017, most of them in the park. Hluhluwe–Imfolozi has implemented Smart Park which facilitates
976-425: A favoured position in the group, either stalking prey on the "wing", then attacking, or moving a smaller distance in the centre of the group and capturing prey fleeing from other lionesses. Males attached to prides do not usually participate in group hunting. Some evidence suggests, however, that males are just as successful as females; they are typically solo hunters who ambush prey in small bushland. They may join in
1098-943: A federal government, Canada recognizes 55 National Wildlife Areas, containing species of ecological significance. This area is protected by legislation known as the Canada Wildlife Act, which is overseen by the Ministry of Environment and Climate Change. About half of the habitat is marine habitat, and the remainder is terrestrial habitat. Nature Conservancy Canada, Ducks Unlimited, and Escarpment Biosphere Conservancy, among others, protect nature reserves in Canada as well. As part of Canada's Ecological Gifts Program, these charities protect wilderness on privately owned lands. There are 30 nature reserves in Egypt which cover 12% of Egyptian land. Those nature reserves were built according to
1220-470: A few reported cases of old and wounded lions falling prey to wild dogs. Most lionesses reproduce by the time they are four years of age. Lions do not mate at a specific time of year and the females are polyestrous . Like those of other cats, the male lion's penis has spines that point backward. During withdrawal of the penis, the spines rake the walls of the female's vagina, which may cause ovulation . A lioness may mate with more than one male when she
1342-632: A general system for sanctuaries to follow. Among them, the American Sanctuary Association monitors and aids in various facilities to care for exotic wildlife. The number of sanctuaries has substantially increased over the past few years. In Australia , a nature reserve is the title of a type of protected area used in the jurisdictions of the Australian Capital Territory , New South Wales , Tasmania and Western Australia . The term "nature reserve"
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#17327829413591464-548: A heavier mane. On average, Asiatic lions have sparser manes than African lions. This feature likely evolved to signal the fitness of males to females. Males with darker manes appear to have greater reproductive success and are more likely to remain in a pride for longer. They have longer and thicker hair and higher testosterone levels, but they are also more vulnerable to heat stress. The core body temperature does apparently not increase regardless of sex, season, feeding time, length and colour of mane, but only surface temperature
1586-602: A high affinity to P. l. melanochaita . This result suggested that the taxonomic position of lions in Central Africa may require revision. Other lion subspecies or sister species to the modern lion existed in prehistoric times: The Panthera lineage is estimated to have genetically diverged from the common ancestor of the Felidae around 9.32 to 4.47 million years ago to 11.75 to 0.97 million years ago . Results of analyses differ in
1708-501: A kill or are being harassed, while the latter tend to visibly react to the presence of lions with or without the presence of food. In the Ngorongoro crater , lions subsist largely on kills stolen from hyenas, causing them to increase their kill rate. In Botswana's Chobe National Park, the situation is reversed as hyenas there frequently challenge lions and steal their kills, obtaining food from 63% of all lion kills. When confronted on
1830-514: A kill, hyenas may either leave or wait patiently at a distance of 30–100 m (98–328 ft) until the lions have finished. Hyenas may feed alongside lions and force them off a kill. The two species attack one another even when there is no food involved for no apparent reason. Lions can account for up to 71% of hyena deaths in Etosha National Park . Hyenas have adapted by frequently mobbing lions that enter their home ranges. When
1952-519: A kill. Crippled lions and cubs may fall victim to hyenas and leopards or be trampled by buffalo or elephants. Careless lions may be maimed when hunting prey. Nile crocodiles may also kill and eat lions, evidenced by the occasional lion claw found in crocodile stomachs. Ticks commonly infest the ears, neck and groin regions of the lions. Adult forms of several tapeworm species of the genus Taenia have been isolated from lion intestines, having been ingested as larvae in antelope meat. Lions in
2074-558: A lion makes when sniffing chemical signals and involves an open mouth with bared teeth, raised muzzle, wrinkled nose, closed eyes and relaxed ears. Lions also use chemical and visual marking; males spray urine and scrape plots of ground and objects within the territory. The lion's repertoire of vocalisations is large; variations in intensity and pitch appear to be central to communication. Most lion vocalisations are variations of growling , snarling , meowing and roaring. Other sounds produced include puffing, bleating and humming. Roaring
2196-473: A longer evolutionary history than genetically less diverse lion samples from Asia and West and Central Africa. A whole genome-wide sequence of lion samples showed that samples from West Africa shared alleles with samples from Southern Africa, and samples from Central Africa shared alleles with samples from Asia. This phenomenon indicates that Central Africa was a melting pot of lion populations after they had become isolated, possibly migrating through corridors in
2318-399: A loose pride while females associate with up to 12 other females, forming a stronger pride together with their cubs. Female and male lions associate only when mating. Coalitions of males hold territory for a longer time than single lions. Males in coalitions of three or four individuals exhibit a pronounced hierarchy, in which one male dominates the others and mates more frequently. The lion
2440-433: A nomad is a "range". Males associated with a pride patrol the fringes. Both males and females defend the pride against intruders, but the male lion is better-suited for this purpose due to its stockier, more powerful build. Some individuals consistently lead the defense against intruders, while others lag behind. Lions tend to assume specific roles in the pride; slower-moving individuals may provide other valuable services to
2562-554: A nomadic phase before gaining residence in a pride. A study undertaken in the Serengeti National Park revealed that nomadic coalitions gain residency at between 3.5 and 7.3 years of age. In Kruger National Park, dispersing male lions move more than 25 km (16 mi) away from their natal pride in search of their own territory. Female lions stay closer to their natal pride. Therefore, female lions in an area are more closely related to each other than male lions in
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#17327829413592684-425: A reed-bed, a cave, or some other sheltered area, usually away from the pride. She will often hunt alone while the cubs are still helpless, staying relatively close to the den. Lion cubs are born blind, their eyes opening around seven days after birth. They weigh 1.2–2.1 kg (2.6–4.6 lb) at birth and are almost helpless, beginning to crawl a day or two after birth and walking around three weeks of age. To avoid
2806-566: A rhino has been killed in Hluhluwe Imfolozi." In 1981, the Natal Parks board (now Ezemvelo KZN Wildlife) attempted to reintroduce African wild dogs into the park. Twenty-three dogs were released in the reserve, most of which had been bred in zoos. However this met with limited success and by 2015, the population had fluctuated between 3 and 30 individuals. Further dogs were released into the park in 2022. The first visitor camp
2928-640: A royal hunting ground for the Zulu kingdom in the time of Shaka . The southern white rhino, first identified by Western naturalist William John Burchell in 1812, was virtually eliminated during the 19th century by European hunters, and by 1895 was believed to be extinct. A population of between 20 and 100 was identified in South Africa and preserved by establishing the Umfolozi Junction Reserve and Hluhluwe Reserve, which are now parts of
3050-565: A short, rounded head, a reduced neck, and round ears; males have broader heads. The fur varies in colour from light buff to silvery grey, yellowish red, and dark brown. The colours of the underparts are generally lighter. A new-born lion has dark spots , which fade as the cub reaches adulthood, although faint spots may still be seen on the legs and underparts. The tail of all lions ends in a dark, hairy tuft that, in some lions, conceals an approximately 5 mm (0.20 in)-long, hard "spine" or "spur" composed of dermal papillae. The functions of
3172-769: A total area of approximately 1,330 km (510 sq mi). Section 21 of the National Parks and Access to the Countryside Act 1949 gives local authorities the exclusive statutory power to establish a local nature reserve in consultation with the SNH. The biosphere reserves of Iran have a total land area of 1.64 million km . The "reserves" support more than 8,000 recorded species of plants (almost 2,421 are endemic ), 502 species of birds , 164 species of mammals , 209 species of reptiles , and 375 species of butterflies . India 's 18 biospheres extend over
3294-656: A total of 46 protected areas, which represented 6,807.89 km (2,628.54 sq mi) of land and 463.94 km (179.13 sq mi) of marine surfaces. Among the protected areas, nine are classified as nature reserve ( Portuguese : reserva natural ). About 5.18% of the area of Romania has a protected status (12,360 km (4,770 sq mi)), including the Danube Delta , which makes up half of this area (2.43% of Romania's total area). There are 15 National Parks, and around 90 Natural Parks in Spain. Spain
3416-693: A total of 85,940 km (33,180 sq mi) and protect larger areas than typical national parks in other countries. The first national reserve of India was established in 1986. Israel 's national parks are declared historic sites or nature reserves, which are mostly operated and maintained by the National Nature and Parks Authority. As of 2019, Israel maintains more than 490 nature reserves that protect 2,500 species of indigenous wild plants, 20 species of fish, 530 species of birds and 70 species of mammals. In total, they cover 6,400 km (2,500 sq mi) of nature reserves, approximately 28% of
3538-630: A wildlife refuge since very few people live in the area. Wildlife has flourished in the zone since the Chernobyl nuclear accident in 1986. South Africa is well known for its many nature reserves. The oldest nature reserve in the country (and oldest conservation area in the world) is the Groenkloof Nature Reserve that was established in 1892 in the capital city Pretoria in the old South African Republic and current Republic of South Africa. The country has many national parks but
3660-429: A year old and begin to hunt effectively when nearing the age of two. Single lions are capable of bringing down zebra and wildebeest, while larger prey like buffalo and giraffe are riskier. In Chobe National Park , large prides have been observed hunting African bush elephants up to around 15 years old in exceptional cases, with the victims being calves, juveniles, and even subadults. In typical hunts, each lioness has
3782-837: Is Rye Harbour Nature Reserve in East Sussex, where a network of footpaths enables visitors to explore shingle, saltmarsh, saline lagoon, reedbed, and grazing marsh habitats. Through the Natural Heritage (Scotland) Act 1991 the Scottish Natural Heritage (SNH) was established in 1992 as a government body, responsible to the Scottish Government Ministers and through them to the Scottish Parliament. At 31 March 2008, there were 65 Scottish national nature reserves with
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3904-404: Is in heat . Lions of both sexes may be involved in group homosexual and courtship activities. Males will also head-rub and roll around with each other before mounting each other. Generation length of the lion is about seven years. The average gestation period is around 110 days; the female gives birth to a litter of between one and four cubs in a secluded den, which may be a thicket,
4026-905: Is a generalist hypercarnivore and is considered to be both an apex and keystone predator due to its wide prey spectrum. Its prey consists mainly of medium-sized to large ungulates , particularly blue wildebeest , plains zebra , African buffalo , gemsbok and giraffe . It also frequently takes common warthog despite it being much smaller. In India, chital and sambar deer are the most common wild prey, while livestock contributes significantly to lion kills outside protected areas. It usually avoids fully grown adult elephants , rhinoceros and hippopotamus and small prey like dik-dik , hyraxes , hares and monkeys . Unusual prey include porcupines and small reptiles. Lions kill other predators but seldom consume them. Young lions first display stalking behaviour at around three months of age, although they do not participate in hunting until they are almost
4148-493: Is a social species , forming groups called prides. A lion's pride consists of a few adult males, related females, and cubs. Groups of female lions usually hunt together, preying mostly on medium-sized and large ungulates . The lion is an apex and keystone predator ; although some lions scavenge when opportunities occur and have been known to hunt humans , lions typically do not actively seek out and prey on humans. The lion inhabits grasslands , savannahs , and shrublands . It
4270-553: Is a mixture of dry savannah forest and very dry, deciduous scrub forest . In Africa, the range of the lion originally spanned most of the central African rainforest zone and the Sahara desert. In the 1960s, it became extinct in North Africa, except in the southern part of Sudan. During the mid- Holocene , around 8,000-6,000 years ago, the range of lions expanded into Southeastern and Eastern Europe, partially re-occupying
4392-953: Is affected. Unlike in other felid species, female lions consistently interact with multiple males at once. Another hypothesis suggests that the mane also serves to protect the neck in fights, but this is disputed. During fights, including those involving maneless females and adolescents, the neck is not targeted as much as the face, back, and hindquarters. Injured lions also begin to lose their manes. Almost all male lions in Pendjari National Park are either maneless or have very short manes. Maneless lions have also been reported in Senegal, in Sudan 's Dinder National Park and in Tsavo East National Park , Kenya. Castrated lions often have little to no mane because
4514-457: Is also protected, and hunting is not allowed, but products and by-products from research may be sold. Wildlife refuges in Brazil have as their objective the protection of natural flora and fauna where conditions are assured for the existence and reproduction of species or communities of the local flora and the resident or migratory fauna. The refuges can consist of privately owned land, as long as
4636-881: Is defined in the relevant statutes used in those states and territories rather than by a single national statute. As of 2016, 1767 out of a total of 11044 protected areas listed within the Australian National Reserve System used the term "nature reserve" in their names. In Brazil, nature reserves are classified by the National System of Nature Conservation Units as ecological stations ( Portuguese : estações ecológicas ) or biological reserves ( Portuguese : reservas biológicas ). Their main objectives are preserving fauna and flora and other natural attributes, excluding direct human interference. Visits are allowed only with permission, and only for educational or scientific purposes. Changes to
4758-399: Is generally mutual and the recipient appears to express pleasure. The head and neck are the most common parts of the body licked; this behaviour may have arisen out of utility because lions cannot lick these areas themselves. Lions have an array of facial expressions and body postures that serve as visual gestures. A common facial expression is the "grimace face" or flehmen response , which
4880-404: Is home to 86 species including: Nile crocodile , hippopotamus , south African cheetah , spotted hyena , blue wildebeest , jackal , South African giraffe , Burchell's zebra , waterbuck , nyala , common eland , greater kudu , impala , duiker , suni , reedbuck , common warthog , bushpig , mongoose , baboon , monkeys , a variety of tortoises , terrapins , snakes and lizards . It
5002-480: Is no evidence for gene flow between the two lineages, indicating that they did not share the same geographic area. The Eurasian and American cave lions became extinct at the end of the last glacial period without mitochondrial descendants on other continents. The modern lion was probably widely distributed in Africa during the Middle Pleistocene and started to diverge in sub-Saharan Africa during
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5124-495: Is one of the world's top spots for viewing nyala . The park is a prime birding destination and is home to 340 bird species. The Hluhluwe River floodplain is one of the few areas in South Africa where yellow-throated longclaw , rosy-throated longclaw and Cape longclaw co-occur. Birdlife include black-crowned night heron , Wahlberg's eagle , Shelley's francolin , black-bellied bustard , Temminck's courser , Klaas's cuckoo , little bee-eater and crested barbet . The park has
5246-519: Is partly in a low-risk malaria area. This area is situated between the two Umfolozi Rivers where they divide into the Mfolozi emnyama ('Black Umfolozi') to the north and the Mfolozi emhlophe ('White Umfolozi') to the south. This area is to the south of the park and is generally hot in summer, and mild to cool in winter, although cold spells do occur. The topography in the Umfolozi section ranges from
5368-415: Is short and powerful; it attempts to catch prey with a fast rush and final leap, usually pulls it down by the rump, and kills with a clamping bite to the throat or muzzle . It can hold the prey's throat for up to 13 minutes, until the prey stops moving. It has a bite force from 1593.8 to 1768 Newtons at the canine tip and up 4167.6 Newtons at the carnassial notch. Lions typically consume prey at
5490-569: Is specifically defined in the Reserves Act to mean a reserve that prioritizes the protection of rare flora and fauna, to the extent that public access is by permit only. Some of these reserves include Ecological Islands , a comparatively new concept in wildlife preservation, pioneered in New Zealand to help rebuild the populations of nearly extinct birds, and other species that are heavily threatened by introduced predators. In Nicaragua ,
5612-590: Is the Tsavo lion pride that always has just one adult male. Prides act as fission–fusion societies , and members will split into subgroups that keep in contact with roars . Nomadic lions range widely and move around sporadically, either in pairs or alone. Pairs are more frequent among related males. A lion may switch lifestyles; nomads can become residents and vice versa. Interactions between prides and nomads tend to be hostile, although pride females in estrus allow nomadic males to approach them. Males spend years in
5734-624: Is the country with the most sites listed in the World Network of Biosphere Reserves . There are 4 biosphere reserves (two of them are dated 1927 and 1874) and 17 nature reserves in Ukraine, covering 160,000 ha (400,000 acres). Ukraine administers 40 national parks , 2632 habitat management areas, 3025 nature monuments, and 1430 other preservations. There are some differences between the regulations for England , Northern Ireland , Scotland and Wales , which are separately managed. At
5856-615: Is the most social of all wild felid species, living in groups of related individuals with their offspring. Such a group is called a " pride ". Groups of male lions are called "coalitions". Females form the stable social unit in a pride and do not tolerate outside females. The majority of females remain in their birth prides while all males and some females will disperse . The average pride consists of around 15 lions, including several adult females and up to four males and their cubs of both sexes. Large prides, consisting of up to 30 individuals, have been observed. The sole exception to this pattern
5978-480: Is typically brownish and tinged with yellow, rust, and black hairs. Mutations in the genes microphthalmia-associated transcription factor and tyrosinase are possibly responsible for the colour of manes. It starts growing when lions enter adolescence, when testosterone levels increase, and reach their full size at around four years old. Cool ambient temperatures in European and North American zoos may result in
6100-426: Is unable to consume all of the kill, it rests for a few hours before continuing to eat. On hot days, the pride retreats to shade with one or two males standing guard. Lions defend their kills from scavengers such as vultures and hyenas. Lions scavenge on carrion when the opportunity arises, scavenging animals dead from natural causes such as disease or those that were killed by other predators. Scavenging lions keep
6222-663: Is usually more diurnal than other wild cats, but when persecuted, it adapts to being active at night and at twilight . During the Neolithic period, the lion ranged throughout Africa and Eurasia, from Southeast Europe to India, but it has been reduced to fragmented populations in sub-Saharan Africa and one population in western India . It has been listed as Vulnerable on the IUCN Red List since 1996 because populations in African countries have declined by about 43% since
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#17327829413596344-647: Is yet to be officially classified), and seven specific types of "reserve", each of which prioritize various degrees of protection to different amenities such as scenery, recreation, flora and fauna, scientific value, or history. Land is often sub-categorised beneath its general classification, as defined in law between the Reserves Act of 1977 , the National Parks Act of 1980 , and the Conservation Act of 1987 . Under these classifications,
6466-470: The Department of Conservation administers more than 80,000 square kilometres (31,000 sq mi)—nearly 30% of the nation's total area—with at least some degree of protection. This land is composed of 14 National Parks, 30 Conservation Parks, and approximately 8,900 discrete areas of land in total. Although the most public land is strongly protected for natural preservation, the term nature reserve
6588-651: The Ministry of the Environment and Natural Resources (MARENA) is in charge of environmental protection and of the study, planning, and management of Nicaragua's natural resources. Nearly one-fifth of the territory is designated as protected areas like national parks, nature reserves (including the Bosawás Biosphere Reserve ), and biological reserves. Nicaragua has 78 protected areas that cover 22,422 km (8,657 sq mi), about 17.3% of
6710-512: The Narmada River in central India. Lions spend much of their time resting; they are inactive for about twenty hours per day. Although lions can be active at any time, their activity generally peaks after dusk with a period of socialising, grooming, and defecating. Intermittent bursts of activity continue until dawn, when hunting most often takes place. They spend an average of two hours a day walking and fifty minutes eating. The lion
6832-484: The Nile Basin during the early Holocene . In zoos, lions have been bred with tigers to create hybrids for the curiosity of visitors or for scientific purpose. The liger is bigger than a lion and a tiger, whereas most tigons are relatively small compared to their parents because of reciprocal gene effects. The leopon is a hybrid between a lion and leopard. The lion is a muscular, broad-chested cat with
6954-413: The big five game animals can be found. Thanks to conservation efforts, the park now has one of the largest populations of white rhinoceros in the world, although this population remains severely threatened by the poaching of hundreds of rhinos every year in the park. Throughout the park there are many signs of stone age settlements and iron smelting sites. The area is claimed to have been declared
7076-775: The Asiatic lion population. In 2017, the Cat Classification Task Force of the Cat Specialist Group revised lion taxonomy , and recognises two subspecies based on results of several phylogeographic studies on lion evolution , namely: However, there seems to be some degree of overlap between both groups in northern Central Africa. DNA analysis from a more recent study indicates that Central African lions are derived from both northern and southern lions, as they cluster with P. leo leo in mtDNA-based phylogenies whereas their genomic DNA indicates
7198-581: The British established the Yob Wildlife Reserve in northern Eritrea specifically to protect significant populations of Nubian ibex in the area. Denmark has three national parks and several nature reserves, some of them inside the national park areas. The largest single reserve is Hanstholm Nature Reserve, which covers 40 km (9,900 acres) and is part of Thy National Park . In Sweden , there are 30 national parks . The first of them
7320-545: The Hluhluwe–Imfolozi Park. Historically, tsetse flies carrying the nagana disease protected the area from colonial hunters. Later, as the Zululand area was settled by white farmers, wildlife in the reserves was blamed for the prevalence of the tsetse fly, and the reserves became experimental areas in the efforts to eradicate the fly. Farmers called for the slaughter of game and over 100,000 animals were killed in
7442-629: The Late Pleistocene. Lion populations in East and Southern Africa became separated from populations in West and North Africa when the equatorial rainforest expanded 183,500 to 81,800 years ago. They shared a common ancestor probably between 98,000 and 52,000 years ago. Due to the expansion of the Sahara between 83,100 and 26,600 years ago, lion populations in West and North Africa became separated. As
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#17327829413597564-566: The Ngorongoro Crater were afflicted by an outbreak of stable fly ( Stomoxys calcitrans ) in 1962, resulting in lions becoming emaciated and covered in bloody, bare patches. Lions sought unsuccessfully to evade the biting flies by climbing trees or crawling into hyena burrows; many died or migrated and the local population dropped from 70 to 15 individuals. A more recent outbreak in 2001 killed six lions. Captive lions have been infected with canine distemper virus (CDV) since at least
7686-571: The United States is Pelican Island National Wildlife Refuge and was established by Theodore Roosevelt in 1903 as part of his Square Deal campaign to improve the country. At the time, setting aside land for wildlife was not a constitutional right of the president. In 2006, a bi-partisan group of US House of Representatives members established the Congressional Wildlife Refuge Caucus to further support
7808-636: The adjacent Timbavati Private Game Reserve in eastern South Africa. They were removed from the wild in the 1970s, thus decreasing the white lion gene pool . Nevertheless, 17 births have been recorded in five prides between 2007 and 2015. White lions are selected for breeding in captivity. They have reportedly been bred in camps in South Africa for use as trophies to be killed during canned hunts . African lions live in scattered populations across sub-Saharan Africa. The lion prefers grassy plains and savannahs , scrub bordering rivers, and open woodlands with bushes. It rarely enters closed forests. On Mount Elgon ,
7930-424: The age of two. When a new male lion takes over a pride, adolescents both male and female may be evicted. Lions may live 12–17 years in the wild. Although adult lions have no natural predators, evidence suggests most die violently from attacks by humans or other lions. Lions often inflict serious injuries on members of other prides they encounter in territorial disputes or members of the home pride when fighting at
8052-642: The ancient world. According to stone inscriptions found in the vicinity, the king commanded the people not to harm animals or destroy trees within the area. In the United States, the U.S. Fish and Wildlife Service , managed by the Secretary of the Interior , is responsible for managing many of the federal nature reserves including the National Wildlife Refuge System . The National Wildlife Refuge System includes areas administered for
8174-544: The animal's expressive movements are highly developed. The most common peaceful, tactile gestures are head rubbing and social licking , which have been compared with the role of allogrooming among primates. Head rubbing, nuzzling the forehead, face and neck against another lion appears to be a form of greeting and is seen often after an animal has been apart from others or after a fight or confrontation. Males tend to rub other males, while cubs and females rub females. Social licking often occurs in tandem with head rubbing; it
8296-499: The attack. They take advantage of factors that reduce visibility; many kills take place near some form of cover or at night. One study in 2018 recorded a lion running at a top speed of 74.1 km/h (46.0 mph). The lion accelerates at the start of the chase by 9.5 m/s² , whereas zebras, wildebeest and Thomson's gazelle accelerate by 5 m/s², 5.6 m/s² and 4.5 m/s², respectively; acceleration appears to be more important than steady displacement speed in lion hunts. The lion's attack
8418-809: The best-known is the Kruger National Park , which was announced in 1898, and is the largest, at nearly 2,000,000 ha (20,000 km ). The Kruger Park and Table Mountain National Park are two of South Africa's most visited tourist attractions, along with the Addo Elephant National Park . South Africa also has 10 World Heritage Sites , including four natural sites and one mixed site. And it has provincial game reserves including Shamwari , Londolozi , Sanbona and Lalibela. The country currently has 20 national parks covering 3,700,000 ha (37,000 km ), about 3% of
8540-497: The corridor between the Hluhluwe and Imfolozi reserves was added in order to join the separate reserves into the current single park. The park is located in the province of KwaZulu-Natal on the east coast of South Africa . The park is closest to the town of Mtubatuba , Hluhluwe village and Hlabisa village. The geography of the area differs from the north, or Hluhluwe area, to the south, or Umfolozi area. Hluhluwe–Imfolozi Park
8662-908: The country's land area. In 1984, the two areas with the highest number of nature reserves were the South (15.2%) and Samaria (the Shomron, 13.5%). Under the Nature Conservation Law, places can be designated as 'wilderness areas', 'nature conservation areas' and 'prefectural nature conservation areas'. In 1995, when the Japanese Government published its information in English, there were 5 wilderness areas, 10 nature conservation areas and 516 prefectural nature conservation areas. There are seven nature reserves in Jordan . In 1966
8784-646: The country's total area. A few of them predate the October Revolution of 1917, but most have been created during the Soviet Union era. There are also natural protected areas where only certain species are protected, or only certain activities are prohibited; those are known as zakaznik (Russian: заказник ). Unofficial sanctuaries can also occur as a result of human accidents; the Chernobyl Exclusion Zone has in practice become
8906-411: The cubs varies—one male could patiently let the cubs play with his tail or his mane, while another may snarl and bat the cubs away. Pride lionesses often synchronise their reproductive cycles and communal rearing and suckling of the young, which suckle indiscriminately from any or all of the nursing females in the pride. The synchronisation of births is advantageous because the cubs grow to being roughly
9028-448: The early 1990s. Lion populations are untenable outside designated protected areas. Although the cause of the decline is not fully understood, habitat loss and conflicts with humans are the greatest causes for concern. One of the most widely recognised animal symbols in human culture, the lion has been extensively depicted in sculptures and paintings, on national flags, and in literature and films. Lions have been kept in menageries since
9150-424: The ecosystems in both types of reserve are allowed to restore and preserve the natural balance, biological diversity and natural ecological processes. Ecological stations are also allowed to change the environment within strictly defined limits (e.g. affecting no more than three percent of the area or 1,500 hectares (3,700 acres), whichever is less) for the purpose of scientific research. A wildlife reserve in Brazil
9272-709: The end of March 2004, there were 215 national nature reserves in England with a total area of 879 km (339 sq mi). The reserves are scattered through England, from Lindisfarne in Northumberland to The Lizard in Cornwall . Nearly every rural county has at least one. Many national nature reserves contain nationally important populations of rare flowers , ferns and mosses , butterflies and other insects , and nesting and wintering birds . Examples include unique alpine plants at Upper Teesdale and
9394-504: The field of snake's head fritillaries at North Meadow, Cricklade , Wiltshire . There are now over 1,050 local nature reserves in England. They range from windswept coastal headlands, ancient woodlands and flower-rich meadows to former inner-city railways, long-abandoned landfill sites and industrial areas now re-colonized by wildlife. In total, they cover almost 40,000 ha (99,000 acres)—a natural resource which makes an important contribution to England's biodiversity . A good example
9516-534: The group. Alternatively, there may be rewards associated with being a leader that fends off intruders; the rank of lionesses in the pride is reflected in these responses. The male or males associated with the pride must defend their relationship with the pride from outside males who may attempt to usurp them. Dominance hierarchies do not appear to exist among individuals of either sex in a pride. Asiatic lion prides differ in group composition. Male Asiatic lions are solitary or associate with up to three males, forming
9638-570: The hunting of large, slower-moving prey like buffalo; and even hunt them on their own. Moderately-sized hunting groups generally have higher success rates than lone females and larger groups. Lions are not particularly known for their stamina. For instance, a lioness's heart comprises only 0.57% of her body weight and a male's is about 0.45% of his body weight, whereas a hyena's heart comprises almost 1% of its body weight. Thus, lions run quickly only in short bursts at about 48–59 km/h (30–37 mph) and need to be close to their prey before starting
9760-510: The integration of systems, including drone technology, for early detection and rapid response of reaction units. On 6 March 2020 two of three suspected rhino poachers were killed in a shootout after they were detected by infrared cameras. In 2022, 244 rhinos were poached in the province, 228 of them in the park. In 2023, the reserve was the worst hit in South Africa; The Minister of Environment, Forestry and Fisheries , Barbara Creecy, said "We have to be concerned that almost every single day this year
9882-585: The largest total areas being in Bavaria with 1,416 km (547 sq mi) and Lower Saxony with 1,275 km (492 sq mi). In Hungary, there are 10 national parks , more than 15 nature reserves and more than 250 protected areas. Hortobágy National Park is the largest continuous natural grassland in Europe and the oldest national park in Hungary. It is situated on the eastern part of Hungary, on
10004-564: The laws no. 102/1983 and 4/1994 for protection of the Egyptian nature reserve. Egypt announced a plan from to build 40 nature reserves from 1997 to 2017, to help protect the natural resources and the culture and history of those areas. The largest nature reserve in Egypt is Gebel Elba (35,600 square kilometres (13,700 sq mi)) in the southeast, on the Red Sea coast. On 16 March 1959,
10126-506: The level of protection afforded by local laws. Normally it is more strictly protected than a nature park . Various jurisdictions may use other terminology, such as ecological protection area or private protected area in legislation and in official titles of the reserves. Cultural practices that roughly equate to the establishment and maintenance of reserved areas for animals date back to antiquity, with King Devanampiya Tissa of Sri Lanka establishing Mihintale wildlife sanctuary, one of
10248-722: The lion has been recorded up to an elevation of 3,600 m (11,800 ft) and close to the snow line on Mount Kenya . Savannahs with an annual rainfall of 300 to 1,500 mm (12 to 59 in) make up the majority of lion habitat in Africa, estimated at 3,390,821 km (1,309,203 sq mi) at most, but remnant populations are also present in tropical moist forests in West Africa and montane forests in East Africa. The Asiatic lion now survives only in and around Gir National Park in Gujarat, western India. Its habitat
10370-810: The lion population in Kenya's Masai Mara National Reserve declined, the spotted hyena population increased rapidly. Lions tend to dominate cheetahs and leopards, steal their kills and kill their cubs and even adults when given the chance. Cheetahs often lose their kills to lions or other predators. A study in the Serengeti ecosystem revealed that lions killed at least 17 of 125 cheetah cubs born between 1987 and 1990. Cheetahs avoid their competitors by hunting at different times and habitats. Leopards, by contrast, do not appear to be motivated by an avoidance of lions, as they use heavy vegetation regardless of whether lions are present in an area and both cats are active around
10492-551: The location of the hunt but sometimes drag large prey into cover. They tend to squabble over kills, particularly the males. Cubs suffer most when food is scarce but otherwise all pride members eat their fill, including old and crippled lions, which can live on leftovers. Large kills are shared more widely among pride members. An adult lioness requires an average of about 5 kg (11 lb) of meat per day while males require about 7 kg (15 lb). Lions gorge themselves and eat up to 30 kg (66 lb) in one session. If it
10614-516: The lowlands of the Umfolozi River beds to steep hilly country, which includes some wide and deep valleys. Habitats in this area are primarily grasslands, which extend into acacia savannah and woodlands. The Hluhluwe region has hilly topography where altitudes range from 80 to 540 metres (260 to 1,770 ft) above sea level. The high ridges support coastal scarp forests in a well-watered region with valley bushveld at lower levels. The north of
10736-491: The mid-1970s. CDV is spread by domestic dogs and other carnivores; a 1994 outbreak in Serengeti National Park resulted in many lions developing neurological symptoms such as seizures. During the outbreak, several lions died from pneumonia and encephalitis . Feline immunodeficiency virus and lentivirus also affect captive lions. When resting, lion socialisation occurs through a number of behaviours;
10858-549: The nation's landmass. Private nature reserves exist with land excluded from private land trusts and maintained at the sole cost of the proprietor. For example, O Parks, WildLife, and Recreation, Or El Ostional Private Wildlife Reserve, was established within the Mesoamerican Biological Corridor by retired FDNY firefighter Kevin Michael Shea, who purchased 46 acres (0.19 km ) of land for
10980-549: The needs of the National Wildlife Refuge System in Congress . Lion The lion ( Panthera leo ) is a large cat of the genus Panthera , native to Africa and India . It has a muscular, broad-chested body ; a short, rounded head; round ears; and a dark, hairy tuft at the tip of its tail. It is sexually dimorphic ; adult male lions are larger than females and have a prominent mane. It
11102-557: The objectives of the unit are compatible with the landowners' usage of the land and natural resources. Public visits are subject to the conditions and restrictions established by the management plan of the unit and are subject to authorisation by and regulations of the main administrative and scientific research body. The Niagara Escarpment and the St. Lawrence River in Ontario are among the 18 nature reserves recognized by UNESCO in Canada. As
11224-563: The organization that would later start Jordan's nature reserves, the Royal Society for the Conservation of Nature , was founded. RSCN's first efforts involved bringing back severely endangered species. In 1973, RSCN was given the right to issue hunting licenses, giving RSCN an upper hand in preventing extinction. The first step was the founding of Jordan's first nature reserve, Shaumari Wildlife Reserve , in 1975. The primary purpose
11346-463: The park is more rugged and mountainous with forests and grasslands and is known as the Hluhluwe area, while the Umfolozi area is found to the south near the Black and White Umfolozi rivers where there is open savanna . The park is home to Africa's big five game : African bush elephant , south-central black rhinoceros and southern white rhinoceros , African buffalo , lion and African leopard . It
11468-434: The phylogenetic relationship of the lion; it was thought to form a sister group with the jaguar that diverged 3.46 to 1.22 million years ago , but also with the leopard that diverged 3.1 to 1.95 million years ago to 4.32 to 0.02 million years ago . Hybridisation between lion and snow leopard ancestors possibly continued until about 2.1 million years ago. The lion-leopard clade
11590-585: The plain of the Alföld. It was established in 1972. There are alkaline grasslands interrupted by marshes. They have a sizable importance because there are the fishponds. One of the most spectacular sights of the park is the autumn migration of cranes. Some famous Hungarian animal species live in Hortobágy National Park, such as the grey cattle, racka long-wool sheep living only in Hungary, Hungarian horses and buffalo. Hortobágy National Park has been
11712-612: The previous males associated with a pride, the victors often kill any existing young cubs , perhaps because females do not become fertile and receptive until their cubs mature or die. Females often fiercely defend their cubs from a usurping male but are rarely successful unless a group of three or four mothers within a pride join forces against the male. Cubs also die from starvation and abandonment, and predation by leopards, hyenas and wild dogs. Male cubs are excluded from their maternal pride when they reach maturity at around two or three years of age, while some females may leave when they reach
11834-499: The protection and conservation of fish and wildlife that are threatened with extinction, as well as wildlife ranges, game ranges, wildlife management areas, and waterfowl production areas. The first North American wildlife refuge, Lake Merritt Wildlife sanctuary at Lake Merritt , was established by Samuel Merritt and enacted in California state law in 1870 as the first government owned refuge. The first federally owned refuge in
11956-531: The purpose of restoring the ecological system of a dry tropical forest , ravaged during the Nicaraguan Revolution . The park provides a private nature reserve, wildlife corridor and verified carbon credits . There are around 100 nature reserves ( Russian : заповедник , romanized : zapovednik ) in Russia, covering some 330,000 km (130,000 sq mi), or about 1.4% of
12078-589: The rainforest decreased and thus gave rise to more open habitats, lions moved from West to Central Africa. Lions from North Africa dispersed to southern Europe and Asia between 38,800 and 8,300 years ago. Extinction of lions in southern Europe, North Africa and the Middle East interrupted gene flow between lion populations in Asia and Africa. Genetic evidence revealed numerous mutations in lion samples from East and Southern Africa, which indicates that this group has
12200-457: The range of the now extinct cave lion . In Hungary , the modern lion was present from about 4,500 to 3,200 years Before Present . In Ukraine , the modern lion was present from about 6,400 to 2,000 years Before Present. In Greece, it was common, as reported by Herodotus in 480 BC; it was considered rare by 300 BC and extirpated by AD 100. In Asia the lion once ranged in regions where climatic conditions supported an abundance of prey. It
12322-488: The removal of the gonads inhibits testosterone production. Rarely, both wild and captive lionesses have manes. Increased testosterone may be the cause of maned lionesses reported in northern Botswana. The white lion is a rare morph with a genetic condition called leucism , which is caused by a double recessive allele . It is not albino; it has normal pigmentation in the eyes and skin. White lions have occasionally been encountered in and around Kruger National Park and
12444-541: The reserves between 1919 and 1950, although the rhino population was spared. The introduction of DDT spraying in 1945 virtually eliminated the tsetse fly from the reserves, although subsequent outbreaks have occurred. By the 1950s the white rhino population of the reserve had recovered to around 400, and the park's warden, Ian Player , established Operation Rhino in the 1950s and 60s, with the park's Rhino Capture Unit relocating hundreds of rhinos to establish populations in other reserves across their historic range. In 1989,
12566-429: The same area. The evolution of sociability in lions was likely driven both by high population density and the clumped resources of savannah habitats. The larger the pride, the more high-quality territory they can defend; "hotspots" being near river confluences , where the cats have better access to water, prey and shelter (via vegetation). The area occupied by a pride is called a "pride area" whereas that occupied by
12688-420: The same size and have an equal chance of survival, and sucklings are not dominated by older cubs. Weaning occurs after six or seven months. Male lions reach maturity at about three years of age and at four to five years are capable of challenging and displacing adult males associated with another pride. They begin to age and weaken at between 10 and 15 years of age at the latest. When one or more new males oust
12810-406: The same time of day. In addition, there is no evidence that lions affect leopard abundance. Leopards take refuge in trees, though lionesses occasionally attempt to climb up and retrieve their kills. Lions similarly dominate African wild dogs , taking their kills and dispatching pups or adult dogs. Population densities of wild dogs are low in areas where lions are more abundant. However, there are
12932-420: The same time. When first introduced to the rest of the pride, lion cubs lack confidence when confronted with adults other than their mother. They soon begin to immerse themselves in the pride life, however, playing among themselves or attempting to initiate play with the adults. Lionesses with cubs of their own are more likely to be tolerant of another lioness's cubs than lionesses without cubs. Male tolerance of
13054-422: The spur are unknown. The tuft is absent at birth and develops at around 5 + 1 ⁄ 2 months of age. It is readily identifiable at the age of seven months. Its skull is very similar to that of the tiger, although the frontal region is usually more depressed and flattened and has a slightly shorter postorbital region and broader nasal openings than those of the tiger. Due to the amount of skull variation in
13176-413: The tiger in size. The size and weight of adult lions vary across its range and habitats. Accounts of a few individuals that were larger than average exist from Africa and India. The male lion's mane is the most recognisable feature of the species. It may have evolved around 320,000–190,000 years ago. It grows downwards and backwards, covering most of the head, neck, shoulders, and chest. The mane
13298-585: The time of the Roman Empire and have been a key species sought for exhibition in zoological gardens across the world since the late 18th century. Cultural depictions of lions were prominent in Ancient Egypt , and depictions have occurred in virtually all ancient and medieval cultures in the lion's historic and current range. The English word lion is derived via Anglo-Norman liun from Latin leōnem (nominative: leō ), which in turn
13420-468: The total area of South Africa. The Prince Edward Islands , which are South African territories in the Southern Ocean , have been declared a special nature reserve. It is a highly protected area from which all human activity is excluded, except for conservation and scientific research. The area around Mihintale , Sri Lanka , was a sanctuary for wildlife, probably the first of its generation in
13542-407: The two species, usually only the structure of the lower jaw can be used as a reliable indicator of species. The skeletal muscles of the lion make up 58.8% of its body weight and represent the highest percentage of muscles among mammals. The lion has a high concentration of fast twitch muscle fibers, giving them quick bursts of speed but less stamina. Among felids, the lion is second only to
13664-441: The world's earliest wildlife sanctuaries , in the 3rd-century-BC Anuradhapura Ancient Kingdom. Early reservations often had a religious underpinning, such as the 'evil forest' areas of West Africa which were forbidden to humans, who were threatened with spiritual attack if they went there. Sacred areas taboo from human entry to fishing and hunting are known by many ancient cultures worldwide. The world's first modern nature reserve
13786-627: Was Yellowstone National Park in Wyoming, United States , followed by the Royal National Park near Sydney , Australia, and the Barguzin Nature Reserve of Imperial Russia , the first of zapovedniks set up by a federal government entirely for the scientific study of nature. There are several national and international organizations that oversee the numerous non-profit animal sanctuaries and refuges to provide
13908-400: Was a borrowing from Ancient Greek λέων léōn . The Hebrew word לָבִיא lavi may also be related. The generic name Panthera is traceable to the classical Latin word 'panthēra' and the ancient Greek word πάνθηρ 'panther'. Felis leo was the scientific name used by Carl Linnaeus in 1758, who described the lion in his work Systema Naturae . The genus name Panthera
14030-460: Was built at Hilltop in 1934, and is now the main camp in the northern (Hluhluwe) section of the park. The main camp in the southern (Umfolozi) section is Mpila. The reserve has a 300-kilometre (190 mi) road network. The park ecosystems are threatened by the proximity of coal mines, with ongoing applications for prospecting and mine expansions on the western and southern edges of the park. Nature reserve A nature reserve (also known as
14152-593: Was coined by Lorenz Oken in 1816. Between the mid-18th and mid-20th centuries, 26 lion specimens were described and proposed as subspecies, of which 11 were recognised as valid in 2005. They were distinguished mostly by the size and colour of their manes and skins. In the 19th and 20th centuries, several lion type specimens were described and proposed as subspecies , with about a dozen recognised as valid taxa until 2017. Between 2008 and 2016, IUCN Red List assessors used only two subspecific names: P. l. leo for African lion populations, and P. l. persica for
14274-505: Was described by David Attenborough as "one of the first people anywhere to recognise not only that the natural world was of great importance but that it needed protection as humanity made more and more demands on it". Drachenfels was protected as the first state-designated nature reserve in modern-day Germany; the site was bought by the Prussian State in 1836 to protect it from further quarrying. The first major nature reserve
14396-520: Was distributed in the Asian and African Palearctic since at least the early Pliocene . The earliest fossils recognisable as lions were found at Olduvai Gorge in Tanzania and are estimated to be up to 2 million years old. Estimates for the divergence time of the modern and cave lion lineages range from 529,000 to 392,000 years ago based on mutation rate per generation time of the modern lion. There
14518-634: Was established in 1821 by the naturalist and explorer Charles Waterton around his estate in Walton Hall, West Yorkshire . He spent £9000 on the construction of a three-mile long, 9 ft tall wall to enclose his park against poachers . He tried to encourage bird life by planting trees and hollowing out trunks for owls to nest in. Waterton invented artificial nest boxes to house starlings , western jackdaws and sand martins ; and unsuccessfully attempted to introduce little owls from Italy. Waterton allowed local people access to his reserve and
14640-829: Was established in 1909. There are almost 4,000 nature reserves in Sweden. They comprise about 85% of the surface that is protected by the Swedish Environmental Code. In Estonia, there are five national parks , more than 100 nature reserves, and around 130 landscape protection areas. The largest nature reserve in Estonia is Alam-Pedja Nature Reserve , which covers 342 km (85,000 acres). As of 2017, France counts 10 national parks , around 50 regional nature parks , and 8 marine parks . In 1995 Germany had 5,314 nature reserves ( German : Naturschutzgebiete ) covering 6,845 km (2,643 sq mi),
14762-951: Was present in the Caucasus until the 10th century. It lived in Palestine until the Middle Ages and in Southwest Asia until the late 19th century. By the late 19th century, it had been extirpated in most of Turkey. The last live lion in Iran was sighted in 1942, about 65 km (40 mi) northwest of Dezful , although the corpse of a lioness was found on the banks of the Karun river in Khuzestan province in 1944. It once ranged from Sind and Punjab in Pakistan to Bengal and
14884-615: Was to create means to breed endangered species, specifically: the Arabian oryx , gazelles , ostriches , and Persian onagers in their natural environment. By the end of 2009 there were 10 nature reserves ( Kyrgyz : корук , koruk ) in Kyrgyzstan covering 600,000 hectares (6,000 km ) or about three percent of the total area of the country. New Zealand has a variety of types of reserve, including national parks , various types of conservation areas (including stewardship land that
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