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102-671: The Iberian wolf ( Canis lupus signatus , or Canis lupus lupus , Spanish and Portuguese : Lobo ibérico ), is a subspecies of grey wolf . It inhabits the northwest of the Iberian Peninsula , which includes northwestern Spain and northern Portugal . It is home to 2,200-2,700 wolves which have been isolated from mixing with other wolf populations for over a century. They form the largest wolf population in Western Europe . Due to population controls and damage to livestock , Iberian wolves were, as of September 2021,
204-1054: A dagger (†); groups of uncertain placement are labelled with a question mark (?) and dashed lines (- - - - -). Jawless fishes (118 species: hagfish , lampreys ) [REDACTED] † Thelodonti , † Conodonta , † Anaspida [REDACTED] [REDACTED] [REDACTED] † Galeaspida [REDACTED] † Osteostraci [REDACTED] † Placodermi [REDACTED] † Acanthodii [REDACTED] (>1,100 species: sharks , rays , chimaeras ) [REDACTED] (2 species: coelacanths ) [REDACTED] Dipnoi (6 species: lungfish ) [REDACTED] Tetrapoda (>38,000 species, not considered fish: amphibians, reptiles, birds, mammals) [REDACTED] (14 species: bichirs , reedfish ) [REDACTED] (27 species: sturgeons , paddlefish ) [REDACTED] Ginglymodi (7 species: gars , alligator gars ) [REDACTED] Halecomorphi (2 species: bowfin , eyetail bowfin ) [REDACTED] (>32,000 species) [REDACTED] Fishes (without tetrapods) are
306-579: A paraphyletic group and for this reason, the class Pisces seen in older reference works is no longer used in formal classifications. Traditional classification divides fish into three extant classes (Agnatha, Chondrichthyes, and Osteichthyes), and with extinct forms sometimes classified within those groups, sometimes as their own classes. Fish account for more than half of vertebrate species. As of 2016, there are over 32,000 described species of bony fish, over 1,100 species of cartilaginous fish, and over 100 hagfish and lampreys. A third of these fall within
408-613: A Spanish-based creole language called Chavacano developed in the southern Philippines. However, it is not mutually intelligible with Spanish. The number of Chavacano-speakers was estimated at 1.2 million in 1996. The local languages of the Philippines also retain significant Spanish influence, with many words derived from Mexican Spanish , owing to the administration of the islands by Spain through New Spain until 1821, until direct governance from Madrid afterwards to 1898. Fish A fish ( pl. : fish or fishes )
510-603: A considerable number of words from Arabic , as well as a minor influence from the Germanic Gothic language through the period of Visigoth rule in Iberia. In addition, many more words were borrowed from Latin through the influence of written language and the liturgical language of the Church. The loanwords were taken from both Classical Latin and Renaissance Latin , the form of Latin in use at that time. According to
612-522: A dramatic change in the pronunciation of its sibilant consonants , known in Spanish as the reajuste de las sibilantes , which resulted in the distinctive velar [x] pronunciation of the letter ⟨j⟩ and—in a large part of Spain—the characteristic interdental [θ] ("th-sound") for the letter ⟨z⟩ (and for ⟨c⟩ before ⟨e⟩ or ⟨i⟩ ). See History of Spanish (Modern development of
714-482: A first language by Spaniards and educated Filipinos ( Ilustrados ). Despite a public education system set up by the colonial government, by the end of Spanish rule in 1898, only about 10% of the population had knowledge of Spanish, mostly those of Spanish descent or elite standing. Spanish continued to be official and used in Philippine literature and press during the early years of American administration after
816-478: A long history in the territory of the current-day United States dating back to the 16th century. In the wake of the 1848 Guadalupe Hidalgo Treaty , hundreds of thousands of Spanish speakers became a minoritized community in the United States. The 20th century saw further massive growth of Spanish speakers in areas where they had been hitherto scarce. According to the 2020 census, over 60 million people of
918-436: A more spherical lens . Their retinas generally have both rods and cones (for scotopic and photopic vision ); many species have colour vision , often with three types of cone. Teleosts can see polarized light ; some such as cyprinids have a fourth type of cone that detects ultraviolet . Amongst jawless fish , the lamprey has well-developed eyes, while the hagfish has only primitive eyespots. Hearing too
1020-508: A primarily Hassaniya Arabic -speaking territory, Spanish was officially spoken as the language of the colonial administration during the late 19th and 20th centuries. Today, Spanish is present in the partially-recognized Sahrawi Arab Democratic Republic as its secondary official language, and in the Sahrawi refugee camps in Tindouf ( Algeria ), where the Spanish language is still taught as
1122-419: A protective bony cover or operculum . They are able to oxygenate their gills using muscles in the head. Some 400 species of fish in 50 families can breathe air, enabling them to live in oxygen-poor water or to emerge on to land. The ability of fish to do this is potentially limited by their single-loop circulation, as oxygenated blood from their air-breathing organ will mix with deoxygenated blood returning to
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#17327937679881224-617: A second language features characteristics involving the variability of the vowel system. While far from its heyday during the Spanish protectorate in Morocco , the Spanish language has some presence in northern Morocco , stemming for example from the availability of certain Spanish-language media. According to a 2012 survey by Morocco's Royal Institute for Strategic Studies (IRES), penetration of Spanish in Morocco reaches 4.6% of
1326-531: A second language, largely by Cuban educators. The number of Spanish speakers is unknown. Spanish is also an official language of the African Union . Spanish was an official language of the Philippines from the beginning of Spanish administration in 1565 to a constitutional change in 1973. During Spanish colonization , it was the language of government, trade, and education, and was spoken as
1428-441: A sense of touch and of hearing . Blind cave fish navigate almost entirely through the sensations from their lateral line system. Some fish, such as catfish and sharks, have the ampullae of Lorenzini , electroreceptors that detect weak electric currents on the order of millivolt. Vision is an important sensory system in fish. Fish eyes are similar to those of terrestrial vertebrates like birds and mammals, but have
1530-663: A study of mitochondrial DNA sequences of both modern and ancient wolves indicated that in Europe, the two most genetically distinct haplotypes form the Italian wolf , and separately, the Iberian wolf. The National Center for Biotechnology Information / Genbank lists the Iberian wolf under Canis lupus signatus . In 2020, a genomic study of Eurasian wolves found that the populations of the Dinaric Alps - Balkan Mountains region,
1632-485: A tail fin, jaws, skin covered with scales , and lays eggs. Each criterion has exceptions, creating a wide diversity in body shape and way of life. For example, some fast-swimming fish are warm-blooded, while some slow-swimming fish have abandoned streamlining in favour of other body shapes. Fish species are roughly divided equally between freshwater and marine (oceanic) ecosystems; there are some 15,200 freshwater species and around 14,800 marine species. Coral reefs in
1734-589: A true "land fish" as this worm-like catfish strictly lives among waterlogged leaf litter . Cavefish of multiple families live in underground lakes , underground rivers or aquifers . Like other animals, fish suffer from parasitism . Some species use cleaner fish to remove external parasites. The best known of these are the bluestreak cleaner wrasses of coral reefs in the Indian and Pacific oceans. These small fish maintain cleaning stations where other fish congregate and perform specific movements to attract
1836-422: A typical fish is adapted for efficient swimming by alternately contracting paired sets of muscles on either side of the backbone. These contractions form S-shaped curves that move down the body. As each curve reaches the tail fin, force is applied to the water, moving the fish forward. The other fins act as control surfaces like an aircraft's flaps, enabling the fish to steer in any direction. Since body tissue
1938-411: A white stroke on the cheek that joins the white throat and with a reddish snout; and C. l. deitanus was much smaller and with more colouring than the wolves to the north. C. l. signatus is described by Cabrera as being 140–180 cm in head and body length, and 70–80 cm height at the shoulders. The Iberian wolf differs from the more common Eurasian wolf with its slighter frame, white marks on
2040-416: Is cognate to the root word of satisfacer ("to satisfy"), and hecho ("made") is similarly cognate to the root word of satisfecho ("satisfied"). Compare the examples in the following table: Some consonant clusters of Latin also produced characteristically different results in these languages, as shown in the examples in the following table: In the 15th and 16th centuries, Spanish underwent
2142-436: Is Latin in origin, including Latin borrowings from Ancient Greek. Alongside English and French , it is also one of the most taught foreign languages throughout the world. Spanish is well represented in the humanities and social sciences . Spanish is also the third most used language on the internet by number of users after English and Chinese and the second most used language by number of websites after English. Spanish
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#17327937679882244-674: Is a global language with about 500 million native speakers, mainly in the Americas and Spain , and about 600 million when including second language speakers. Spanish is the official language of 20 countries , as well as one of the six official languages of the United Nations . Spanish is the world's second-most spoken native language after Mandarin Chinese ; the world's fourth-most spoken language overall after English , Mandarin Chinese, and Hindustani ( Hindi - Urdu ); and
2346-469: Is a system of rugged mountain ranges in the south of the Iberian Peninsula where a very small wolf population has lived in isolation for half a century. A breeding pair of wolves was last sighted in 2013, however an extensive survey in 2016 could not find a breeding population. The DNA of a road-killed wolf from 2003 indicated high levels of inbreeding and 30% of the genome being that of dog. It
2448-423: Is an aquatic , anamniotic , gill -bearing vertebrate animal with swimming fins and a hard skull , but lacking limbs with digits . Fish can be grouped into the more basal jawless fish and the more common jawed fish , the latter including all living cartilaginous and bony fish , as well as the extinct placoderms and acanthodians . Most fish are cold-blooded , their body temperature varying with
2550-465: Is an important sensory system in fish. Fish sense sound using their lateral lines and otoliths in their ears, inside their heads. Some can detect sound through the swim bladder. Some fish, including salmon, are capable of magnetoreception ; when the axis of a magnetic field is changed around a circular tank of young fish, they reorient themselves in line with the field. The mechanism of fish magnetoreception remains unknown; experiments in birds imply
2652-662: Is classified as Canis lupus lupus by Mammal Species of the World . Some authors claim that the south-eastern Spanish wolf, last sighted in Murcia in the 1930s, was a different subspecies called Canis lupus deitanus . It was even smaller and more reddish in color, without dark spots. Both subspecies were nominated by the Spanish-born zoologist Ángel Cabrera in 1907. The Iberian wolf's skull morphometrics , mtDNA , and microsatellites differ from other European wolves. In 2016,
2754-431: Is denser than water, fish must compensate for the difference or they will sink. Many bony fish have an internal organ called a swim bladder that allows them to adjust their buoyancy by increasing or decreasing the amount of gas it contains. The scales of fish provide protection from predators at the cost of adding stiffness and weight. Fish scales are often highly reflective; this silvering provides camouflage in
2856-399: Is estimated that about 486 million people speak Spanish as a native language , making it the second most spoken language by number of native speakers . An additional 75 million speak Spanish as a second or foreign language , making it the fourth most spoken language in the world overall after English, Mandarin Chinese, and Hindi with a total number of 538 million speakers. Spanish is also
2958-761: Is heavily influenced by Venezuelan Spanish. In addition to sharing most of its borders with Spanish-speaking countries, the creation of Mercosur in the early 1990s induced a favorable situation for the promotion of Spanish language teaching in Brazil . In 2005, the National Congress of Brazil approved a bill, signed into law by the President , making it mandatory for schools to offer Spanish as an alternative foreign language course in both public and private secondary schools in Brazil. In September 2016 this law
3060-785: Is often due to varying biodiversity throughout its range. Wolf populations strongly declined across Europe during the 18th and 19th centuries largely due to human persecution, and by the end of the Second World War they had been eradicated from all of Central Europe and almost all of Northern Europe . Their population decline continued until the 1960s, with isolated populations surviving in Italy , Spain, Portugal, Greece , and Finland . Wolf populations have commenced recovering naturally since then. The Iberian wolf consists of over 2,000 individuals in over 350 packs distributed across 140,000 km. Their densities in some regions are among
3162-844: Is spoken by very small communities in Angola due to Cuban influence from the Cold War and in South Sudan among South Sudanese natives that relocated to Cuba during the Sudanese wars and returned for their country's independence. Spanish is also spoken in the integral territories of Spain in Africa, namely the cities of Ceuta and Melilla and the Canary Islands , located in the Atlantic Ocean some 100 km (62 mi) off
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3264-465: Is the biggest part of the brain; it is small in hagfish and lampreys , but very large in mormyrids , processing their electrical sense . The brain stem or myelencephalon controls some muscles and body organs, and governs respiration and osmoregulation . The lateral line system is a network of sensors in the skin which detects gentle currents and vibrations, and senses the motion of nearby fish, whether predators or prey. This can be considered both
3366-813: Is the de facto national language of the country, Spanish is often used in public services and notices at the federal and state levels. Spanish is also used in administration in the state of New Mexico . The language has a strong influence in major metropolitan areas such as those of Los Angeles , Miami , San Antonio , New York , San Francisco , Dallas , Tucson and Phoenix of the Arizona Sun Corridor , as well as more recently, Chicago , Las Vegas , Boston , Denver , Houston , Indianapolis , Philadelphia , Cleveland , Salt Lake City , Atlanta , Nashville , Orlando , Tampa , Raleigh and Baltimore-Washington, D.C. due to 20th- and 21st-century immigration. Although Spanish has no official recognition in
3468-488: Is the official Spanish language of the State. ... The other Spanish languages shall also be official in their respective Autonomous Communities... The Royal Spanish Academy ( Real Academia Española ), on the other hand, currently uses the term español in its publications. However, from 1713 to 1923, it called the language castellano . The Diccionario panhispánico de dudas (a language guide published by
3570-550: Is thought that as the population of these wolves declined, the inability to find a mate led to inbreeding and hybridization with dogs. The wolves in this region may now be extinct. Spanish language Spanish ( español ) or Castilian ( castellano ) is a Romance language of the Indo-European language family that evolved from the Vulgar Latin spoken on the Iberian Peninsula of Europe . Today, it
3672-542: Is used as an official language by many international organizations , including the United Nations , European Union , Organization of American States , Union of South American Nations , Community of Latin American and Caribbean States , African Union , among others. In Spain and some other parts of the Spanish-speaking world, Spanish is called not only español but also castellano (Castilian),
3774-578: Is usually assumed to be derived from castillo ('castle'). In the Middle Ages , the language spoken in Castile was generically referred to as Romance and later also as Lengua vulgar . Later in the period, it gained geographical specification as Romance castellano ( romanz castellano , romanz de Castiella ), lenguaje de Castiella , and ultimately simply as castellano (noun). Different etymologies have been suggested for
3876-565: The Gnathostomata or (for bony fish) Osteichthyes , also contains the clade of tetrapods (four-limbed vertebrates, mostly terrestrial), which are usually not considered fish. Some tetrapods, such as cetaceans and ichthyosaurs , have secondarily acquired a fish-like body shape through convergent evolution . Fishes of the World comments that "it is increasingly widely accepted that tetrapods, including ourselves, are simply modified bony fishes, and so we are comfortable with using
3978-723: The Indo-Pacific constitute the center of diversity for marine fishes, whereas continental freshwater fishes are most diverse in large river basins of tropical rainforests , especially the Amazon , Congo , and Mekong basins. More than 5,600 fish species inhabit Neotropical freshwaters alone, such that Neotropical fishes represent about 10% of all vertebrate species on the Earth. Fish are abundant in most bodies of water. They can be found in nearly all aquatic environments, from high mountain streams (e.g., char and gudgeon ) to
4080-681: The Silurian , with giant armoured placoderms such as Dunkleosteus . Jawed fish, too, appeared during the Silurian: the cartilaginous Chondrichthyes and the bony Osteichthyes . During the Devonian , fish diversity greatly increased, including among the placoderms, lobe-finned fishes, and early sharks, earning the Devonian the epithet "the age of fishes". Fishes are a paraphyletic group, since any clade containing all fish, such as
4182-568: The Spanish as a First Foreign Language (SAFFL) initiative in March 2005. Spanish has historically had a significant presence on the Dutch Caribbean islands of Aruba , Bonaire and Curaçao ( ABC Islands ) throughout the centuries and in present times. The majority of the populations of each island (especially Aruba) speaking Spanish at varying although often high degrees of fluency. The local language Papiamentu (Papiamento on Aruba)
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4284-508: The Spanish–American War but was eventually replaced by English as the primary language of administration and education by the 1920s. Nevertheless, despite a significant decrease in influence and speakers, Spanish remained an official language of the Philippines upon independence in 1946, alongside English and Filipino , a standardized version of Tagalog . Spanish was briefly removed from official status in 1973 but reimplemented under
4386-477: The abyssal and even hadal depths of the deepest oceans (e.g., cusk-eels and snailfish ), although none have been found in the deepest 25% of the ocean. The deepest living fish in the ocean so far found is a cusk-eel, Abyssobrotula galatheae , recorded at the bottom of the Puerto Rico Trench at 8,370 m (27,460 ft). In terms of temperature, Jonah's icefish live in cold waters of
4488-425: The end-Devonian extinction wiped out the apex placoderms. Bony fish are further divided into the lobe-finned and ray-finned fish . About 96% of all living fish species today are teleosts , a crown group of ray-finned fish that can protrude their jaws . The tetrapods , a mostly terrestrial clade of vertebrates that have dominated the top trophic levels in both aquatic and terrestrial ecosystems since
4590-400: The intertidal zone , are facultative air breathers, able to breathe air when out of water, as may occur daily at low tide , and to use their gills when in water. Some coastal fish like rockskippers and mudskippers choose to leave the water to feed in habitats temporarily exposed to the air. Some catfish absorb air through their digestive tracts. The digestive system consists of a tube,
4692-405: The ostracoderms , had heavy bony plates that served as protective exoskeletons against invertebrate predators . The first fish with jaws , the placoderms, appeared in the Silurian and greatly diversified during the Devonian , the "Age of Fishes". Bony fish, distinguished by the presence of swim bladders and later ossified endoskeletons , emerged as the dominant group of fish after
4794-425: The stout infantfish . Swimming performance varies from fish such as tuna, salmon , and jacks that can cover 10–20 body-lengths per second to species such as eels and rays that swim no more than 0.5 body-lengths per second. A typical fish is cold-blooded , has a streamlined body for rapid swimming, extracts oxygen from water using gills, has two sets of paired fins, one or two dorsal fins, an anal fin and
4896-423: The 1930s, Iberian wolves were relatively spread throughout Portugal, but destruction of habitat, loss of wild ungulates and the persecution by humans made it lose most of its territory (from around 44,100 km in 1900–1930, to only 16,300 km in 2002–2003) though populations have grown to about 29,000 km since. Some Spanish naturalists and conservationists such as Félix Rodríguez de la Fuente called for
4998-404: The 2021–2022 school year alone. The local business process outsourcing industry has also helped boost the language's economic prospects. Today, while the actual number of proficient Spanish speakers is around 400,000, or under 0.5% of the population, a new generation of Spanish speakers in the Philippines has likewise emerged, though speaker estimates vary widely. Aside from standard Spanish,
5100-595: The 20th century, Spanish was introduced to Equatorial Guinea and the Western Sahara , and to areas of the United States that had not been part of the Spanish Empire, such as Spanish Harlem in New York City . For details on borrowed words and other external influences upon Spanish, see Influences on the Spanish language . Spanish is the primary language in 20 countries worldwide. As of 2023, it
5202-564: The 9th century, and the first systematic written use of the language happened in Toledo , a prominent city of the Kingdom of Castile , in the 13th century. Spanish colonialism in the early modern period spurred the introduction of the language to overseas locations, most notably to the Americas. As a Romance language, Spanish is a descendant of Latin. Around 75% of modern Spanish vocabulary
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#17327937679885304-654: The Equatoguinean education system and is the primary language used in government and business. Whereas it is not the mother tongue of virtually any of its speakers, the vast majority of the population is proficient in Spanish. The Instituto Cervantes estimates that 87.7% of the population is fluent in Spanish. The proportion of proficient Spanish speakers in Equatorial Guinea exceeds the proportion of proficient speakers in other West and Central African nations of their respective colonial languages. Spanish
5406-637: The Iberian Peninsula. These languages included Proto-Basque , Iberian , Lusitanian , Celtiberian and Gallaecian . The first documents to show traces of what is today regarded as the precursor of modern Spanish are from the 9th century. Throughout the Middle Ages and into the modern era , the most important influences on the Spanish lexicon came from neighboring Romance languages — Mozarabic ( Andalusi Romance ), Navarro-Aragonese , Leonese , Catalan/Valencian , Portuguese , Galician , Occitan , and later, French and Italian . Spanish also borrowed
5508-462: The Iberian peninsula, and Italy diverged from each other 10,500 years ago followed by negligible gene flow between them. Their long-term isolation may explain the morphological and genetic differences between them. Cabrera identified two types of wolves living in Spain. In comparison, C. l. lupus had a pale, undefined stroke on the cheek that contrasts with the white on the throat; C. l. signatus had
5610-515: The Iberian wolf primarily (82%) fed on wild animals, with domesticated animals only making up a small (18%) portion of the diet. Wild boar (44%) and roe deer (35%) were the most important prey items, with ibex (3%) and red deer (0.3%) barely being targeted. The structure and availability of prey populations, as well as the size of its prey, often affects the diet of the Iberian wolf as well, as it has been observed to occasionally consume smaller prey such as rodents and birds. The consumption of unusual prey
5712-624: The Late Paleozoic , evolved from lobe-finned fish during the Carboniferous , developing air-breathing lungs homologous to swim bladders. Despite the cladistic lineage, tetrapods are usually not considered to be fish, making "fish" a paraphyletic group. Fish have been an important natural resource for humans since prehistoric times, especially as food . Commercial and subsistence fishers harvest fish in wild fisheries or farm them in ponds or in breeding cages in
5814-540: The Old Spanish sibilants) for details. The Gramática de la lengua castellana , written in Salamanca in 1492 by Elio Antonio de Nebrija , was the first grammar written for a modern European language. According to a popular anecdote, when Nebrija presented it to Queen Isabella I , she asked him what was the use of such a work, and he answered that language is the instrument of empire. In his introduction to
5916-589: The Royal Spanish Academy) states that, although the Royal Spanish Academy prefers to use the term español in its publications when referring to the Spanish language, both terms— español and castellano —are regarded as synonymous and equally valid. The term castellano is related to Castile ( Castilla or archaically Castiella ), the kingdom where the language was originally spoken. The name Castile , in turn,
6018-601: The Southern Ocean, including under the Filchner–Ronne Ice Shelf at a latitude of 79°S, while desert pupfish live in desert springs, streams, and marshes, sometimes highly saline, with water temperatures as high as 36 C. A few fish live mostly on land or lay their eggs on land near water. Mudskippers feed and interact with one another on mudflats and go underwater to hide in their burrows. A single undescribed species of Phreatobius has been called
6120-502: The U.S. population were of Hispanic or Hispanic American by origin. In turn, 41.8 million people in the United States aged five or older speak Spanish at home, or about 13% of the population. Spanish predominates in the unincorporated territory of Puerto Rico , where it is also an official language along with English. Spanish is by far the most common second language in the country, with over 50 million total speakers if non-native or second-language speakers are included. While English
6222-457: The administration of Ferdinand Marcos two months later. It remained an official language until the ratification of the present constitution in 1987, in which it was re-designated as a voluntary and optional auxiliary language. Additionally, the constitution, in its Article XIV, stipulates that the Government shall provide the people of the Philippines with a Spanish-language translation of
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#17327937679886324-707: The animal's protection to the European Parliament Committee on Petitions . Over a few weeks in September and October 2013, about 30 puppies and young wolves were killed by hunters in Asturias, León, and Cantabria. Although hunting is banned in Portugal, about 45% of wolf deaths are due to human activities, including illegal hunting. Calling the rate unsustainable, 10 organisations signed a statement requesting stronger protection. Sierra Morena
6426-556: The attention of the cleaners. Cleaning behaviors have been observed in a number of fish groups, including an interesting case between two cichlids of the same genus, Etroplus maculatus , the cleaner, and the much larger E. suratensis . Fish occupy many trophic levels in freshwater and marine food webs . Fish at the higher levels are predatory , and a substantial part of their prey consists of other fish. In addition, mammals such as dolphins and seals feed on fish, alongside birds such as gannets and cormorants . The body of
6528-550: The body, and produce a concentrated urine. The reverse happens in freshwater fish : they tend to gain water osmotically, and produce a dilute urine. Some fish have kidneys able to operate in both freshwater and saltwater. Fish have small brains relative to body size compared with other vertebrates, typically one-fifteenth the brain mass of a similarly sized bird or mammal. However, some fish have relatively large brains, notably mormyrids and sharks , which have brains about as large for their body weight as birds and marsupials . At
6630-684: The classic hispanus or hispanicus took the suffix -one from Vulgar Latin , as happened with other words such as bretón (Breton) or sajón (Saxon). Like the other Romance languages , the Spanish language evolved from Vulgar Latin , which was brought to the Iberian Peninsula by the Romans during the Second Punic War , beginning in 210 BC. Several pre-Roman languages (also called Paleohispanic languages )—some distantly related to Latin as Indo-European languages , and some that are not related at all—were previously spoken in
6732-500: The consonant written b (a bilabial with plosive and fricative allophones). In modern Spanish, there is no difference between the pronunciation of orthographic b and v . Typical of Spanish (as also of neighboring Gascon extending as far north as the Gironde estuary , and found in a small area of Calabria ), attributed by some scholars to a Basque substratum was the mutation of Latin initial f into h- whenever it
6834-404: The country's constitution. In recent years changing attitudes among non-Spanish speaking Filipinos have helped spur a revival of the language, and starting in 2009 Spanish was reintroduced as part of the basic education curriculum in a number of public high schools, becoming the largest foreign language program offered by the public school system, with over 7,000 students studying the language in
6936-525: The east. Height is 70–90 cm and body length 110–140 cm. Females weigh 25–35 kg while Males weigh 35–55 kg. The Iberian wolf lives in small packs. It is considered to be beneficial because it keeps the population of wild boars stable, thus allowing some respite to the endangered capercaillie populations which suffer greatly from boar predation. It also eats rabbits , roe deer , red deer , ibexes and even small carnivores and fish , depending on location. In Guadarrama National Park ,
7038-525: The end of the hunting and the protection of the animal. Today, the hunting of wolves is banned in Portugal, and in Spain since 2021. The 2003 census estimated the total Iberian population to be 2,000 wolves. Wolves have been reported as returning to Navarre and the Basque Country and to the provinces of Extremadura , Madrid , and Guadalajara . A male wolf was found recently in Catalonia , where
7140-570: The exact root is unknown; some authorities reconstruct a Proto-Indo-European root * peysk- , attested only in Italic , Celtic , and Germanic . About 530 million years ago during the Cambrian explosion , fishlike animals with a notochord and eyes at the front of the body, such as Haikouichthys , appear in the fossil record . During the late Cambrian , other jawless forms such as conodonts appear. Jawed vertebrates appear in
7242-683: The following table: Spanish is marked by palatalization of the Latin double consonants ( geminates ) nn and ll (thus Latin annum > Spanish año , and Latin anellum > Spanish anillo ). The consonant written u or v in Latin and pronounced [w] in Classical Latin had probably " fortified " to a bilabial fricative /β/ in Vulgar Latin. In early Spanish (but not in Catalan or Portuguese) it merged with
7344-466: The former British colony of Belize (known until 1973 as British Honduras ) where English is the sole official language, according to the 2022 census, 54% of the total population are able to speak the language. Due to its proximity to Spanish-speaking countries and small existing native Spanish speaking minority, Trinidad and Tobago has implemented Spanish language teaching into its education system. The Trinidadian and Tobagonian government launched
7446-405: The front of the brain are the olfactory lobes , a pair of structures that receive and process signals from the nostrils via the two olfactory nerves . Fish that hunt primarily by smell, such as hagfish and sharks, have very large olfactory lobes. Behind these is the telencephalon , which in fish deals mostly with olfaction. Together these structures form the forebrain. Connecting the forebrain to
7548-412: The gills flows in the opposite direction to the water, resulting in efficient countercurrent exchange . The gills push the oxygen-poor water out through openings in the sides of the pharynx. Cartilaginous fish have multiple gill openings: sharks usually have five, sometimes six or seven pairs; they often have to swim to oxygenate their gills. Bony fish have a single gill opening on each side, hidden beneath
7650-532: The gills. Oxygen-rich blood then flows without further pumping, unlike in mammals, to the body tissues. Finally, oxygen-depleted blood returns to the heart. Fish exchange gases using gills on either side of the pharynx . Gills consist of comblike structures called filaments. Each filament contains a capillary network that provides a large surface area for exchanging oxygen and carbon dioxide . Fish exchange gases by pulling oxygen-rich water through their mouths and pumping it over their gills. Capillary blood in
7752-604: The grammar, dated 18 August 1492, Nebrija wrote that "... language was always the companion of empire." From the 16th century onwards, the language was taken to the Spanish-discovered America and the Spanish East Indies via Spanish colonization of America . Miguel de Cervantes , author of Don Quixote , is such a well-known reference in the world that Spanish is often called la lengua de Cervantes ("the language of Cervantes"). In
7854-447: The gut, leading from the mouth to the anus. The mouth of most fishes contains teeth to grip prey, bite off or scrape plant material, or crush the food. An esophagus carries food to the stomach where it may be stored and partially digested. A sphincter, the pylorus, releases food to the intestine at intervals. Many fish have finger-shaped pouches, pyloric caeca , around the pylorus, of doubtful function. The pancreas secretes enzymes into
7956-457: The heart from the rest of the body. Lungfish, bichirs, ropefish, bowfins, snakefish, and the African knifefish have evolved to reduce such mixing, and to reduce oxygen loss from the gills to oxygen-poor water. Bichirs and lungfish have tetrapod-like paired lungs, requiring them to surface to gulp air, and making them obligate air breathers. Many other fish, including inhabitants of rock pools and
8058-599: The highest in Europe, with up to 7 wolves/100 km. In 2018, a study identified that this wolf could be categorised into 11 genetic groups. Four groups were identified in the Portuguese regions of Alto Minho , Alto Trás-os-Montes , and one group located south of the Douro river . Seven groups were identified in the Spanish regions of western Galicia , eastern Galicia, western Asturias , central Asturias, eastern Asturias, south-eastern Asturias, and Castilla y León . Until
8160-455: The intestine to digest the food; other enzymes are secreted directly by the intestine itself. The liver produces bile which helps to break up fat into an emulsion which can be absorbed in the intestine. Most fish release their nitrogenous wastes as ammonia . This may be excreted through the gills or filtered by the kidneys . Salt is excreted by the rectal gland. Saltwater fish tend to lose water by osmosis ; their kidneys return water to
8262-632: The language from the Kingdom of Castile , contrasting it with other languages spoken in Spain such as Galician , Basque , Asturian , Catalan/Valencian , Aragonese , Occitan and other minor languages. The Spanish Constitution of 1978 uses the term castellano to define the official language of the whole of Spain, in contrast to las demás lenguas españolas (lit. "the other Spanish languages "). Article III reads as follows: El castellano es la lengua española oficial del Estado. ... Las demás lenguas españolas serán también oficiales en las respectivas Comunidades Autónomas... Castilian
8364-549: The language's hegemony in an intensely centralising Spanish state were established from the 18th century onward. Other European territories in which it is also widely spoken include Gibraltar and Andorra . Spanish is also spoken by immigrant communities in other European countries, such as the United Kingdom , France , Italy , and Germany . Spanish is an official language of the European Union . Today,
8466-502: The last native wolf was killed in 1929. However, this animal was not a member of the Iberian subspecies, but an Italian wolf ( C. l. italicus ) migrating from France . As of 2013, an estimated 300 individuals remain in Portugal. In October 2013, Ecologists in Action called for an urgent review of the Iberian wolf census, which may overestimate their numbers. Later that month, wolf association Lobo Marley sent 198,000 signatures calling for
8568-694: The majority of the Spanish speakers live in Hispanic America . Nationally, Spanish is the official language—either de facto or de jure —of Argentina , Bolivia (co-official with 36 indigenous languages), Chile , Colombia , Costa Rica , Cuba , Dominican Republic , Ecuador , El Salvador , Guatemala , Honduras , Mexico (co-official with 63 indigenous languages), Nicaragua , Panama , Paraguay (co-official with Guaraní ), Peru (co-official with Quechua , Aymara , and "the other indigenous languages"), Puerto Rico (co-official with English), Uruguay , and Venezuela . Spanish language has
8670-419: The midbrain is the diencephalon ; it works with hormones and homeostasis . The pineal body is just above the diencephalon; it detects light, maintains circadian rhythms, and controls color changes. The midbrain contains the two optic lobes . These are very large in species that hunt by sight, such as rainbow trout and cichlids . The hindbrain controls swimming and balance.The single-lobed cerebellum
8772-443: The nine largest families; from largest to smallest, these are Cyprinidae , Gobiidae , Cichlidae , Characidae , Loricariidae , Balitoridae , Serranidae , Labridae , and Scorpaenidae . About 64 families are monotypic , containing only one species. Fish range in size from the huge 16-metre (52 ft) whale shark to some tiny teleosts only 8-millimetre (0.3 in) long, such as the cyprinid Paedocypris progenetica and
8874-679: The northwest of the African mainland. The Spanish spoken in the Canary Islands traces its origins back to the Castilian conquest in the 15th century , and, in addition to a resemblance to Western Andalusian speech patterns, it also features strong influence from the Spanish varieties spoken in the Americas, which in turn have also been influenced historically by Canarian Spanish. The Spanish spoken in North Africa by native bilingual speakers of Arabic or Berber who also speak Spanish as
8976-500: The ocean. Fish are caught for recreation , or raised by fishkeepers as ornaments for private and public exhibition in aquaria and garden ponds . Fish have had a role in human culture through the ages, serving as deities , religious symbols, and as the subjects of art, books and movies. The word fish is inherited from Proto-Germanic , and is related to German Fisch , the Latin piscis and Old Irish īasc , though
9078-432: The only Western European subspecies of wolf whose hunting remained legal, yet only in Spain. Nonetheless, very few hunting permits were given every year, strictly north of the Douro river . Along with the difficulty of their hunt by virtue of their vigilant nature and the rarity of their sightings, they were strongly desired by many European hunters as a big-game trophy . The Iberian wolf Canis lupus signatus Cabrera 1907
9180-402: The open ocean. Because the water all around is the same colour, reflecting an image of the water offers near-invisibility. Fish have a closed-loop circulatory system . The heart pumps the blood in a single loop throughout the body; for comparison, the mammal heart has two loops, one for the lungs to pick up oxygen, one for the body to deliver the oxygen. In fish, the heart pumps blood through
9282-533: The population. Many northern Moroccans have rudimentary knowledge of Spanish, with Spanish being particularly significant in areas adjacent to Ceuta and Melilla. Spanish also has a presence in the education system of the country (through either selected education centers implementing Spain's education system, primarily located in the North, or the availability of Spanish as foreign language subject in secondary education). In Western Sahara , formerly Spanish Sahara ,
9384-507: The surrounding water, though some large active swimmers like white shark and tuna can hold a higher core temperature . Many fish can communicate acoustically with each other, such as during courtship displays . The earliest fish appeared during the Cambrian as small filter feeders ; they continued to evolve through the Paleozoic , diversifying into many forms. The earliest fish with dedicated respiratory gills and paired fins ,
9486-415: The taxon Osteichthyes as a clade, which now includes all tetrapods". The biodiversity of extant fish is unevenly distributed among the various groups; teleosts , bony fishes able to protrude their jaws , make up 96% of fish species. The cladogram shows the evolutionary relationships of all groups of living fishes (with their respective diversity ) and the tetrapods. Extinct groups are marked with
9588-537: The term español (Spanish). According to the Royal Spanish Academy, español derives from the Occitan word espaignol and that, in turn, derives from the Vulgar Latin * hispaniolus ('of Hispania'). Hispania was the Roman name for the entire Iberian Peninsula . There are other hypotheses apart from the one suggested by the Royal Spanish Academy. Spanish philologist Ramón Menéndez Pidal suggested that
9690-444: The theories of Ramón Menéndez Pidal , local sociolects of Vulgar Latin evolved into Spanish, in the north of Iberia, in an area centered in the city of Burgos , and this dialect was later brought to the city of Toledo , where the written standard of Spanish was first developed, in the 13th century. In this formative stage, Spanish developed a strongly differing variant from its close cousin, Leonese , and, according to some authors,
9792-640: The third most used language on the Internet , after English and Chinese. Spanish is the official language of Spain . Upon the emergence of the Castilian Crown as the dominant power in the Iberian Peninsula by the end of the Middle Ages, the Romance vernacular associated with this polity became increasingly used in instances of prestige and influence, and the distinction between "Castilian" and "Spanish" started to become blurred. Hard policies imposing
9894-652: The upper lips, the dark marks on the tail, and a pair of dark marks in its front legs that give it its subspecies name, signatus ("marked"). The subspecies differentiation may have developed at the end of the Pleistocene Ice Ages due to the isolation of the Iberian Peninsula when glacier barriers grew in the Pyrenees and eventually reached the Gulf of Biscay in the west and the Mediterranean in
9996-596: The world's most widely spoken Romance language. The country with the largest population of native speakers is Mexico . Spanish is part of the Ibero-Romance language group , in which the language is also known as Castilian ( castellano ). The group evolved from several dialects of Vulgar Latin in Iberia after the collapse of the Western Roman Empire in the 5th century. The oldest Latin texts with traces of Spanish come from mid-northern Iberia in
10098-634: Was developed in the cities of Toledo , in the 13th to 16th centuries, and Madrid , from the 1570s. The development of the Spanish sound system from that of Vulgar Latin exhibits most of the changes that are typical of Western Romance languages , including lenition of intervocalic consonants (thus Latin vīta > Spanish vida ). The diphthongization of Latin stressed short e and o —which occurred in open syllables in French and Italian, but not at all in Catalan or Portuguese—is found in both open and closed syllables in Spanish, as shown in
10200-622: Was distinguished by a heavy Basque influence (see Iberian Romance languages ). This distinctive dialect spread to southern Spain with the advance of the Reconquista , and meanwhile gathered a sizable lexical influence from the Arabic of Al-Andalus , much of it indirectly, through the Romance Mozarabic dialects (some 4,000 Arabic -derived words, make up around 8% of the language today). The written standard for this new language
10302-713: Was followed by a vowel that did not diphthongize. The h- , still preserved in spelling, is now silent in most varieties of the language, although in some Andalusian and Caribbean dialects, it is still aspirated in some words. Because of borrowings from Latin and neighboring Romance languages, there are many f -/ h - doublets in modern Spanish: Fernando and Hernando (both Spanish for "Ferdinand"), ferrero and herrero (both Spanish for "smith"), fierro and hierro (both Spanish for "iron"), and fondo and hondo (both words pertaining to depth in Spanish, though fondo means "bottom", while hondo means "deep"); additionally, hacer ("to make")
10404-462: Was revoked by Michel Temer after the impeachment of Dilma Rousseff . In many border towns and villages along Paraguay and Uruguay, a mixed language known as Portuñol is spoken. Equatorial Guinea is the only Spanish-speaking country located entirely in Africa, with the language introduced during the Spanish colonial period . Enshrined in the constitution as an official language (alongside French and Portuguese), Spanish features prominently in
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