108-423: Anaplectidae Blaberidae Blattidae Corydiidae Cryptocercidae Ectobiidae Lamproblattidae Nocticolidae Tryonicidae Cockroaches (or roaches ) are insects belonging to the order Blattodea (Blattaria). About 30 cockroach species out of 4,600 are associated with human habitats . Some species are well-known pests . Modern cockroaches are an ancient group that first appeared during
216-585: A communist-led revolution in Afghanistan, Soviet forces invaded the country, ultimately starting the Soviet–Afghan War . In May 1988, the Soviets started to withdraw from Afghanistan , due to international opposition, persistent anti-Soviet guerrilla warfare, and a lack of support by Soviet citizens. From 1985 onwards, the last Soviet leader Mikhail Gorbachev , who sought to enact liberal reforms in
324-653: A troika was designated to take charge. Eventually Joseph Stalin , the General Secretary of the Communist Party , managed to suppress all opposition factions and consolidate power in his hands to become the country's dictator by the 1930s. Leon Trotsky , the main proponent of world revolution , was exiled from the Soviet Union in 1929, and Stalin's idea of Socialism in One Country became
432-611: A Varangian from the Rus' people , named Rurik , was elected ruler of Novgorod in 862. In 882, his successor Oleg ventured south and conquered Kiev , which had been previously paying tribute to the Khazars . Rurik's son Igor and Igor's son Sviatoslav subsequently subdued all local East Slavic tribes to Kievan rule, destroyed the Khazar Khaganate, and launched several military expeditions to Byzantium and Persia . In
540-408: A common shelter, overlapping generations, non-closure of groups, equal reproductive potential of group members, an absence of task specialization, high levels of social dependence, central place foraging, social information transfer, kin recognition , and a meta-population structure. There is evidence that a few species of group-living roaches in the genera Melyroidea and Aclavoidea may exhibit
648-510: A crisis. By 1991, economic and political turmoil began to boil over as the Baltic states chose to secede from the Soviet Union. On 17 March, a referendum was held, in which the vast majority of participating citizens voted in favour of changing the Soviet Union into a renewed federation . In June 1991, Boris Yeltsin became the first directly elected President in Russian history when he
756-736: A decisive role for the Allies in World War II by leading large-scale efforts on the Eastern Front . With the onset of the Cold War , it competed with the United States for ideological dominance and international influence . The Soviet era of the 20th century saw some of the most significant Russian technological achievements , including the first human-made satellite and the first human expedition into outer space . In 1991,
864-664: A global superpower. After World War II, according to the Potsdam Conference , the Red Army occupied parts of Eastern and Central Europe, including East Germany and the eastern regions of Austria . Dependent communist governments were installed in the Eastern Bloc satellite states. After becoming the world's second nuclear power , the Soviet Union established the Warsaw Pact alliance, and entered into
972-469: A group. In one study, isolated cockroaches were less likely to leave their shelters and explore, spent less time eating, interacted less with conspecifics when exposed to them, and, among males, took longer to recognize receptive females. Because these changes occurred in many contexts, the authors suggested them as constituting a behavioral syndrome . These effects might have been due either to reduced metabolic and developmental rates in isolated individuals or
1080-800: A large confederacy, which was subsequently taken over by the Cumans and the Kipchaks . The ancestors of Russians are among the Slavic tribes that separated from the Proto-Indo-Europeans, who appeared in the northeastern part of Europe c. 1500 years ago. The East Slavs gradually settled western Russia (approximately between modern Moscow and Saint-Petersburg ) in two waves: one moving from Kiev towards present-day Suzdal and Murom and another from Polotsk towards Novgorod and Rostov . Prior to Slavic migration, that territory
1188-500: A member state of the G20 , SCO , BRICS , APEC , OSCE , and WTO ; and the leading member state of post-Soviet organisations such as CIS , CSTO , and EAEU/EEU . It possesses the largest stockpile of nuclear weapons and has the third-highest military expenditure . Russia is generally considered a great power and is a regional power . Internationally, Russia ranks very low in measurements of democracy , human rights and freedom of
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#17327982972781296-545: A million inhabitants. Its capital and largest city is Moscow . Saint Petersburg is Russia's second-largest city and its cultural capital . The East Slavs emerged as a recognised group in Europe between the 3rd and 8th centuries CE. The first East Slavic state, Kievan Rus' , arose in the 9th century, and in 988, it adopted Orthodox Christianity from the Byzantine Empire . Kievan Rus' ultimately disintegrated;
1404-640: A more conservative approach, proposing to retain the termites as the Termitoidae , an epifamily within the order. Such a measure preserves the classification of termites at family level and below. Most species of cockroach are about the size of a thumbnail , but several species are notably larger. The world's heaviest cockroach is the Australian giant burrowing cockroach Macropanesthia rhinoceros , which can reach 8 centimetres (3 in) in length and weigh up to 35 grams (1.2 oz). Comparable in size
1512-487: A number of conflicts in former Soviet states and other countries , including its war with Georgia in 2008 and its war with Ukraine since 2014, which has involved the internationally unrecognised annexations of Ukrainian territory including Crimea in 2014 and four other regions in 2022 during an ongoing invasion . Russia is a permanent member of the United Nations Security Council ;
1620-402: A pair of spiracles for respiration. In addition to the spiracles, the final segment consists of a pair of cerci , a pair of anal styles, the anus and the external genitalia. Males have an aedeagus through which they secrete sperm during copulation, while females have spermatheca for storing sperm and an ovipositor through which the oothecae are laid. Cockroaches are abundant throughout
1728-421: A reproductive division of labor, which, if confirmed, would make these the only genuinely eusocial lineage known among roaches, in contrast to the subsocial members of the genus Cryptocercus . Some species make a buzzing noise while other cockroaches make a chirping noise. Gromphadorhina species and Archiblatta hoeveni produce sound through the modified spiracles on the fourth abdominal segment. In
1836-571: A secret protocol within the Molotov–Ribbentrop Pact with Nazi Germany . The Soviet Union later invaded Finland , and occupied and annexed the Baltic states , as well as parts of Romania . On 22 June 1941, Germany invaded the Soviet Union , opening the Eastern Front , the largest theater of World War II. Eventually, some 5 million Red Army troops were captured by the Nazis;
1944-412: A separate order Isoptera to cockroaches. However, recent genetic evidence strongly suggests that they evolved directly from 'true' cockroaches, and many authors now place them as an " epifamily " of Blattodea. This evidence supported a hypothesis suggested in 1934 that termites are closely related to the wood-eating cockroaches (genus Cryptocercus ). This hypothesis was originally based on similarity of
2052-592: A simple set of individual interactions. Daily rhythms may also be regulated by a complex set of hormonal controls of which only a small subset have been understood. In 2005, the role of one of these proteins, pigment dispersing factor (PDF), was isolated and found to be a key mediator in the circadian rhythms of the cockroach. Pest species adapt readily to a variety of environments, but prefer warm conditions found within buildings. Many tropical species prefer even warmer environments. Cockroaches are mainly nocturnal and run away when exposed to light. An exception to this
2160-462: A struggle for global dominance, known as the Cold War , with the rivalling United States and NATO . After Stalin's death in 1953 and a short period of collective rule , the new leader Nikita Khrushchev denounced Stalin and launched the policy of de-Stalinization , releasing many political prisoners from the Gulag labour camps. The general easement of repressive policies became known later as
2268-548: A system of tubes called tracheae which are attached to openings called spiracles on all body segments. When the carbon dioxide level in the insect rises high enough, valves on the spiracles open and carbon dioxide diffuses out and oxygen diffuses in. The tracheal system branches repeatedly, the finest tracheoles bringing air directly to each cell , allowing gaseous exchange to take place. While cockroaches do not have lungs as do vertebrates , and can continue to respire if their heads are removed, in some very large species,
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#17327982972782376-524: Is also sometimes used to refer to extinct cockroach relatives. The earliest cockroach-like fossils ("blattopterans" or " roachoids ") are from the Carboniferous period 320 million years ago. Fossil roachoids are considered the common ancestor of both mantises and modern cockroaches, and are distinguished from the latter by the presence of a long external ovipositor . As the body, hind wings and mouthparts are not preserved in fossils frequently,
2484-424: Is and how many other cockroaches there are. A study used specially scented roach-sized robots that seem real to the roaches to demonstrate that once there are enough insects in a place to form a critical mass , the roaches accepted the collective decision on where to hide, even if this was an unusually lit place. When reared in isolation, German cockroaches show behavior that is different from behavior when reared in
2592-729: Is the Asian cockroach , which flies mostly at night but is attracted to brightly lit surfaces and pale colors. Gregarious cockroaches display collective decision-making when choosing food sources. When a sufficient number of individuals (a "quorum") exploits a food source, this signals to newcomer cockroaches that they should stay there longer rather than leave for elsewhere. Other mathematical models have been developed to explain aggregation dynamics and conspecific recognition. Cooperation and competition are balanced in cockroach group decision-making behavior. Cockroaches appear to use just two pieces of information to decide where to go, namely how dark it
2700-468: Is the Central American giant cockroach Blaberus giganteus . The longest cockroach species is Megaloblatta longipennis , which can reach 97 mm ( 3 + 7 ⁄ 8 in) in length and 45 mm ( 1 + 3 ⁄ 4 in) across. A Central and South American species, Megaloblatta blaberoides , has the largest wingspan of up to 185 mm ( 7 + 1 ⁄ 4 in). At
2808-628: The Oxford English Dictionary , the English name Russia first appeared in the 14th century, borrowed from Medieval Latin : Russia , used in the 11th century and frequently in 12th-century British sources, in turn derived from Russi , 'the Russians'; and the suffix -ia . In modern historiography, this state is usually denoted as Kievan Rus' after its capital city. Another Medieval Latin name for Rus'
2916-554: The Battle of Kursk in the summer of 1943. Another German failure was the Siege of Leningrad , in which the city was fully blockaded on land between 1941 and 1944 by German and Finnish forces, and suffered starvation and more than a million deaths, but never surrendered. Soviet forces steamrolled through Eastern and Central Europe in 1944–1945 and captured Berlin in May 1945. In August 1945,
3024-765: The Brusilov Offensive of the Imperial Russian Army almost completely destroyed the Austro-Hungarian Army . However, the already-existing public distrust of the regime was deepened by the rising costs of war, high casualties , and rumors of corruption and treason. All this formed the climate for the Russian Revolution of 1917, carried out in two major acts. In early 1917, Nicholas II was forced to abdicate ; he and his family were imprisoned and later executed during
3132-475: The Congress of Vienna , which defined the map of post-Napoleonic Europe. The officers who pursued Napoleon into Western Europe brought ideas of liberalism back to Russia, and attempted to curtail the tsar's powers during the abortive Decembrist revolt of 1825. At the end of the conservative reign of Nicholas I (1825–1855), a zenith period of Russia's power and influence in Europe, was disrupted by defeat in
3240-693: The Crimean War . Nicholas's successor Alexander II (1855–1881) enacted significant changes throughout the country, including the emancipation reform of 1861 . These reforms spurred industrialisation, and modernised the Imperial Russian Army, which liberated much of the Balkans from Ottoman rule in the aftermath of the 1877–1878 Russo-Turkish War . During most of the 19th and early 20th century, Russia and Britain colluded over Afghanistan and its neighbouring territories in Central and South Asia;
3348-592: The Grand Duchy of Moscow led the unification of Russian lands, leading to the proclamation of the Tsardom of Russia in 1547. By the early 18th century, Russia had vastly expanded through conquest, annexation, and the efforts of Russian explorers , developing into the Russian Empire , which remains the third-largest empire in history . However, with the Russian Revolution in 1917, Russia's monarchic rule
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3456-614: The Great Northern War (1700–1721), securing Russia's access to the sea and sea trade. In 1703, on the Baltic Sea, Peter founded Saint Petersburg as Russia's new capital. Throughout his rule, sweeping reforms were made , which brought significant Western European cultural influences to Russia. He was succeeded by Catherine I (1725–1727), followed by Peter II (1727–1730), and Anna . The reign of Peter I's daughter Elizabeth in 1741–1762 saw Russia's participation in
3564-701: The Khrushchev Thaw . At the same time, Cold War tensions reached its peak when the two rivals clashed over the deployment of the United States Jupiter missiles in Turkey and Soviet missiles in Cuba . In 1957, the Soviet Union launched the world's first artificial satellite , Sputnik 1 , thus starting the Space Age . Russian cosmonaut Yuri Gagarin became the first human to orbit
3672-491: The Late Jurassic , with their ancestors, known as " roachoids ", likely originating during the Carboniferous period around 320 million years ago. Those early ancestors, however, lacked the internal ovipositors of modern roaches. Cockroaches are somewhat generalized insects lacking special adaptations (such as the sucking mouthparts of aphids and other true bugs ); they have chewing mouthparts and are probably among
3780-707: The Oldowan period in the early Lower Paleolithic . About 2 million years ago, representatives of Homo erectus migrated to the Taman Peninsula in southern Russia. Flint tools, some 1.5 million years old, have been discovered in the North Caucasus . Radiocarbon dated specimens from Denisova Cave in the Altai Mountains estimate the oldest Denisovan specimen lived 195–122,700 years ago. Fossils of Denny , an archaic human hybrid that
3888-568: The Rurik dynasty that ruled Kievan Rus' collectively. Kiev's dominance waned, to the benefit of Vladimir-Suzdal in the north-east, the Novgorod Republic in the north, and Galicia-Volhynia in the south-west. By the 12th century, Kiev lost its pre-eminence and Kievan Rus' had fragmented into different principalities. Prince Andrey Bogolyubsky sacked Kiev in 1169 and made Vladimir his base, leading to political power being shifted to
3996-843: The Russian Civil War . The monarchy was replaced by a shaky coalition of political parties that declared itself the Provisional Government , and proclaimed the Russian Republic . On 19 January [ O.S. 6 January], 1918, the Russian Constituent Assembly declared Russia a democratic federal republic (thus ratifying the Provisional Government's decision). The next day the Constituent Assembly
4104-632: The Seven Years' War (1756–1763). During the conflict, Russian troops overran East Prussia , reaching Berlin. However, upon Elizabeth's death, all these conquests were returned to the Kingdom of Prussia by pro-Prussian Peter III of Russia . Catherine II ("the Great"), who ruled in 1762–1796, presided over the Russian Age of Enlightenment . She extended Russian political control over
4212-852: The Siberian River Routes , and by the mid-17th century, there were Russian settlements in eastern Siberia, on the Chukchi Peninsula , along the Amur River , and on the coast of the Pacific Ocean. In 1648, Semyon Dezhnyov became the first European to navigate through the Bering Strait . Under Peter the Great , Russia was proclaimed an empire in 1721, and established itself as one of the European great powers. Ruling from 1682 to 1725, Peter defeated Sweden in
4320-653: The Soviet Union , by joining the Russian SFSR into a single state with the Byelorussian , Transcaucasian , and Ukrainian republics. Eventually internal border changes and annexations during World War II created a union of 15 republics ; the largest in size and population being the Russian SFSR, which dominated the union politically, culturally, and economically. Following Lenin's death in 1924,
4428-440: The cuticle may enable cockroaches to distinguish between different populations of cockroach by odor. The behaviors involved have been studied in only a few species, but German cockroaches leave fecal trails with an odor gradient. Other cockroaches follow such trails to discover sources of food and water, and where other cockroaches are hiding. Thus, cockroaches have emergent behavior , in which group or swarm behavior emerges from
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4536-666: The 10th to 11th centuries, Kievan Rus' became one of the largest and most prosperous states in Europe. The reigns of Vladimir the Great (980–1015) and his son Yaroslav the Wise (1019–1054) constitute the Golden Age of Kiev, which saw the acceptance of Orthodox Christianity from Byzantium , and the creation of the first East Slavic written legal code , the Russkaya Pravda . The age of feudalism and decentralisation had come, marked by constant in-fighting between members of
4644-566: The 16th century, Russia expanded east of the Ural Mountains . However, the Tsardom was weakened by the long and unsuccessful Livonian War against the coalition of the Kingdom of Poland and the Grand Duchy of Lithuania (later the united Polish–Lithuanian Commonwealth ), the Kingdom of Sweden , and Denmark–Norway for access to the Baltic coast and sea trade. In 1572, an invading army of Crimean Tatars were thoroughly defeated in
4752-599: The Byzantine double-headed eagle his own, and eventually Russia's, coat-of-arms. Vasili III united all of Russia by annexing the last few independent Russian states in the early 16th century. In development of the Third Rome ideas, the grand duke Ivan IV ("the Terrible") was officially crowned the first tsar of Russia in 1547. The tsar promulgated a new code of laws ( Sudebnik of 1550 ), established
4860-616: The Earth, aboard the Vostok ;1 crewed spacecraft on 12 April 1961 . Following the ousting of Khrushchev in 1964, another period of collective rule ensued, until Leonid Brezhnev became the leader. The era of the 1970s and the early 1980s was later designated as the Era of Stagnation . The 1965 Kosygin reform aimed for partial decentralisation of the Soviet economy . In 1979, after
4968-415: The Polish–Lithuanian Commonwealth and annexed most of its territories into Russia , making it the most populous country in Europe. In the south, after the successful Russo-Turkish Wars against the Ottoman Empire , Catherine advanced Russia's boundary to the Black Sea, by dissolving the Crimean Khanate , and annexing Crimea . As a result of victories over Qajar Iran through the Russo-Persian Wars , by
5076-692: The Provisional Government and gave full governing power to the soviets, leading to the creation of the world's first socialist state . The Russian Civil War broke out between the anti-communist White movement and the Bolsheviks with its Red Army . In the aftermath of signing the Treaty of Brest-Litovsk that concluded hostilities with the Central Powers of World War I ; Bolshevist Russia surrendered most of its western territories, which hosted 34% of its population, 54% of its industries, 32% of its agricultural land, and roughly 90% of its coal mines. The Allied powers launched an unsuccessful military intervention in support of anti-communist forces. In
5184-406: The Red Army invaded Manchuria and ousted the Japanese from Northeast Asia, contributing to the Allied victory over Japan. The 1941–1945 period of World War II is known in Russia as the Great Patriotic War . The Soviet Union, along with the United States, the United Kingdom and China were considered the Big Four of Allied powers in World War II, and later became the Four Policemen , which
5292-436: The Russian SFSR emerged from the dissolution of the Soviet Union as the Russian Federation. A new constitution was adopted, which established a federal semi-presidential system . Since the turn of the century, Russia's political system has been dominated by Vladimir Putin , under whom the country has experienced democratic backsliding and become an authoritarian dictatorship . Russia has been militarily involved in
5400-403: The Russian autocratic state. The destruction of Kievan Rus' saw the eventual rise of the Grand Duchy of Moscow , initially a part of Vladimir-Suzdal . While still under the domain of the Mongol - Tatars and with their connivance, Moscow began to assert its influence in the region in the early 14th century, gradually becoming the leading force in the "gathering of the Russian lands". When
5508-476: The Russian parliament culminated in a constitutional crisis which ended violently through military force. During the crisis, Yeltsin was backed by Western governments, and over 100 people were killed. In December, a referendum was held and approved, which introduced a new constitution, giving the president enormous powers. The 1990s were plagued by armed conflicts in the North Caucasus , both local ethnic skirmishes and separatist Islamist insurrections. From
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#17327982972785616-408: The Soviet Union led Russia into a deep and prolonged depression. During and after the disintegration of the Soviet Union, wide-ranging reforms including privatisation and market and trade liberalisation were undertaken, including radical changes along the lines of " shock therapy ". The privatisation largely shifted control of enterprises from state agencies to individuals with inside connections in
5724-523: The Soviet Union. The transitional disorganisation of the country's agriculture, combined with the harsh state policies and a drought, led to the Soviet famine of 1932–1933 ; which killed 5.7 to 8.7 million, 3.3 million of them in the Russian SFSR. The Soviet Union, ultimately, made the costly transformation from a largely agrarian economy to a major industrial powerhouse within a short span of time. The Soviet Union entered World War II on 17 September 1939 with its invasion of Poland , in accordance with
5832-406: The Soviet system, introduced the policies of glasnost (openness) and perestroika (restructuring) in an attempt to end the period of economic stagnation and to democratise the government . This, however, led to the rise of strong nationalist and separatist movements across the country. Prior to 1991, the Soviet economy was the world's second-largest, but during its final years, it went into
5940-493: The Ukrainian leader, Bohdan Khmelnytsky , offered to place Ukraine under the protection of the Russian tsar, Alexis ; whose acceptance of this offer led to another Russo-Polish War . Ultimately, Ukraine was split along the Dnieper , leaving the eastern part, ( Left-bank Ukraine and Kiev ) under Russian rule. In the east, the rapid Russian exploration and colonisation of vast Siberia continued, hunting for valuable furs and ivory. Russian explorers pushed eastward primarily along
6048-516: The Volga-Dnieper region of southern Russia and Ukraine as the urheimat of the Proto-Indo-Europeans . Early Indo-European migrations from the Pontic–Caspian steppe of Ukraine and Russia spread Yamnaya ancestry and Indo-European languages across large parts of Eurasia. Nomadic pastoralism developed in the Pontic–Caspian steppe beginning in the Chalcolithic . Remnants of these steppe civilizations were discovered in places such as Ipatovo , Sintashta , Arkaim , and Pazyryk , which bear
6156-408: The body musculature may contract rhythmically to forcibly move air in and out of the spiracles; this may be considered a form of breathing. Anaplectidae Anaplectidae is a family of cockroaches in the order Blattodea . Previously placed as a subfamily of the Ectobiidae there are presently (2020) two genera and more than 90 described species in Anaplectidae. These two genera belong to
6264-423: The capital Moscow. In 1612, the Poles were forced to retreat by the Russian volunteer corps, led by merchant Kuzma Minin and prince Dmitry Pozharsky . The Romanov dynasty acceded to the throne in 1613 by the decision of the Zemsky Sobor, and the country started its gradual recovery from the crisis. Russia continued its territorial growth through the 17th century, which was the age of the Cossacks . In 1654,
6372-459: The continent. 450 species are found in Australia . Only about four widespread species are commonly regarded as pests. Cockroaches occupy a wide range of habitats. Many live in leaf litter , among the stems of matted vegetation, in rotting wood, in holes in stumps, in cavities under bark, under log piles and among debris. Some live in arid regions and have developed mechanisms to survive without access to water sources. Others are aquatic, living near
6480-414: The control of the Golden Horde and consolidated the whole of northern Rus' under Moscow's dominion, and was the first Russian ruler to take the title "Grand Duke of all Rus'". After the fall of Constantinople in 1453, Moscow claimed succession to the legacy of the Eastern Roman Empire . Ivan III married Sophia Palaiologina , the niece of the last Byzantine emperor Constantine XI , and made
6588-782: The country was more often referred to by its inhabitants as Rus ' , the Russian land ( Russkaia zemlia ), or the Muscovite state ( Moskovskoe gosudarstvo ), among other variations. In 1721, Peter the Great changed the name of the state from Tsardom of Russia ( Russian : Русское царство , romanized : Russkoye tsarstvo ) or Tsardom of Muscovy ( Russian : Московское царство , romanized : Moskovskoye tsarstvo ) to Russian Empire ( Rossiiskaia imperiia ). There are several words in Russian which translate to "Russians" in English. The noun and adjective русский , russkiy refers to ethnic Russians . The adjective российский , rossiiskiy denotes Russian citizens regardless of ethnicity. The same applies to
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#17327982972786696-400: The crucial Battle of Molodi . The death of Ivan's sons marked the end of the ancient Rurik dynasty in 1598, and in combination with the disastrous famine of 1601–1603 , led to a civil war, the rule of pretenders, and foreign intervention during the Time of Troubles in the early 17th century. The Polish–Lithuanian Commonwealth , taking advantage, occupied parts of Russia, extending into
6804-616: The day in crevices, among dead leaves, in bird and insect nests or among epiphytes , emerging at night to feed. Cockroaches are social insects; a large number of species are either gregarious or inclined to aggregate, and a slightly smaller number exhibit parental care. It used to be thought that cockroaches aggregated because they were reacting to environmental cues, but it is now believed that pheromones are involved in these behaviors. Some species secrete these in their feces with gut microbial symbionts being involved, while others use glands located on their mandibles . Pheromones produced by
6912-405: The earliest known traces of horses in warfare . The genetic makeup of speakers of the Uralic language family in northern Europe was shaped by migration from Siberia that began at least 3,500 years ago. In the 3rd to 4th centuries CE, the Gothic kingdom of Oium existed in southern Russia, which was later overrun by Huns . Between the 3rd and 6th centuries CE, the Bosporan Kingdom , which
7020-644: The fact that the isolated individuals had not had a training period to learn about what others were like via their antennae. Individual American cockroaches appear to have consistently different "personalities" regarding how they seek shelter. In addition, group personality is not simply the sum of individual choices, but reflects conformity and collective decision-making. The gregarious German and American cockroaches have elaborate social structure , chemical signaling, and "social herd" characteristics. Lihoreau and his fellow researchers stated: The social biology of domiciliary cockroaches ... can be characterized by
7128-445: The family Anaplectidae: Russia Russia , or the Russian Federation , is a country spanning Eastern Europe and North Asia . It is the largest country in the world by area , extending across eleven time zones and sharing land borders with fourteen countries . It is the world's ninth-most populous country and Europe's most populous country . Russia is a highly urbanised country including 16 population centres with over
7236-417: The first Europeans to reach and colonise Alaska . In 1803–1806, the first Russian circumnavigation was made. In 1820, a Russian expedition discovered the continent of Antarctica . During the Napoleonic Wars , Russia joined alliances with various European powers, and fought against France. The French invasion of Russia at the height of Napoleon's power in 1812 reached Moscow, but eventually failed as
7344-399: The first Russian feudal representative body (the Zemsky Sobor ), revamped the military, curbed the influence of the clergy, and reorganised local government. During his long reign, Ivan nearly doubled the already large Russian territory by annexing the three Tatar khanates: Kazan and Astrakhan along the Volga , and the Khanate of Sibir in southwestern Siberia. Ultimately, by the end of
7452-418: The first half of the 19th century, Russia also conquered the Caucasus . Catherine's successor, her son Paul , was unstable and focused predominantly on domestic issues . Following his short reign, Catherine's strategy was continued with Alexander I's (1801–1825) wresting of Finland from the weakened Sweden in 1809, and of Bessarabia from the Ottomans in 1812. In North America, the Russians became
7560-520: The former species, several different hisses are produced, including disturbance sounds, produced by adults and larger nymphs; and aggressive, courtship and copulatory sounds produced by adult males. Henschoutedenia epilamproides has a stridulatory organ between its thorax and abdomen, but the purpose of the sound produced is unclear. Several Australian species practice acoustic and vibration behaviour as an aspect of courtship. They have been observed producing hisses and whistles from air forced through
7668-420: The front legs are the shortest and the hind legs the longest, providing the main propulsive power when the insect runs. The spines on the legs were earlier considered to be sensory, but observations of the insect's gait on sand and wire meshes have demonstrated that they help in locomotion on difficult terrain. The structures have been used as inspiration for robotic legs. The abdomen has ten segments, each having
7776-485: The government, which led to the rise of Russian oligarchs . Many of the newly rich moved billions in cash and assets outside of the country in an enormous capital flight . The depression of the economy led to the collapse of social services—the birth rate plummeted while the death rate skyrocketed, and millions plunged into poverty; while extreme corruption, as well as criminal gangs and organised crime rose significantly. In late 1993, tensions between Yeltsin and
7884-424: The gregarious German cockroach , have an elaborate social structure involving common shelter, social dependence, information transfer and kin recognition. Cockroaches have appeared in human culture since classical antiquity . They are popularly depicted as large, dirty pests, although the majority of species are small and inoffensive and live in a wide range of habitats around the world. Cockroaches are members of
7992-575: The insect is to utilize cellulose; however, some species secrete cellulase in their saliva , and the wood-eating cockroach, Panesthia cribrata , is able to survive indefinitely on a diet of crystallized cellulose while being free of microorganisms. The similarity of these symbionts in the genus Cryptocercus to those in termites are such that these cockroaches have been suggested to be more closely related to termites than to other cockroaches, and current research strongly supports this hypothesis about their relationships. All species studied so far carry
8100-535: The latter deliberately starved to death or otherwise killed 3.3 million Soviet POWs , and a vast number of civilians, as the " Hunger Plan " sought to fulfil Generalplan Ost . Although the Wehrmacht had considerable early success, their attack was halted in the Battle of Moscow . Subsequently, the Germans were dealt major defeats first at the Battle of Stalingrad in the winter of 1942–1943, and then in
8208-479: The light", which in classical Latin was applied not only to cockroaches, but also to mantids . Historically, the name Blattaria was used largely interchangeably with the name Blattodea, but whilst Blattaria was used to refer to 'true' cockroaches exclusively, the Blattodea also includes the termites. The current catalogue of world cockroach species uses the name Blattodea for the group. Another name, Blattoptera ,
8316-686: The meantime, both the Bolsheviks and White movement carried out campaigns of deportations and executions against each other, known respectively as the Red Terror and White Terror . By the end of the violent civil war, Russia's economy and infrastructure were heavily damaged, and as many as 10 million perished during the war, mostly civilians. Millions became White émigrés , and the Russian famine of 1921–1922 claimed up to five million victims. On 30 December 1922, Lenin and his aides formed
8424-609: The more recently coined noun россиянин , rossiianyn , "Russian" in the sense of citizen of the Russian state. According to the Primary Chronicle , the word Rus' is derived from the Rus' people , who were a Swedish tribe, and where the three original members of the Rurikid dynasty came from. The Finnish word for Swedes, ruotsi , has the same origin. Later archeological studies mostly confirmed this theory. The first human settlement on Russia dates back to
8532-625: The most primitive of living Neopteran insects. They are common and hardy insects capable of tolerating a wide range of climates , from Arctic cold to tropical heat. Tropical cockroaches are often much larger than temperate species. Modern cockroaches are not considered to be a monophyletic group, as it has been found based on genetics that termites are deeply nested within the group, with some groups of cockroaches more closely related to termites than they are to other cockroaches, thus rendering Blattaria paraphyletic . Both cockroaches and termites are included into Blattodea. Some species, such as
8640-641: The north-east. Led by Prince Alexander Nevsky , Novgorodians repelled the invading Swedes in the Battle of the Neva in 1240, as well as the Germanic crusaders in the Battle on the Ice in 1242. Kievan Rus' finally fell to the Mongol invasion of 1237–1240, which resulted in the sacking of Kiev and other cities, as well as the death of a major part of the population. The invaders, later known as Tatars , formed
8748-572: The obligate mutualistic endosymbiont bacterium Blattabacterium , with the exception of Nocticola , an Australian cave-dwelling genus without eyes, pigment or wings, which recent genetic studies indicate is a very primitive cockroach. It had previously been thought that all five families of cockroach were descended from a common ancestor that was infected with B. cuenoti . It may be that N. australiensis subsequently lost its symbionts, or alternatively this hypothesis will need to be re-examined. Like other insects, cockroaches breathe through
8856-620: The obstinate resistance in combination with the bitterly cold Russian winter led to a disastrous defeat of invaders, in which the pan-European Grande Armée faced utter destruction. Led by Mikhail Kutuzov and Michael Andreas Barclay de Tolly , the Imperial Russian Army ousted Napoleon and drove throughout Europe in the War of the Sixth Coalition , ultimately entering Paris. Alexander I controlled Russia's delegation at
8964-654: The official line. The continued internal struggle in the Bolshevik party culminated in the Great Purge . Under Stalin's leadership, the government launched a command economy , industrialisation of the largely rural country , and collectivisation of its agriculture . During this period of rapid economic and social change, millions of people were sent to penal labour camps , including many political convicts for their suspected or real opposition to Stalin's rule; and millions were deported and exiled to remote areas of
9072-522: The other end of the size scale, Attaphila cockroaches that live with leaf-cutter ants include some of the world's smallest species, growing to about 3.5 mm in length. Cockroaches are generalized insects with few special adaptations, and may be among the most primitive living Neopteran insects. They have a relatively small head and a broad, flattened body, and most species are reddish-brown to dark brown. They have large compound eyes , two ocelli , and long, flexible antennae . The mouthparts are on
9180-407: The press ; the country also has high levels of perceived corruption . As of 2024, Russia has a high-income economy which ranks eleventh in the world by nominal GDP and fourth at purchasing power parity , relying on its vast mineral and energy resources ; the world's second-largest for oil production and natural gas production . Russia is home to 32 UNESCO World Heritage Sites . According to
9288-492: The relationship of these roachoids and modern cockroaches remains disputed. The earliest definitive fossils of modern crown group cockroaches, specifically Corydiidae , are known from the Late Jurassic rocks of Russia . The evolutionary relationships of the Blattodea (cockroaches and termites) shown in the cladogram are based on Inward, Beccaloni and Eggleton (2007). The cockroach families Anaplectidae , Lamproblattidae , and Tryonicidae are not shown but are placed within
9396-589: The rivalry between the two major European empires came to be known as the Great Game . The late 19th century saw the rise of various socialist movements in Russia. Alexander II was assassinated in 1881 by revolutionary terrorists. The reign of his son Alexander III (1881–1894) was less liberal but more peaceful. Under last Russian emperor, Nicholas II (1894–1917), the Revolution of 1905
9504-876: The seat of the Metropolitan of the Russian Orthodox Church moved to Moscow in 1325, its influence increased. Moscow's last rival, the Novgorod Republic , prospered as the chief fur trade centre and the easternmost port of the Hanseatic League . Led by Prince Dmitry Donskoy of Moscow, the united army of Russian principalities inflicted a milestone defeat on the Mongol-Tatars in the Battle of Kulikovo in 1380. Moscow gradually absorbed its parent duchy and surrounding principalities, including formerly strong rivals such as Tver and Novgorod . Ivan III ("the Great") threw off
9612-422: The second and third thoracic segments. The tegmina , or first pair of wings, are tough and protective; these lay as a shield on top of the membranous hind wings , which are used in flight. All four wings have branching longitudinal veins, as well as multiple cross-veins . The three pairs of legs are sturdy, with large coxae and five claws each. They are attached to each of the three thoracic segments. Of these,
9720-834: The spiracles. Furthermore, in the presence of a potential mate, some cockroaches tap the substrate in a rhythmic, repetitive manner. Acoustic signals may be of greater prevalence amongst perching species, particularly those that live on low vegetation in Australia's tropics. Cockroaches are generally omnivorous ; the American cockroach ( Periplaneta americana ), for example, feeds on a great variety of foodstuffs including bread, fruit, leather, starch in book bindings, paper, glue, skin flakes, hair, dead insects and soiled clothing. Many species of cockroach harbor in their gut symbiotic protozoans and bacteria which are able to digest cellulose . In many species, these symbionts may be essential if
9828-656: The state of the Golden Horde , which ruled over Russia for the next two centuries. Only the Novgorod Republic escaped foreign occupation after it agreed to pay tribute to the Mongols. Galicia-Volhynia would later be absorbed by Lithuania and Poland , while the Novgorod Republic continued to prosper in the north. In the northeast, the Byzantine-Slavic traditions of Kievan Rus' were adapted to form
9936-638: The superfamily Blattoidea. The cockroach families Corydiidae and Ectobiidae were previously known as the Polyphagidae and Blattellidae. Termitidae Rhinotermitidae Kalotermitidae Archotermopsidae Hodotermitidae Mastotermitidae Cryptocercidae (brown-hooded cockroaches) Blattidae (Oriental, American and other cockroaches) Blaberidae (giant cockroaches) Ectobiidae (part) Ectobiidae (part) Corydiidae (sand cockroaches, etc) Nocticolidae (cave cockroaches, etc) Mantodea (mantises) Termites were previously regarded as
10044-541: The superorder Dictyoptera , which includes the termites and mantids , a group of insects once thought to be separate from cockroaches. Currently, 4,600 species and over 460 genera are described worldwide. The name " cockroach " comes from the Spanish word for cockroach, cucaracha , transformed by 1620s English folk etymology into "cock" and "roach". The scientific name derives from the Latin blatta , "an insect that shuns
10152-411: The surface of water bodies, including bromeliad phytotelmata , and diving to forage for food. Most of these respire by piercing the water surface with the tip of the abdomen which acts as a snorkel , but some carry a bubble of air under their thoracic shield when they submerge. Others live in the forest canopy where they may be one of the main types of invertebrate present. Here they may hide during
10260-505: The symbiotic gut flagellates in termites regarded as living fossils and wood-eating cockroaches. Additional evidence emerged when F. A. McKittrick (1965) noted similar morphological characteristics between some termites and cockroach nymphs. The similarities among these cockroaches and termites have led some scientists to reclassify termites as a single family, the Termitidae , within the order Blattodea. Other scientists have taken
10368-523: The time Chechen separatists declared independence in the early 1990s, an intermittent guerrilla war was fought between the rebel groups and Russian forces. Terrorist attacks against civilians were carried out by Chechen separatists, claiming the lives of thousands of Russian civilians. After the dissolution of the Soviet Union, Russia assumed responsibility for settling the latter's external debts. In 1992, most consumer price controls were eliminated, causing extreme inflation and significantly devaluing
10476-449: The underside of the head and include generalized chewing mandibles , salivary glands and various touch and taste receptors. The body is divided into a thorax of three segments and a ten-segmented abdomen. The external surface has a tough exoskeleton which contains calcium carbonate ; this protects the inner organs and provides attachment to muscles. This external exoskeleton is coated with wax to repel water. The wings are attached to
10584-460: The world and live in a wide range of environments, especially in the tropics and subtropics . Cockroaches can withstand extremely low temperatures, allowing them to live in the Arctic . Some species are capable of surviving temperatures of −122 °C (−188 °F) by manufacturing an antifreeze made out of glycerol . In North America , 50 species separated into five families are found throughout
10692-515: Was Ruthenia . In Russian, the current name of the country, Россия ( Rossiya ), comes from the Byzantine Greek name for Rus', Ρωσία ( Rosía ). A new form of the name Rus ' , Росия ( Rosiya ), was borrowed from the Greek term and first attested in 1387. The name Rossiia appeared in Russian sources in the late 15th century, but until the end of the 17th century
10800-449: Was elected President of the Russian SFSR. In August 1991, a coup d'état attempt by members of Gorbachev's government, directed against Gorbachev and aimed at preserving the Soviet Union, instead led to the end of the Communist Party of the Soviet Union. On 25 December 1991, following the dissolution of the Soviet Union , along with contemporary Russia, fourteen other post-Soviet states emerged. The economic and political collapse of
10908-687: Was a Hellenistic polity that succeeded the Greek colonies, was also overwhelmed by nomadic invasions led by warlike tribes such as the Huns and Eurasian Avars . The Khazars , who were of Turkic origin , ruled the steppes between the Caucasus in the south, to the east past the Volga river basin, and west as far as Kyiv on the Dnieper river until the 10th century. After them came the Pechenegs who created
11016-415: Was abolished and eventually replaced by the Russian SFSR —the world's first constitutionally socialist state . Following the Russian Civil War , the Russian SFSR established the Soviet Union with three other Soviet republics , within which it was the largest and principal constituent. At the expense of millions of lives , the Soviet Union underwent rapid industrialisation in the 1930s and later played
11124-670: Was dissolved by the All-Russian Central Executive Committee . An alternative socialist establishment co-existed, the Petrograd Soviet , wielding power through the democratically elected councils of workers and peasants, called soviets . The rule of the new authorities only aggravated the crisis in the country instead of resolving it, and eventually, the October Revolution , led by Bolshevik leader Vladimir Lenin , overthrew
11232-515: Was found at Kostyonki–Borshchyovo , and at Sungir , dating back to 34,600 years ago—both in western Russia . Humans reached Arctic Russia at least 40,000 years ago, in Mamontovaya Kurya . Ancient North Eurasian populations from Siberia genetically similar to Mal'ta–Buret' culture and Afontova Gora were an important genetic contributor to Ancient Native Americans and Eastern Hunter-Gatherers . The Kurgan hypothesis places
11340-581: Was half Neanderthal and half Denisovan, and lived some 90,000 years ago, was also found within the latter cave. Russia was home to some of the last surviving Neanderthals, from about 45,000 years ago, found in Mezmaiskaya cave . The first trace of an early modern human in Russia dates back to 45,000 years, in Western Siberia . The discovery of high concentration cultural remains of anatomically modern humans , from at least 40,000 years ago,
11448-584: Was populated by Finno-Ugrian peoples. From the 7th century onwards, the incoming East Slavs slowly assimilated the native Finno-Ugrians. The establishment of the first East Slavic states in the 9th century coincided with the arrival of Varangians , the Vikings who ventured along the waterways extending from the eastern Baltic to the Black and Caspian Seas. According to the Primary Chronicle ,
11556-487: Was the foundation of the United Nations Security Council . During the war, Soviet civilian and military death were about 26–27 million , accounting for about half of all World War II casualties . The Soviet economy and infrastructure suffered massive devastation, which caused the Soviet famine of 1946–1947 . However, at the expense of a large sacrifice, the Soviet Union emerged as
11664-695: Was triggered by the humiliating failure of the Russo-Japanese War . The uprising was put down, but the government was forced to concede major reforms ( Russian Constitution of 1906 ), including granting freedoms of speech and assembly , the legalisation of political parties, and the creation of an elected legislative body, the State Duma . In 1914, Russia entered World War I in response to Austria-Hungary 's declaration of war on Russia's ally Serbia , and fought across multiple fronts while isolated from its Triple Entente allies. In 1916,
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