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Peopling of the Americas

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The peopling of the Americas began when Paleolithic hunter-gatherers ( Paleo-Indians ) entered North America from the North Asian Mammoth steppe via the Beringia land bridge , which had formed between northeastern Siberia and western Alaska due to the lowering of sea level during the Last Glacial Maximum (26,000 to 19,000 years ago). These populations expanded south of the Laurentide Ice Sheet and spread rapidly southward, occupying both North and South America by 12,000 to 14,000 years ago. The earliest populations in the Americas, before roughly 10,000 years ago, are known as Paleo-Indians . Indigenous peoples of the Americas have been linked to Siberian populations by proposed linguistic factors , the distribution of blood types , and in genetic composition as reflected by molecular data, such as DNA .

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84-616: While there is general agreement that the Americas were first settled from Asia, the pattern of migration and the place(s) of origin in Eurasia of the peoples who migrated to the Americas remain unclear. The traditional theory is that Ancient Beringians moved when sea levels were significantly lowered due to the Quaternary glaciation , following herds of now-extinct Pleistocene megafauna along ice-free corridors that stretched between

168-615: A Paleo-Siberian ( Early Paleo-Eskimo ) lineage before about 2,500 years ago. The discovery was made from archaeogenetic analyses on the remains of two female infants discovered in 2013 at the Upward Sun River site (USR). The USR site is affiliated with the Denali Complex , a dispersed archaeological culture of the American Arctic. The genomic analysis of nuclear DNA of the older of the two infants (USR1)

252-824: A Native American source population related to the Ainu ancestors, the Jōmon . Paleo-Indian skeletons in the Americas such as Kennewick Man (Washington State), Hoya Negro skeleton (Yucatán), Luzia Woman and other skulls from the Lagoa Santa site (Brazil), Buhl Woman (Idaho), Peñon Woman III , two skulls from the Tlapacoya site (Mexico City), and 33 skulls from Baja California have exhibited certain craniofacial traits distinct from most modern Native Americans, leading physical anthropologists to posit an earlier "Paleoamerican" population wave. The most basic measured distinguishing trait

336-676: A branch of Ancient East Asians migrated to Northeastern Siberia, and mixed with descendants of the ANE, leading to the emergence of Ancient Paleo-Siberian and Native American populations in Extreme Northeastern Asia. However, the Beringian standstill hypothesis is not supported by paternal DNA evidence, which may reflect different population histories for paternal and maternal lineages in Native Americans, which

420-614: A daughter population of ancient East Asians, who they encountered around 25,000 years ago, which led to the emergence of Native American ancestral populations. However, the exact location where the admixture took place is unknown, and the migratory movements that united the two populations are a matter of debate. One theory supposes that Ancient North Eurasians migrated south to East Asia , or Southern Siberia , where they would have encountered and mixed with ancient East Asians. Genetic evidence from Lake Baikal in Russia supports this area as

504-749: A durable and extensive geographic feature connecting Siberia with Alaska. With the rise of sea level after the Last Glacial Maximum (LGM), the Beringian land bridge was again submerged. Estimates of the final re-submergence of the Beringian land bridge based purely on present bathymetry of the Bering Strait and eustatic sea level curve place the event around 11,000 years BP (Figure 1). Ongoing research reconstructing Beringian paleogeography during deglaciation could change that estimate and possible earlier submergence could further constrain models of human migration into North America. The onset of

588-619: A high-rise central business district . The satellite cities are primarily suburban , featuring a small downtown core and a small industrial area or port . The suburbs consist mostly of residences, strip malls , and shopping centers. The region is also home to numerous ports. The two largest and busiest are the Port of Seattle and Port of Tacoma , which, if combined, comprise the third largest container port in North America after Los Angeles/Long Beach and New York/New Jersey. As defined by

672-478: A lineage found among Native Americans and Han Chinese, emerged around 20,000 BP, constraining the emergence of D4h3 to post-LGM. Age estimates based on Y-chromosome micro-satellite diversity place origin of the American Haplogroup Q1a3a (Y-DNA) at around 15,000 to 10,000 BP. Greater consistency of DNA molecular evolution rate models with each other and with archaeological data may be gained by

756-400: A lure for coastal migration. Reconstruction of the southern Beringian coastline also suggests potential for a highly productive coastal marine environment. Pollen data indicate a warm period culminating between 17,000 and 13,000 BP followed by cooling between 13,000 and 11,500 BP. Coastal areas deglaciated rapidly as coastal alpine glaciers, then lobes of Cordilleran ice, retreated. The retreat

840-495: A much earlier date, possibly 40,000 years ago, followed by a much later second wave of immigrants. The Clovis First theory, which dominated thinking on New World anthropology for much of the 20th century, was challenged in the 2000s by the secure dating of archaeological sites in the Americas to before 13,000 years ago. The archaeological sites in the Americas with the oldest dates that have gained broad acceptance are all compatible with an age of about 15,000 years. This includes

924-586: A small Australasian ancestry component of c. 3% was detected, which remains unexplained by the current state of research (as of 2021), but may be explained by the presence of the more basal Tianyuan-related ancestry , a deep East Asian lineage which did not directly contribute to modern East Asians but may have contributed to the ancestors of Native Americans in Siberia, as such ancestry is also found among previous Paleolithic Siberians ( Ancient North Eurasians ). Ancient Beringian The Ancient Beringian (AB)

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1008-510: Is a human archaeogenetic lineage, based on the genome of an infant found at the Upward Sun River site (dubbed USR1), dated to 11,500 years ago. The AB lineage diverged from the Ancestral Native American (ANA) lineage about 20,000 years ago. The ANA lineage was estimated as having been formed between 20,000 and 25,000 years ago by a mixture of East Asian (~65%) and Ancient North Eurasian (~35%) lineages, consistent with

1092-503: Is a long-standing open question. While advances in archaeology , Pleistocene geology , physical anthropology , and DNA analysis have progressively shed more light on the subject, significant questions remain unresolved. The " Clovis first theory" refers to the hypothesis that the Clovis culture represents the earliest human presence in the Americas about 13,000 years ago. Evidence of pre-Clovis cultures has accumulated and pushed back

1176-654: Is due to the Aleutian Low coming in from the northwest, and variation in winter rain from year to year is mostly due to variation in the strength of the Aleutian Low (strong = dry and weak = wet). The dry season (May-Sep) is caused by a subtropical high moving north from California. The driest part of the region is in the rain shadow of the Olympic Mountains around Sequim and Whidbey Island , receiving 40-75 cm of precipitation per year. The wettest part

1260-824: Is in the foothills of both the Cascade and the Olympic Mts, and on the west side of the Sound, receiving 125-1000+ cm of precipitation per year. North Pacific Oak Woodland is one of the principal plant associations of the Puget Trough, where many of the soils are well drained mesic. Counties of the Puget Sound region: In addition, the San Juan Islands (all of San Juan County plus a few islands belonging to Whatcom County ) are often considered part of

1344-536: Is less than 100%. The HTLV virus genome has been mapped, allowing identification of four major strains and analysis of their antiquity through mutations. The highest geographic concentrations of the strain HLTV-1 are in sub-Saharan Africa and Japan. In Japan, it occurs in its highest concentration on Kyushu . It is also present among African descendants and native populations in the Caribbean region and South America. It

1428-623: Is not uncommon and has been observed in other populations. A 2019 study suggested that Native Americans are the closest living relatives to 10,000-year-old fossils found near the Kolyma River in northeastern Siberia. A study published in July 2022 suggested that people in southern China may have contributed to the Native American gene pool, based on the discovery and DNA analysis of 14,000-year-old human fossils. The contrast between

1512-511: Is now Western Canada , would have allowed migration before the beginning of the Holocene . However, a 2016 study has argued against this, suggesting that the peopling of North America via such a corridor is unlikely to significantly pre-date the earliest Clovis sites. The study concludes that the ice-free corridor in what is now Alberta and British Columbia "was gradually taken over by a boreal forest dominated by spruce and pine trees" and that

1596-803: Is rare in Central America and North America. Its distribution in the Americas has been regarded as due to importation with the slave trade. The Ainu have developed antibodies to HTLV-1, indicating its endemicity to the Ainu and its antiquity in Japan. A subtype "A" has been defined and identified among the Japanese (including Ainu ), and among Caribbean and South American isolates. A subtype "B" has been identified in Japan and India. In 1995, Native Americans in coastal British Columbia were found to have both subtypes A and B. Bone marrow specimens from an Andean mummy about 1500 years old were reported to have shown

1680-536: Is the dolichocephaly of the skull. Some modern isolated populations such as the Pericúes of Baja California and the Fuegians of Tierra del Fuego exhibit that same morphological trait. Other anthropologists advocate an alternative hypothesis that evolution of an original Beringian phenotype gave rise to a distinct morphology that was similar in all known Paleoamerican skulls, followed by later convergence towards

1764-607: The Alaskan Peninsula isolated the interior of Beringia from the Pacific coast. Coastal alpine glaciers and lobes of Cordilleran ice coalesced into piedmont glaciers that covered large stretches of the coastline as far south as Vancouver Island and formed an ice lobe across the Straits of Juan de Fuca by 18,000 BP. Coastal alpine glaciers started to retreat around 19,000 BP while Cordilleran ice continued advancing in

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1848-495: The Alexander Archipelago . The now-submerged coastal plain has potential for more refugia. Pollen data indicate mostly herb/shrub tundra vegetation in unglaciated areas, with some boreal forest towards the southern end of the range of Cordilleran ice. The coastal marine environment remained productive, as indicated by fossils of pinnipeds . The highly productive kelp forests over rocky marine shallows may have been

1932-519: The Boeing Company became established in the area. During World War II the Puget Sound area became a focus for the war industry, with Boeing producing many of the nation's heavy bombers and the ports of Seattle, Bremerton and Tacoma available for shipbuilding. The most important yards in the Sound during World War II were Seattle-Tacoma Shipbuilding 's Seattle and Tacoma yards, also known as Todd Pacific , Todd Seattle and Todd Tacoma and

2016-809: The Buttermilk Creek Complex in Texas, the Meadowcroft Rockshelter site in Pennsylvania and the Monte Verde site in southern Chile. Archaeological evidence of pre- Clovis people points to the South Carolina Topper Site being 16,000 years old, at a time when the glacial maximum would have theoretically allowed for lower coastlines. It has often been suggested that an ice-free corridor, in what

2100-458: The Laurentide and Cordilleran ice sheets. Another route proposed is that, either on foot or using boats , they migrated down the Pacific coast to South America as far as Chile . Any archaeological evidence of coastal occupation during the last Ice Age would now have been covered by the sea level rise , up to a hundred metres since then. The precise date for the peopling of the Americas

2184-543: The North American extinction event that occurred at the end of the Pleistocene. Their genome , however, contains evidence of a bottleneck – something that can be used to test hypothesis on migrations between the two continents. Early human groups were largely nomadic , relying on following food sources for survival. Mobility was part of what made humans successful. As nomadic groups, early humans likely followed

2268-645: The Oregon Country , and became U.S. territory when the 1846 Oregon Treaty was signed. After arriving along the Oregon Trail , many settlers wandered north to what is now Washington and settled the Puget Sound area. The first non-indigenous settlement was New Market (now known as Tumwater ) in 1846. In 1853 Washington Territory was formed from part of Oregon Territory . In 1888 the Northern Pacific railroad line reached Puget Sound, linking

2352-887: The Puget Sound Navy Yard . They produced a significant portion of destroyers and escort carriers . Smaller operations included Winslow , Associated Shipbuilders and the Lake Washington Shipyard . Since 1995, Puget Sound has been recognized as an American Viticultural Area by the Alcohol and Tobacco Tax and Trade Bureau . The urban region designated the Puget Sound Region is centered on Seattle and consists of nine counties , two urban center cities and four satellite cities making up what has been dubbed "Pugetopolis." Both urban core cities have large industrial areas and seaports plus

2436-560: The Puget lowlands up to 16,800 BP. Even during the maximum extent of coastal ice, unglaciated refugia persisted on present-day islands, that supported terrestrial and marine mammals. As deglaciation occurred, refugia expanded until the coast became ice-free by 15,000 BP. The retreat of glaciers on the Alaskan Peninsula provided access from Beringia to the Pacific coast by around 17,000 BP. The ice barrier between interior Alaska and

2520-623: The Sea of Okhotsk , shows a marked shift from tree and shrub pollen to herb pollen prior to 30,000 BP, as herb tundra replaced boreal forest and shrub steppe going into the LGM. A similar record of tree/shrub pollen being replaced with herb pollen as the LGM approached was recovered near the Kolyma River in Arctic Siberia. The abandonment of the northern regions of Siberia due to rapid cooling or

2604-518: The Seattle–Tacoma combined statistical area (CSA), which encompasses most of the Puget Sound region. The population of this wider region is 4,269,349—almost two-thirds of Washington's population—as of 2012 . The Seattle CSA is the 12th largest CSA , and the 13th largest primary census statistical area in the country. The additional metropolitan and micropolitan areas included are: A state-run ferry system, Washington State Ferries , connects

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2688-592: The U.S. Census Bureau and the Office of Management and Budget , the Seattle metropolitan area is officially the Seattle–Tacoma–Bellevue, WA metropolitan statistical area (MSA) and consists of: Based on commuting patterns, the adjacent metropolitan areas of Olympia , Bremerton , and Mount Vernon , along with a few smaller satellite urban areas, are grouped together in a wider labor market region known as

2772-680: The Ulchis of the lower Amur River region (4 among 87 sampled, or 4.6%), along with Subhaplogroup C1a (1 among 87, or 1.1%). Subhaplogroup C1a is regarded as a close sister clade of the Native American Subhaplogroup C1b. Subhaplogroup D1a has also been found among ancient Jōmon skeletons from Hokkaido The modern Ainu are regarded as descendants of the Jōmon. The occurrence of the Subhaplogroups D1a and C1a in

2856-647: The Vashon Glaciation , the Puget Sound region was covered by a lobe of the Cordilleran Ice Sheet . The glacier that covered the area was about 3,000 feet (900 m) thick within the vicinity of Seattle. By the time Captain George Vancouver found the Sound in 1792, early native people had already been there for over 5,000 years. Logging started as early as 1853. In the 1880s logging railroads cut their way into Puget Sound. 1886

2940-761: The Wisconsin glaciation , the Earth's ocean water was, to varying degrees over time, stored in glacier ice. As water accumulated in glaciers, the volume of water in the oceans correspondingly decreased, resulting in lowering of global sea level . The variation of sea level over time has been reconstructed using oxygen isotope analysis of deep sea cores, the dating of marine terraces, and high-resolution oxygen isotope sampling from ocean basins and modern ice caps. A drop of eustatic sea level by about 60 to 120 metres (200 to 390 ft) from present-day levels, commencing around 30,000 years Before Present (BP), created Beringia ,

3024-506: The "Clovis people likely came from the south, not the north, perhaps following wild animals such as bison ". An alternative hypothesis for the peopling of America is coastal migration , which may have been feasible along the deglaciated (but now submerged) coastline of the Pacific Northwest from about 16,000 years ago. Pre-LGM migration across Beringia has been proposed to explain purported pre-LGM ages of archaeological sites in

3108-464: The Americas followed by isolation of the northern population following closure of the ice-free corridor. Evidence of Australo-Melanesians admixture in Amazonian populations was found by Skoglund and Reich (2016). A study of the diversification of mtDNA Haplogroups C and D from southern Siberia and eastern Asia, respectively, suggests that the parent lineage (Subhaplogroup D4h) of Subhaplogroup D4h3,

3192-462: The Americas occurring around 10,000 to 15,000 years after isolation of the small founding population . Another model (Kitchen et al. 2008) proposes that migration into Beringia occurred approximately 36,000 BP, followed by 20,000 years of isolation in Beringia. A third model (Nomatto et al. 2009) proposes that migration into Beringia occurred between 40,000 and 30,000 BP, with a pre-LGM migration into

3276-831: The Americas such as Bluefish Caves and Old Crow Flats in the Yukon Territory , and Meadowcroft Rock Shelter in Pennsylvania. The oldest archaeological sites on the Alaskan side of Beringia date to around 14,000 BP. It is possible that a small founder population had entered Beringia before that time. However, archaeological sites that date closer to the LGM on either the Siberian or the Alaskan side of Beringia are lacking. Biomarker and microfossil analyses of sediments from Lake E5 and Burial Lake in northern Alaska suggest human presence in eastern Beringia as early as 34,000 years ago. These sedimentary analyses have been suggested to be

3360-632: The LGM, due to cold and dry conditions. Coastal environments during the Last Glacial Maximum were complex. The lowered sea level, and an isostatic bulge equilibrated with the depression beneath the Cordilleran Ice Sheet, exposed the continental shelf to form a coastal plain. While much of the coastal plain was covered with piedmont glaciers, unglaciated refugia supporting terrestrial mammals have been identified on Haida Gwaii , Prince of Wales Island , and outer islands of

3444-544: The Last Glacial Maximum after 30,000 years BP saw the expansion of alpine glaciers and continental ice sheets that blocked migration routes out of Beringia. By 21,000 years BP, and possibly thousands of years earlier, the Cordilleran and Laurentide ice sheets coalesced east of the Rocky Mountains , closing off a potential migration route into the center of North America. Alpine glaciers in the coastal ranges and

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3528-563: The Last Glacial Maximum, climates in eastern Siberia fluctuated between conditions approximating present day conditions and colder periods. The pre-LGM warm cycles in Arctic Siberia saw flourishes of megafaunas. The oxygen isotope record from the Greenland Ice Cap suggests that these cycles after about 45,000 BP lasted anywhere from hundreds to between one and two thousand years, with greater duration of cold periods starting around 32,000 BP. The pollen record from Elikchan Lake, north of

3612-557: The Pacific coast broke up starting around 16,200 BP. The ice-free corridor to the interior of North America opened between 13,000 and 12,000 BP. Glaciation in eastern Siberia during the LGM was limited to alpine and valley glaciers in mountain ranges and did not block access between Siberia and Beringia. The paleoclimates and vegetation of eastern Siberia and Alaska during the Wisconsin glaciation have been deduced from high resolution oxygen isotope data and pollen stratigraphy . Prior to

3696-680: The Puget Sound region "Ish River country", owing to its numerous rivers with names ending in "ish", such as the Duwamish , Samish , Sammamish , Skokomish , Skykomish , Snohomish , and the Stillaguamish . The ish ending is from Salishan languages and means "people of". The Puget Sound region was formed by the collision and attachment of many terranes ("microcontinents") to the North American Plate between about 50 to 10 million years ago. About 15,000 years ago during

3780-447: The St. Helens fire burned 300,000 acres (1,200 km ). Mount Rainier National Park started in 1899. The 1902 Yacolt Burn burned 600,000 acres (2,400 km ). Olympic National Park was established in 1938. George Vancouver explored Puget sound in 1792. Vancouver claimed it for Great Britain on June 4, 1792, naming it for one of his officers, Lieutenant Peter Puget . It became part of

3864-522: The chronology of the controversial Pedra Furada rock shelter in Piauí , Brazil . More recently, studies at the archaeological sites Santa Elina (27000-10000 years BP) in the midwest, and Rincão I (20000-12000 years BP) in southeastern Brazil also show associations of evidence of human presence with sediments dating from before the LGM. A 2003 study dated evidence for the controlled use of fire to before 40,000 years ago. Additional evidence has been adduced from

3948-483: The cooling period that led into the LGM. A compilation of archaeological site dates throughout eastern Siberia suggest that the cooling period caused a retreat of humans southwards. Pre-LGM lithic evidence in Siberia indicate a settled lifestyle that was based on local resources, while post-LGM lithic evidence indicate a more migratory lifestyle. A 2021 discovery of human footprints in relict lake sediments near White Sands National Park in New Mexico demonstrated there

4032-571: The deglaciated landscape increased slowly. The earliest possible viability of the ice-free corridor as a human migration route has been estimated at 11,500 BP. Birch forests were advancing across former herb tundra in Beringia by 17,000 BP in response to climatic amelioration, indicating increased productivity of the landscape. Analyses of biomarkers and microfossils preserved in sediments from Lake E5 and Burial Lake in northern Alaska suggest early humans burned Beringian landscapes as early as 34,000 years ago. The authors of these studies suggest that fire

4116-560: The development of human Y-chromosome DNA haplogroups (yDNA haplogroups ) and human mitochondrial DNA haplogroups (mtDNA haplogroups) characteristic of Native American populations. Models of molecular evolution rates were used to estimate the ages at which Native American DNA lineages branched off from their parent lineages in Asia and to deduce the ages of demographic events. One model (Tammetal 2007) based on Native American mtDNA Haplotypes (Figure 2) proposes that migration into Beringia occurred between 30,000 and 25,000 BP, with migration into

4200-508: The dispersal associated with the early peopling of the Americas . Puget Sound region The Puget Sound region is a coastal area of the Pacific Northwest in the U.S. state of Washington , including Puget Sound , the Puget Sound lowlands, and the surrounding region roughly west of the Cascade Range and east of the Olympic Mountains . It is characterized by a complex array of saltwater bays, islands, and peninsulas carved out by prehistoric glaciers. Poet Robert Sund called

4284-468: The early 21st century, the models of the chronology of migration are divided into two general approaches. The first is the short chronology theory , that the first migration occurred after the LGM, which went into decline after about 19,000 years ago, and was then followed by successive waves of immigrants. The second theory is the long chronology theory, which proposes that the first group of people entered Beringia, including ice-free parts of Alaska, at

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4368-454: The early human groups who hunted them. Bison , a type of megafauna, have been identified as an ideal candidate for the tracing of human migrations out of Europe because of both their abundance in North America as well as being one of the first megafauna for which ancient DNA was used to trace patterns of population movement. Unlike other types of fauna that moved between the Americas and Eurasia ( mammoths , horses , and lions ), Bison survived

4452-423: The five mtDNA haplogroups found in Indigenous Americans. Native Americans mostly belong to the X2a clade, which has never been found in the Old World . According to Jennifer Raff , X2a probably originated in the same Siberian population as the other four founding maternal lineages, and that there is no compelling reason to believe it is related to X lineages found in Europe or West Eurasia. The Kennewick man fossil

4536-400: The food from Eurasia to the Americas – part of the reason why tracing megafaunal DNA is so helpful for garnering insight to these migratory patterns. The grey wolf originated in the Americas and migrated into Eurasia prior to the Last Glacial Maximum – during which it was believed that remaining populations of the grey wolf residing in North America faced extinction and were isolated from

4620-513: The genetic profiles of the Hokkaido Jōmon skeletons and the modern Ainu illustrates another uncertainty in source models derived from modern DNA samples. The development of high-resolution genomic analysis has provided opportunities to further define Native American subclades and narrow the range of Asian subclades that may be parent or sister subclades. The common occurrence of the mtDNA Haplogroups A, B, C, and D among eastern Asian and Native American populations has long been recognized, along with

4704-569: The geologic association of bones at the Bluefish Cave and Old Crow Flats sites, and the related Bonnet Plume site, have been called into question. No evidence of human remains have been discovered at these sites. In addition to disputed archaeological sites, support for pre-LGM human presence has been found in lake sediment records of northern Alaska. Biomarker and microfossil analyses of sediments from Lake E5 and Burial Lake in suggest human presence in eastern Beringia as early as 34,000 years ago. These analyses are indeed compelling in that they corroborate

4788-555: The ice sheets, the oldest such sites occur in association with the Clovis complex. If humans managed to breach the continental ice sheets significantly before 13,000 BP, there should be clear evidence for it in the form of at least some stratigraphically discrete archaeological components with a relatively high artifact count. So far, no such evidence exists." Genetic studies have used high resolution analytical techniques applied to DNA samples from modern Native Americans and Asian populations regarded as their source populations to reconstruct

4872-558: The inferences made from the Bluefish Cave and Old Crow Flats sites. In 2020, evidence emerged for a new pre-LGM site in North-Central Mexico . Chiquihuite cave , an archaeological site in Zacatecas State, has been dated to 26,000 years BP based on numerous lithic artefacts discovered there. However, there is scholarly debate over whether the artifacts should be considered evidence of human activity or if they were formed naturally. No evidence of human DNA or hearth have been unearthed. Pre-LGM human presence in South America rests partly on

4956-420: The larger islands to the Washington mainland, as well as both sides of the sound, allowing cars and people to move about the greater Puget Sound region. The region has a Csb ( warm summer mediterranean ) climate with some areas in the far east (western foothills of the Cascades) having an oceanic ( Cfb ) climate if the 30 millimetres (1.2 in) isohyet is used. The wet season is from October to April and

5040-399: The location where the admixture took place. However, a third theory, the "Beringian standstill hypothesis", suggests that East Asians instead migrated north to Northeastern Siberia, where they mixed with ANE, and later diverged in Beringia, where distinct Native American lineages formed. This theory is supported by maternal and nuclear DNA evidence. According to Grebenyuk, after 20,000 BP,

5124-462: The lower Amur region suggests a source population from that region distinct from the Altai-Baikal source populations, where sampling did not reveal those two particular subclades. The conclusions regarding Subhaplogroup D1 indicating potential source populations in the lower Amur and Hokkaido areas stand in contrast to the single-source migration model. Subhaplogroup D4h3 has been identified among Han Chinese . Subhaplogroup D4h3 from China does not have

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5208-451: The model of the peopling of the Americas via Beringia during the Last Glacial Maximum . The Ancient Beringian lineage is extinct, and is not found as a contribution to modern indigenous lineages in Alaska . The 2018 study suggests that the AB lineage was replaced by or absorbed in a back-migration of "North Native American" (NNA) to Alaska. The modern Athabaskan populations are derived from an admixture of this NNA back-migration and

5292-407: The modern Native American phenotype. Archaeogenetic studies do not support a two-wave model or the Paleoamerican hypothesis of an Australo-Melanesian origin, and firmly assign all Paleo-Indians and modern Native Americans to one ancient population that entered the Americas in a single migration from Beringia. Only in one ancient specimen (Lagoa Santa) and a few modern populations in the Amazon region,

5376-411: The morphology of Luzia Woman fossil, which was described as Australo-Melanesian . This interpretation was challenged in a 2003 review which concluded the features in question could also have arisen by genetic drift. In November 2018, scientists of the University of São Paulo and Harvard University released a study that contradicts the alleged Australo-Melanesian origin of Luzia. Using DNA sequencing,

5460-425: The most closely related subclades grow more specific. Subhaplogroups D1 and D4h3 have been regarded as Native American specific based on their absence among a large sampling of populations regarded as potential descendants of source populations, over a wide area of Asia. Among the 3,764 samples, the Sakhalin –lower Amur region was represented by 61 Oroks . In another study, Subhaplogroup D1a has been identified among

5544-407: The only possibly recoverable remnants of humans living in Alaska during the last Glacial period. At Old Crow Flats, mammoth bones have been found that are broken in distinctive ways indicating human butchery. The radiocarbon dates on these vary between 25,000 and 40,000 BP. Also, stone microflakes have been found in the area indicating tool production. However, the interpretations of butcher marks and

5628-436: The period of the Last Glacial Maximum along with genetic evidence found from early human remains in the Americas provides evidence to support pre-Clovis migrations into the Americas. The Native American source population was formed in Siberia by the mixing of two distinct populations: Ancient North Eurasians and an ancient East Asian (ESEA) population. According to Jennifer Raff, the Ancient North Eurasian population mixed with

5712-402: The possible date of the first peopling of the Americas. Academics generally believe that humans reached North America south of the Laurentide Ice Sheet at some point between 15,000 and 20,000 years ago. Some new controversial archaeological evidence suggests the possibility that human arrival in the Americas may have occurred prior to the Last Glacial Maximum more than 20,000 years ago. During

5796-448: The presence of haplogroup X . As a whole, the greatest frequency of the four Native American associated haplogroups occurs in the Altai - Baikal region of southern Siberia. Some subclades of C and D closer to the Native American subclades occur among Mongolian, Amur, Japanese, Korean, and Ainu populations. With further definition of subclades related to Native American populations, the requirements for sampling Asian populations to find

5880-405: The presence of the A subtype. The finding ignited controversy, with contention that the sample DNA was insufficiently complete for the conclusion and that the result reflected modern contamination. However, a re-analysis indicated that the DNA sequences were consistent with, but not definitely from, the "cosmopolitan clade" (subtype A). The presence of subtypes A and B in the Americas is suggestive of

5964-416: The rate grew more rapid. The inland Cordilleran and Laurentide ice sheets retreated more slowly than did the coastal glaciers. Opening of an ice-free corridor did not occur until after 13,000 to 12,000 BP. The early environment of the ice-free corridor was dominated by glacial outwash and meltwater, with ice-dammed lakes and periodic flooding from the release of ice-dammed meltwater. Biological productivity of

6048-410: The region to eastern states. For a long period Tacoma was noted for its large smelters where gold, silver, copper and lead ores were treated. Seattle was the primary port for trade with Alaska and the rest of the country and for a time possessed a large shipbuilding industry. The region around eastern Puget Sound developed heavy industry during the period including World War I and World War II , and

6132-490: The rest of the population. This, however, may not be the case. Radiocarbon dating of ancient grey wolf remains found in permafrost deposits in Alaska show a continuous exchange of population from 12,500 radiocarbon years BP to beyond radiocarbon dating capabilities. This indicates that there was viable passage for grey wolf populations to exchange between the two continents. These faunas' ability to exchange populations during

6216-578: The results showed that Luzia's ancestry was entirely Native American. Stones described as probable tools, hammerstones and anvils , have been found in southern California , at the Cerutti Mastodon site , that are associated with a mastodon skeleton which appeared to have been processed by humans. The mastodon skeleton was dated by thorium-230/uranium radiometric analysis, using diffusion–adsorption–decay dating models, to around 130 thousand years ago. No human bones were found and expert reaction

6300-484: The retreat of game species with the onset of the LGM has been proposed to explain the lack of archaeological sites in that region dating to the LGM. The pollen record from the Alaskan side shows shifts between herb/shrub and shrub tundra prior to the LGM, suggesting less dramatic warming episodes than those that allowed forest colonization on the Siberian side. Diverse, though not necessarily plentiful, megafauna were present in those environments. Herb tundra dominated during

6384-464: The same geographic implication as Subhaplotype D1a from Amur-Hokkaido, so its implications for source models are more speculative. Its parent lineage, Subhaplotype D4h, is believed to have emerged in East Asia, rather than Siberia, around 20,000 BP. Subhaplogroup D4h2, a sister clade of D4h3, has also been found among Jōmon skeletons from Hokkaido. D4h3 has a coastal trace in the Americas. X is one of

6468-403: The use of dated fossil DNA to calibrate molecular evolution rates. Although there is no archaeological evidence that can be used to direct support a coastal migration route during the Last Glacial Maximum , genetic analysis has been used to support this thesis. In addition to human genetic lineage, megafaunal DNA lineage can be used to trace movements of megafauna – large mammalian – as well as

6552-665: Was a verifiable human presence in the region dating back to the LGM between 18,000 and 26,000 years ago. Later studies, reported in October 2023, confirmed that the age of the human footprints to be "up to 23,000 years old". The Clovis-first advocates have not accepted the veracity of these findings. In 2022, they said, "The oldest evidence for archaeological sites in the New World with large numbers of artifacts occurring in discrete and minimally disturbed stratigraphic contexts occur in eastern Beringia between 13,000 and 14,200 BP. South of

6636-413: Was accelerated as sea levels rose and floated glacial termini. It has been estimated that the coast range was fully ice-free between 16,000 and 15,000 BP. Littoral marine organisms colonized shorelines as ocean water replaced glacial meltwater. Replacement of herb/shrub tundra by coniferous forests was underway by 15,000 BP north of Haida Gwaii. Eustatic sea level rise caused flooding, which accelerated as

6720-727: Was done at the Centre for Geogenetics at the University of Copenhagen 's Natural History Museum of Denmark . Results from the team's genetic analysis were published in January 2018 in the scientific journal Nature . The analysis compared the infant's genomes with both ancient and contemporary genomes. The results suggested that the pre-"Ancestral Native American" lineage derived from the East Asian lineage after 36 kya, with gene flow until about 25 kya. During 25–20 kya, this lineage

6804-477: Was found to carry the deepest branch of the X2a haplogroup, and he did not have any European ancestry that would be expected for a European origin of the lineage. The Human T cell Lymphotrophic Virus 1 ( HTLV-1 ) is a virus transmitted through exchange of bodily fluids and from mother to child through breast milk. The mother-to-child transmission mimics a hereditary trait, although such transmission from maternal carriers

6888-704: Was later substantially mixed with the Ancient North Eurasian lineage (~35%), who are described as the "result of a palaeolithic admixture" between ancient West Eurasians and ancient East Eurasians , to form the "Ancestral Native American" lineage by 20 kya. The "Ancient Beringian" (AB) lineage derived from ANA and persisted without significant admixture in Alaska until the time of USR1, some 8,000 years later. The lineage of other Paleo-Indians diverged from AB at ca. 20–18 kya, and further divided into "North Native American" (NNA) and "South Native American" lineages between 17.5 kya and 14.6 kya, reflecting

6972-471: Was mixed; claims of tools and bone processing were called "not plausible" by Prof. Tom Dillehay . The Yana River Rhino Horn site (RHS) has dated human occupation of eastern Arctic Siberia to 31,300 BP. That date has been interpreted by some as evidence that migration into Beringia was imminent, lending credence to occupation of Beringia during the LGM. However, the Yana RHS date is from the beginning of

7056-500: Was used as means of hunting megafauna. The Indigenous peoples of the Americas have an ascertained archaeological presence in the Americas dating back to about 15,000 years ago. More recent research, however, suggests a human presence dating to between 18,000 and 26,000 years ago, during the Last Glacial Maximum. There remain uncertainties regarding the precise dating of individual sites and regarding conclusions drawn from population genetics studies of contemporary Native Americans. In

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