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The Silurian ( / s ɪ ˈ lj ʊər i . ən , s aɪ -/ sih- LURE -ee-ən, sy- ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago ( Mya ), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods , the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.

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85-616: The Ontarian River is the term used for the pre-glacial river that began the creation of the valley in Silurian age shales and limestones now occupied by Lake Ontario . The valley was greatly deepened by glacial action during the Ice age . The original flow is thought to have been westward beginning from the Laurentian shield in the present area of the St. Lawrence River to eventually join

170-789: A kingdom in the Holy Roman Empire , and subsequently a part of the Habsburg monarchy and the Austrian Empire . After World War I and the establishment of an independent Czechoslovak state , the whole of Bohemia became a part of Czechoslovakia , defying claims of the German-speaking inhabitants that regions with German-speaking majority should be included in the Republic of German-Austria . Between 1938 and 1945, these border regions were annexed to Nazi Germany as

255-530: A Central European country to be canonized by Pope John Paul II before the " Velvet Revolution " later that year. After the Velvet Divorce in 1993, the territory of Bohemia remained in the Czech Republic. The new Constitution of the Czech Republic provided for higher administrative units to be established, providing for the possibility of Bohemia as an administrative unit, but did not specify

340-512: A German army in 1004, ending the rule of Bolesław I of Poland . The first to use the title of "King of Bohemia" were the Přemyslid dukes Vratislav II (1085) and Vladislaus II (1158), but their heirs would return to the title of duke . The title of king became hereditary under Ottokar I (1198). His grandson Ottokar II (king from 1253 to 1278) conquered a short-lived empire that contained modern Austria and Slovenia . The mid-13th century had

425-586: A Proto-Germanic *Bajahaimaz . Boiohaemum was apparently isolated to the area where King Marobod 's kingdom was centered, within the Hercynian forest . Byzantine Emperor Constantine VII in his 10th-century work De Administrando Imperio also mentioned the region as Boiki (see White Serbia ). The Czech name "Čechy" is derived from the name of the Slavic ethnic group , the Czechs , who settled in

510-630: A diverse range of epibionts, including certain hederelloids as aforementioned. Photosymbiotic scleractinians made their first appearance during the Middle Silurian. Reef abundance was patchy; sometimes, fossils are frequent, but at other points, are virtually absent from the rock record. Bohemia Bohemia ( / b oʊ ˈ h iː m i ə / boh- HEE -mee-ə ; Czech : Čechy [ˈtʃɛxɪ] ; German : Böhmen [ˈbøːmən] ; Upper Sorbian : Čěska [ˈtʃɪska] ; Silesian : Czechy )

595-411: A food web based on as-yet-undiscovered detritivores and grazers on micro-organisms. Millipedes from Cowie Formation such as Cowiedesmus and Pneumodesmus were considered as the oldest millipede from the middle Silurian at 428–430 million years ago, although the age of this formation is later reinterpreted to be from the early Devonian instead by some researchers. Regardless, Pneumodesmus

680-519: A high degree of development in relation to the age of its fossil remains. Fossils of this plant have been recorded in Australia, Canada, and China. Eohostimella heathana is an early, probably terrestrial, "plant" known from compression fossils of Early Silurian (Llandovery) age. The chemistry of its fossils is similar to that of fossilised vascular plants, rather than algae. Fossils that are considered as terrestrial animals are also known from

765-490: A large ocean occupied most of the northern half of the globe. The high sea levels of the Silurian and the relatively flat land (with few significant mountain belts) resulted in a number of island chains, and thus a rich diversity of environmental settings. During the Silurian, Gondwana continued a slow southward drift to high southern latitudes, but there is evidence that the Silurian icecaps were less extensive than those of

850-590: A liberal democratic republic, but serious issues emerged regarding the Czech majority's relationship with the native German and Hungarian minorities. Following the Munich Agreement in 1938, the border regions of Bohemia historically inhabited predominantly by ethnic Germans (the Sudetenland ) were annexed to Nazi Germany . The remnants of Bohemia and Moravia were then annexed by Germany in 1939, while

935-423: A minor mass extinction and associated with rapid sea-level change. Each one leaves a similar signature in the geological record, both geochemically and biologically; pelagic (free-swimming) organisms were particularly hard hit, as were brachiopods , corals , and trilobites , and extinctions rarely occur in a rapid series of fast bursts. The climate fluctuations are best explained by a sequence of glaciations, but

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1020-447: A new nation named and led by them with a united "slavic" ethnic consciousness. Bohemia was made a part of the early Slavic state of Great Moravia , under the rule of Svatopluk I (r. 870–894). After Svatopluk's death Great Moravia was weakened by years of internal conflict and constant warfare, ultimately collapsing and fragmenting because of the continual incursions of the invading nomadic Magyars . Bohemia's initial incorporation into

1105-465: A rebirth as romantic nationalism developed among the Czechs. In 1861, a new elected Bohemian Diet was established. The renewal of the old Bohemian Crown ( Kingdom of Bohemia , Margraviate of Moravia , and Duchy of Upper and Lower Silesia ) became the official political program of both Czech liberal politicians and the majority of Bohemian aristocracy ("state rights program"), while parties representing

1190-525: A second supercontinent known as Euramerica . When the proto-Europe collided with North America, the collision folded coastal sediments that had been accumulating since the Cambrian off the east coast of North America and the west coast of Europe. This event is the Caledonian orogeny , a spate of mountain building that stretched from New York State through conjoined Europe and Greenland to Norway. At

1275-520: A tripartite monarchy (Austria-Hungary-Bohemia) failed in 1871. The "state-rights program" remained the official platform of all Czech political parties (except for social democrats) until 1918. Under the state-rights program, appealing to the stability of Bohemia's borders over many centuries, the Czech emancipation movement claimed the right to the whole of the Bohemian lands over the Germans' right to

1360-561: Is still an important fossil as the oldest definitive evidence of spiracles to breath in the air. The first bony fish, the Osteichthyes , appeared, represented by the Acanthodians covered with bony scales. Fish reached considerable diversity and developed movable jaws , adapted from the supports of the front two or three gill arches. A diverse fauna of eurypterids (sea scorpions)—some of them several meters in length—prowled

1445-603: Is the westernmost and largest historical region of the Czech Republic . Bohemia can also refer to a wider area consisting of the historical Lands of the Bohemian Crown ruled by the Bohemian kings , including Moravia and Czech Silesia , in which case the smaller region is referred to as Bohemia proper as a means of distinction. Bohemia was a duchy of Great Moravia , later an independent principality,

1530-415: The Bohemian crown controlled such diverse lands as Moravia , Silesia , Upper Lusatia and Lower Lusatia , Brandenburg, an area around Nuremberg called New Bohemia, Luxembourg , and several small towns scattered around Germany. From the 13th century on, settlements of Germans developed throughout Bohemia, making Bohemia a bilingual country. The German settlers particularly brought mining technology to

1615-742: The Elbe - Danube watershed . In the second century BC, the Romans were competing for dominance in northern Italy with various peoples, including the Gauls -Celtic tribe Boii . The Romans defeated the Boii at the Battle of Placentia (194 BC) and the Battle of Mutina (193 BC) . Afterward, many of the Boii retreated north across the Alps. Much later Roman authors refer to the area they had once occupied (the "desert of

1700-655: The Mississippi River drainage system. During continental glaciation, the tremendous weight of the glacier depressed the area of the St. Lawrence so much that a lower point of drainage was established eastward through the present river system. The area is still rebounding, however, and the rise could further rearrange drainage within geologic time by raising the outlet of Lake Ontario sufficient for it to drain at some other, lower point. Laurentian River System (Ontario) Silurian One important event in this period

1785-624: The Potsdam Agreement . The Bohemian Germans' property was confiscated by the Czech authorities, and according to contemporary estimates, amounted to a third of the Czechoslovak national income. Germans who were valued for their skills were allowed to stay to pass on their knowledge to the Czech migrants. The expulsion severely depopulated the area and from then on, locales were referred to in only their Czech equivalents regardless of their previous demographic makeup. The resettlement of

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1870-580: The South Pole until they almost disappeared in the middle of Silurian. Layers of broken shells (called coquina ) provide strong evidence of a climate dominated by violent storms generated then as now by warm sea surfaces. The climate and carbon cycle appear to be rather unsettled during the Silurian, which had a higher frequency of isotopic excursions (indicative of climate fluctuations) than any other period. The Ireviken event , Mulde event , and Lau event each represent isotopic excursions following

1955-688: The Sudetenland . The remainder of Czech territory became the Second Czechoslovak Republic and was subsequently occupied as the Protectorate of Bohemia and Moravia until the end of World War II, after which Bohemia became part of the restored Czechoslovakia. In 1969, the Czech lands (including Bohemia) were given autonomy within Czechoslovakia as the Czech Socialist Republic . In 1990, the name

2040-404: The rector of Charles University and a prominent reformer and religious thinker, influenced the rise of modern Czech. During the ecumenical Council of Constance in 1415, Jan Hus was sentenced to be burnt at the stake as a heretic . The verdict was passed despite the fact that Hus was granted formal protection by Emperor Sigismund of Luxembourg prior to the journey. Hus was invited to attend

2125-615: The 1960 and 2000 reforms). However, the three lands are mentioned in the preamble of the Constitution of the Czech Republic : "We, citizens of the Czech Republic in Bohemia, Moravia and Silesia..." Bohemia had an area of 52,065 km (20,102 sq mi), and today is home to about 6.9 million of the Czech Republic's 10.9 million inhabitants. Bohemia was bordered in the south by Upper and Lower Austria (both in Austria ), in

2210-562: The Bohemian aristocracy, started a campaign for restoration of the kingdom's historic rights, whereby Czech was to regain its historical role and replace German as the language of administration. The enlightened absolutism of Joseph II and Leopold II , who introduced minor language concessions, showed promise for the Czech movement, but many of these reforms were later rescinded. During the Revolution of 1848 , many Czech nationalists called for autonomy for Bohemia from Habsburg Austria, but

2295-621: The Bohemians could defeat the Bohemians." Despite an apparent victory for the Catholics, the Bohemian Utraquists were still strong enough to negotiate freedom of religion in 1436. That happened in the so-called Compacts of Basel , declaring peace and freedom between Catholics and Utraquists. It lasted for only a short period of time, as Pope Pius II declared the compacts to be invalid in 1462. In 1458, George of Poděbrady

2380-539: The Boii" as Pliny and Strabo called it ) as Boiohaemum . The earliest mention was by Tacitus ' Germania 28 (written at the end of the first century AD), and later mentions of the same name are in Strabo and Velleius Paterculus . The name appears to consist of the tribal name Boio- plus the Proto-Germanic noun * haimaz "home" (whence Gothic haims , German Heim , Heimat , English home ), indicating

2465-663: The German minority and small part of the aristocracy proclaimed their loyalty to the centralist Constitution (so-called "Verfassungstreue"). After the defeat of Austria in the Austro-Prussian War in 1866, Hungarian politicians achieved the Austro-Hungarian Compromise of 1867 , ostensibly creating equality between the Austrian and Hungarian halves of the empire. An attempt by the Czechs to create

2550-800: The Helvetii and Boii left southern Germany and Bohemia a lightly inhabited "desert" into which Suebic peoples arrived, speaking Germanic languages, and became dominant over remaining Celtic groups. To the south, over the Danube, the Romans extended their empire, and to the southeast, in present-day Hungary, were Dacian peoples. In the area of modern Bohemia, the Marcomanni and other Suebic groups were led by their king, Marobodus , after they had suffered defeat to Roman forces in Germany. He took advantage of

2635-620: The Hussite armies, he used innovative tactics and weapons, such as howitzers, pistols, and fortified wagons, which were revolutionary for the time, and established Žižka as a great general who never lost a battle. After Žižka's death, Prokop the Great took over the command for the army, and under his lead the Hussites were victorious for another ten years, to the sheer terror of Europe. The Hussite cause gradually splintered into two main factions,

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2720-774: The Ilag IV camp for interned civilians from western Allied countries in the region. There were also 17 subcamps of the Flossenbürg concentration camp , in which both men and women, mostly Polish , Soviet and Jewish, but also French, Yugoslav, Czech, Romani and of several other ethnicities, were imprisoned and subjected to forced labor , and 16 subcamps of the Gross-Rosen concentration camp , in which men and women, mostly Polish and Jewish, but also Czechs, Russians, and other people, were similarly imprisoned and subjected to forced labor. Any open opposition to German occupation

2805-750: The Kingdom of the Marcomanni, concerning a queen named Fritigil , is from the fourth century, and she was thought to have lived in or near Pannonia. The Suebian Langobardi , who moved over many generations from the Baltic Sea , via the Elbe and Pannonia to Italy, recorded in a tribal history a time spent in "Bainaib". After the Migration Period , Bohemia was partially repopulated around the sixth century, and eventually Slavic tribes arrived from

2890-683: The Llandovery and Wenlock. Trilobites started to recover in the Rhuddanian, and they continued to enjoy success in the Silurian as they had in the Ordovician despite their reduction in clade diversity as a result of LOME. The Early Silurian was a chaotic time of turnover for crinoids as they rediversified after LOME. Members of Flexibilia, which were minimally impacted by LOME, took on an increasing ecological prominence in Silurian seas. Monobathrid camerates, like flexibles, diversified in

2975-490: The Llandovery, whereas cyathocrinids and dendrocrinids diversified later in the Silurian. Scyphocrinoid loboliths suddenly appeared in the terminal Silurian, shortly before the Silurian-Devonian boundary, and disappeared as abruptly as they appeared very shortly after their first appearance. Endobiotic symbionts were common in the corals and stromatoporoids. Rugose corals especially were colonised and encrusted by

3060-641: The Moravian Empire resulted in the extensive Christianization of the population . A native monarchy arose to the throne, and Bohemia came under the rule of the Přemyslid dynasty , which would rule the Czech lands for the next several hundred years. The Přemyslids secured their frontiers after the collapse of the Moravian state by entering into a state of semivassalage to the Frankish rulers . The alliance

3145-596: The Order in which the Older Sedimentary Strata Succeed each other in England and Wales, which was the germ of the modern geological time scale . As it was first identified, the "Silurian" series when traced farther afield quickly came to overlap Sedgwick's "Cambrian" sequence, however, provoking furious disagreements that ended the friendship. The English geologist Charles Lapworth resolved

3230-751: The Pope. After the death of King Louis II of Hungary and Bohemia in the Battle of Mohács in 1526, Archduke Ferdinand I of Austria became the new king of Bohemia, and the country became a constituent state of the Habsburg monarchy . From 1599 to 1711, Moravia (a Land of the Bohemian Crown ) was frequently subjected to raids by the Ottoman Empire and its vassals (especially the Tatars and Transylvania ). Overall, hundreds of thousands were enslaved whilst tens of thousands were killed. Bohemia enjoyed religious freedom between 1436 and 1620 and became one of

3315-475: The Silures show little correlation ( cf . Geologic map of Wales , Map of pre-Roman tribes of Wales ), Murchison conjectured that their territory included Caer Caradoc and Wenlock Edge exposures - and that if it did not there were plenty of Silurian rocks elsewhere 'to sanction the name proposed'. In 1835 the two men presented a joint paper, under the title On the Silurian and Cambrian Systems, Exhibiting

3400-486: The Silurian was the diversification of jawed fish , which include placoderms , acanthodians (which gave rise to cartilaginous fish ) and osteichthyan ( bony fish , further divided into lobe-finned and ray-finned fishes ), although this corresponded to sharp decline of jawless fish such as conodonts and ostracoderms . The Silurian system was first identified by the Scottish geologist Roderick Murchison , who

3485-518: The Silurian. The definitive oldest record of millipede ever known is Kampecaris obanensis and Archidesmus sp. from the late Silurian (425 million years ago) of Kerrera . There are also other millipedes, centipedes , and trigonotarbid arachnoids known from Ludlow (420 million years ago). Predatory invertebrates would indicate that simple food webs were in place that included non-predatory prey animals. Extrapolating back from Early Devonian biota, Andrew Jeram et al. in 1990 suggested

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3570-809: The Slovak lands became the separate Slovak Republic , a puppet state of Nazi Germany. From 1939 to 1945, Bohemia (without the Sudetenland), together with Moravia, formed the German Protectorate of Bohemia and Moravia . During World War II , the Germans operated the Theresienstadt Ghetto for Jews, the Dulag Luft Ost, Stalag IV-C and Stalag 359 prisoner-of-war camps for French , British, Belgian, Serbian, Dutch, Slovak, Soviet, Romanian, Italian and other Allied POWs, and

3655-650: The Tethys, the Proto-Tethys and Paleo-Tethys , the Rheic Ocean , the Iapetus Ocean (a narrow seaway between Avalonia and Laurentia), and the newly formed Ural Ocean . The Silurian period was once believed to have enjoyed relatively stable and warm temperatures, in contrast with the extreme glaciations of the Ordovician before it and the extreme heat of the ensuing Devonian; however, it is now known that

3740-482: The Winter Queen or Queen of Hearts, was the daughter of King James I of England and VI of Scotland . After Frederick's defeat in the Battle of White Mountain in 1620, 27 Bohemian estates leaders together with Jan Jesenius , rector of the Charles University of Prague, were executed on the Prague's Old Town Square on 21 June 1621, and the rest were exiled from the country; their lands were then given to Catholic loyalists (mostly of Bavarian and Saxon origin). That ended

3825-431: The area during the sixth or seventh century AD. Bohemia, like neighboring Bavaria , is named after the Boii , a large Celtic nation known to the Romans for their migrations and settlement in northern Italy and other places. Another part of the nation moved west with the Helvetii into southern France, which was one of the events leading to the interventions of Julius Caesar's Gaulish campaign of 58 BC. The emigration of

3910-412: The beginning of substantial German immigration, as the court sought to replace losses from the brief Mongol invasion of Europe in 1241. Germans settled primarily along the northern, western, and southern borders of Bohemia, although many lived in towns throughout the kingdom. The House of Luxembourg accepted the invitation to the Bohemian throne with the marriage to the Přemyslid heiress, Elizabeth and

3995-600: The cease-fire agreement, did not support Bohemian German self-defense, while the Czechoslovak army, having status of an Entente army, could freely operate. The absorption of the German-speaking areas in Czechoslovakia was hence a fait accompli . As a result, the totality of Bohemia (as the largest and most populous land) became the core of the newly formed country of Czechoslovakia , which combined Bohemia, Moravia , Czech Silesia , Upper Hungary (present-day Slovakia ) and Carpathian Ruthenia into one state. Under its first president, Tomáš Masaryk , Czechoslovakia became

4080-415: The communists and their Soviet protectors in many of the state's institutions. Gottwald and the communists responded with a coup d'état and installed a pro-Soviet authoritarian state. In 1949, Bohemia ceased to be an administrative unit of Czechoslovakia, as the country was divided into administrative regions that did not follow the historical borders. In 1989, Agnes of Bohemia became the first saint from

4165-444: The conflict by defining a new Ordovician system including the contested beds. An alternative name for the Silurian was "Gotlandian" after the strata of the Baltic island of Gotland . The French geologist Joachim Barrande , building on Murchison's work, used the term Silurian in a more comprehensive sense than was justified by subsequent knowledge. He divided the Silurian rocks of Bohemia into eight stages. His interpretation

4250-411: The council to defend himself and the Czech positions in the religious court, but with the emperor's approval, he was executed on 6 July 1415. The execution of Hus, as well as five consecutive papal crusades against followers of Hus, forced the Bohemians to defend themselves in the Hussite Wars . The uprising against imperial forces was led by a former mercenary, Jan Žižka of Trocnov. As the leader of

4335-441: The crowning subsequent of John I of Bohemia (in the Czech Republic known as Jan Lucemburský ) in 1310. His son, Charles IV , became King of Bohemia in 1346. He founded Charles University in Prague , Central Europe's first university, two years later. His reign brought Bohemia to its peak both politically and in total area, resulting in his being the first king of Bohemia to also be elected as Holy Roman Emperor . Under his rule,

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4420-481: The early Middle Ages , two new Suebic groupings appeared to the west of Bohemia in southern Germany, the Alemanni (in the Helvetian desert), and the Bavarians ( Baiuvarii ). Many Suebic tribes from the Bohemian region took part in such movements westwards, even settling as far away as Spain and Portugal. With them were also tribes who had pushed from the east, such as the Vandals , and Alans . Other groups pushed southwards towards Pannonia . The last known mention of

4505-427: The early 9th century, but became dominant only much later, in the 10th or 11th century. The 9th century was crucial for the future of Bohemia. The manorial system sharply declined, as it did in Bavaria. The influence of the central Fraganeo-Czechs grew, as a result of the important cultic center in their territory. They were Slavic-speaking, thus contributed to the transformation of diverse neighboring populations into

4590-401: The east, and their language began to replace the older Germanic, Celtic, and Sarmatian ones. These are precursors of today's Czechs, but the exact amount of Slavic immigration is a subject of debate. The Slavic influx was divided into two or three waves. The first wave came from the southeast and east, when the Germanic Lombards left Bohemia ( circa 568 AD). Soon after, from the 630s to 660s,

4675-408: The edge of the continental shelf) can be identified, and the highest Silurian sea level was probably around 140 metres (459 ft) higher than the lowest level reached. During this period, the Earth entered a warm greenhouse phase, supported by high CO 2 levels of 4500 ppm, and warm shallow seas covered much of the equatorial land masses. Early in the Silurian, glaciers retreated back into

4760-461: The end of the Silurian, sea levels dropped again, leaving telltale basins of evaporites extending from Michigan to West Virginia, and the new mountain ranges were rapidly eroded. The Teays River , flowing into the shallow mid-continental sea, eroded Ordovician Period strata, forming deposits of Silurian strata in northern Ohio and Indiana. The vast ocean of Panthalassa covered most of the northern hemisphere. Other minor oceans include two phases of

4845-448: The first deep-boring bivalves are known from this period. Chitons saw a peak in diversity during the middle of the Silurian. Hederelloids enjoyed significant success in the Silurian, with some developing symbioses with the colonial rugose coral Entelophyllum . The Silurian was a heyday for tentaculitoids , which experienced an evolutionary radiation focused mainly in Baltoscandia, along with an expansion of their geographic range in

4930-488: The first period to see megafossils of extensive terrestrial biota in the form of moss -like miniature forests along lakes and streams and networks of large, mycorrhizal nematophytes , heralding the beginning of the Silurian-Devonian Terrestrial Revolution. However, the land fauna did not have a major impact on the Earth until it diversified in the Devonian. The first fossil records of vascular plants , that is, land plants with tissues that carry water and food, appeared in

5015-416: The formerly German-settled areas allowed many poorer people to acquire property, thus "equalizing" Czechoslovak society. The Communist Party won the most votes in free elections, but not a simple majority . Klement Gottwald , the communist leader, became prime minister of a coalition government. In February 1948, the non-communist members of the government resigned in protest against arbitrary measures by

5100-451: The global climate underwent many drastic fluctuations throughout the Silurian, evidenced by numerous major carbon and oxygen isotope excursions during this geologic period. Sea levels rose from their Hirnantian low throughout the first half of the Silurian; they subsequently fell throughout the rest of the period, although smaller scale patterns are superimposed on this general trend; fifteen high-stands (periods when sea levels were above

5185-400: The lack of tillites in the middle to late Silurian make this explanation problematic. The Silurian period has been viewed by some palaeontologists as an extended recovery interval following the Late Ordovician mass extinction (LOME), which interrupted the cascading increase in biodiversity that had continuously gone on throughout the Cambrian and most of the Ordovician. The Silurian was

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5270-525: The lands, amounting to a third of Bohemia, where they formed the majority. After World War I , the German Bohemians demanded that the regions with German-speaking majority be included in a German state . Czech political leaders however claimed the entire Bohemian lands, including majority German-speaking areas, for Czechoslovakia. By the end of October, bilingual towns had been occupied by Czech forces. By end of November, many purely German-speaking towns had been occupied. German or Austrian troops, bound by

5355-402: The late-Ordovician glaciation. The southern continents remained united during this period. The melting of icecaps and glaciers contributed to a rise in sea level, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an unconformity . The continents of Avalonia , Baltica , and Laurentia drifted together near the equator , starting the formation of

5440-505: The mass extinction's aftermath, but expanded their range afterwards. The most abundant brachiopods were atrypids and pentamerides; atrypids were the first to recover and rediversify in the Rhuddanian after LOME, while pentameride recovery was delayed until the Aeronian. Bryozoans exhibited significant degrees of endemism to a particular shelf. They also developed symbiotic relationships with cnidarians and stromatolites. Many bivalve fossils have also been found in Silurian deposits, and

5525-416: The moderate Utraquists and the more fanatic Taborites . The Utraquists began to lay the groundwork for an agreement with the Catholic Church and found the more radical views of the Taborites distasteful. Additionally, with general war-weariness and yearning for order, the Utraquists were able to eventually defeat the Taborites in the Battle of Lipany in 1434. Sigismund said after the battle that "only

5610-438: The most liberal countries of the Christian world during that period. In 1609, Holy Roman Emperor Rudolf II , who made Prague again the capital of the empire at the time, himself a Roman Catholic, was moved by the Bohemian nobility to publish Maiestas Rudolphina , which confirmed the older Confessio Bohemica of 1575. After Emperor Matthias II and then King of Bohemia Ferdinand II (later Holy Roman Emperor) began oppressing

5695-409: The mountainous regions of the Sudetes . In the mining town of Sankt Joachimsthal (now Jáchymov ), famous coins called Joachimsthalers were coined, which gave their name to the thaler and the dollar . Meanwhile, Prague German intermediated between Upper German and East Central German , influencing the foundations of modern standard German. At the same time and place, the teachings of Jan Hus ,

5780-412: The natural defenses provided by its mountains and forests. They were able to maintain a strong alliance with neighboring tribes, including (at different times) the Lugii , Quadi , Hermunduri , Semnones , and Buri , which was sometimes partly controlled by the Roman Empire and sometimes in conflict with it; for example, in the second century, they fought Marcus Aurelius . In late classical times and

5865-424: The pro-reformation movement in Bohemia and also ended the role of Prague as ruling city of the Holy Roman Empire. In the so-called "renewed constitution" of 1627, German was established as a second official language in the Czech lands. Czech formally remained the first language in the kingdom, but both German and Latin were widely spoken among the ruling classes, although German became increasingly dominant, and Czech

5950-433: The region. In May 1945, Allied American , Polish, Czechoslovak , Soviet and Romanian troops captured the region, which was then restored to Czechoslovakia. After the war ended in 1945, after initial plans to cede lands to Germany or to create German-speaking cantons had been abandoned, the vast majority of the Bohemian Germans were expelled by the order of the re-established Czechoslovak central government, based on

6035-409: The revolutionaries were defeated. At the same time, German-speaking towns elected representatives for the first German Parliament at Frankfurt . Towns between Karlsbad and Reichenberg chose leftist representatives, while Eger , Rumburg , and Troppau elected conservative representatives. The old Bohemian Diet, one of the last remnants of the independence, was dissolved, although Czech experienced

6120-660: The rights of Protestants in Bohemia, the resulting Bohemian Revolt led to outbreak of the Thirty Years' War in 1618. Elector Frederick V of the Electorate of the Palatinate , a Calvinist Protestant, was elected by the Bohemian nobility to replace Ferdinand on the Bohemian throne and was known as the Winter King. Frederick's wife, the popular Elizabeth Stuart and subsequently, Elizabeth of Bohemia, known as

6205-495: The second half of the Silurian Period. The earliest-known representatives of this group are Cooksonia . Most of the sediments containing Cooksonia are marine in nature. Preferred habitats were likely along rivers and streams. Baragwanathia appears to be almost as old, dating to the early Ludlow (420 million years) and has branching stems and needle-like leaves of 10–20 centimetres (3.9–7.9 in). The plant shows

6290-516: The shallow Silurian seas and lakes of North America; many of their fossils have been found in New York state . Brachiopods were abundant and diverse, with the taxonomic composition, ecology, and biodiversity of Silurian brachiopods mirroring Ordovician ones. Brachiopods that survived the LOME developed novel adaptations for environmental stress, and they tended to be endemic to a single palaeoplate in

6375-606: The territory was taken by Samo 's tribal confederation. His death marked the end of the old "Slavonic" confederation, the second attempt to establish such a Slavonic union after Carantania in Carinthia . Other sources ( Descriptio civitatum et regionum ad septentrionalem plagam Danubii , Bavaria, 800–850) divide the population of Bohemia into the Merehani, Marharaii, Beheimare (Bohemani), and Fraganeo. (The suffix -ani or -ni means "people of-"). Christianity first appeared in

6460-837: The west by Bavaria (in Germany ), and in the north by Saxony and Lusatia (in Germany and Poland , respectively), in the northeast by Silesia (in Poland), and in the east by Moravia (also part of the Czech Republic). Bohemia's borders were mostly marked by mountain ranges such as the Bohemian Forest , the Ore Mountains , and the Giant Mountains ; the Bohemian-Moravian border roughly follows

6545-703: Was brutally suppressed by the Nazi authorities, and many Czech patriots were executed as a result. In 1942, the Czechoslovak resistance carried out the assassination of Reinhard Heydrich , and in reprisal German forces murdered the population of a whole village, Lidice . In the spring of 1945, death marches of prisoners of several subcamps of the Flossenbürg, Gross-Rosen and Buchenwald concentration camps in Saxony and Silesia, and Allied POWs from camps in Austria reached

6630-414: Was changed to the Czech Republic , which became a separate state in 1993 with the breakup of Czechoslovakia . Until 1948, Bohemia was an administrative unit of Czechoslovakia as one of its "lands" ( země ). Since then, administrative reforms have replaced self-governing lands with a modified system of "regions" ( kraje ), which do not follow the borders of the historical Czech lands (or the regions from

6715-526: Was elected to ascend to the Bohemian throne. He is remembered for his attempt to set up a pan-European "Christian League", which would form all the states of Europe into a community based on religion. In the process of negotiating, he appointed Zdeněk Lev of Rožmitál to tour the European courts and to conduct the talks. However, the negotiations were not completed because George's position was substantially damaged over time by his deteriorating relationship with

6800-656: Was examining fossil-bearing sedimentary rock strata in south Wales in the early 1830s. He named the sequences for a Celtic tribe of Wales, the Silures , inspired by his friend Adam Sedgwick , who had named the period of his study the Cambrian , from a Latin name for Wales. Whilst the British rocks now identified as belonging to the Silurian System and the lands now thought to have been inhabited in antiquity by

6885-616: Was facilitated by Bohemia's conversion to Christianity, in the 9th century. Continuing close relations were developed with the East Frankish Kingdom , which devolved from the Carolingian Empire , into East Francia , eventually becoming the Holy Roman Empire . After a decisive victory of the Holy Roman Empire and Bohemia over invading Magyars in the 955 Battle of Lechfeld , Boleslaus I of Bohemia

6970-482: Was granted the Moravia by German emperor Otto the Great . Bohemia would remain a largely autonomous state under the Holy Roman Empire for several decades. The jurisdiction of the Holy Roman Empire was definitively reasserted when Jaromír of Bohemia was granted fief of the Kingdom of Bohemia by Emperor King Henry II of the Holy Roman Empire, with the promise that he hold it as a vassal once he reoccupied Prague with

7055-409: Was questioned in 1854 by Edward Forbes , and the later stages of Barrande; F, G and H have since been shown to be Devonian. Despite these modifications in the original groupings of the strata, it is recognized that Barrande established Bohemia as a classic ground for the study of the earliest Silurian fossils. With the supercontinent Gondwana covering the equator and much of the southern hemisphere,

7140-807: Was spoken in much of the countryside. The formal independence of Bohemia was further jeopardized when the Bohemian Diet approved administrative reform in 1749. It included the indivisibility of the Habsburg Empire and the centralization of rule, which essentially meant the merging of the Royal Bohemian Chancellery with the Austrian Chancellery. At the end of the 18th century, the Czech National Revival movement, in cooperation with part of

7225-520: Was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution : vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods , arachnids and hexapods ) became fully terrestrialized. Another significant evolutionary milestone during

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