166-494: A lone wolf is a wolf not belonging to a pack . As a trope, it refers to an individual who prefers to operate alone. Lone wolf or Lone Wolf , or The Lone Wolf may also refer to: Wolf#Social structure See Subspecies of Canis lupus The wolf ( Canis lupus ; pl. : wolves ), also known as the gray wolf or grey wolf , is a canine native to Eurasia and North America . More than thirty subspecies of Canis lupus have been recognized, including
332-552: A mated pair accompanied by their offspring. Offspring may leave to form their own packs on the onset of sexual maturity and in response to competition for food within the pack. Wolves are also territorial , and fights over territory are among the principal causes of mortality. The wolf is mainly a carnivore and feeds on large wild hooved mammals as well as smaller animals, livestock, carrion , and garbage. Single wolves or mated pairs typically have higher success rates in hunting than do large packs. Pathogens and parasites, notably
498-530: A taxonomic synonym for the dingo . Wozencraft referred to a 1999 mitochondrial DNA (mtDNA) study as one of the guides in forming his decision, and listed the 38 subspecies of C. lupus under the biological common name of "wolf", the nominate subspecies being the Eurasian wolf ( C. l. lupus ) based on the type specimen that Linnaeus studied in Sweden. Studies using paleogenomic techniques reveal that
664-491: A Germanic language", are sometimes referred to as "Germanic-speaking peoples". Today, the term "Germanic" is widely applied to "phenomena including identities, social, cultural or political groups, to material cultural artefacts, languages and texts, and even specific chemical sequences found in human DNA". Several scholars continue to use the term to refer to a culture existing between the 1st to 4th centuries CE, but most historians and archaeologists researching Late Antiquity and
830-721: A Gothic group in modern Ukraine under the rule of Ermanaric , were among the first peoples attacked by the Huns, apparently facing Hunnic pressure for some years. Following Ermanaric's death, the Greuthungi's resistance broke and they moved toward the Dniester river. A second Gothic group, the Tervingi under King Athanaric , constructed a defensive earthwork against the Huns near the Dniester. However, these measures did not stop
996-726: A Gothic ruler of the Amal dynasty, seems to have consolidated power over a large part of the Goths in the Hunnic domain. For the next 20 years, the former subject peoples of the Huns would fight among each other for preeminence. The arrival of the Saxons in Britain is traditionally dated to 449, however, archaeology indicates they had begun arriving in Britain earlier. Latin sources used Saxon generically for seaborne raiders, meaning that not all of
1162-520: A broader snout, shorter ears, a shorter torso and a longer tail. It is slender and powerfully built, with a large, deeply descending rib cage , a sloping back, and a heavily muscled neck. The wolf's legs are moderately longer than those of other canids, which enables the animal to move swiftly, and to overcome the deep snow that covers most of its geographical range in winter, though more short-legged ecomorphs are found in some wolf populations. The ears are relatively small and triangular. The wolf's head
1328-971: A common language. Several ancient sources list subdivisions of the Germanic tribes. Writing in the first century CE, Pliny the Elder lists five Germanic subgroups: the Vandili, the Inguaeones, the Istuaeones (living near the Rhine), the Herminones (in the Germanic interior), and the Peucini Basternae (living on the lower Danube near the Dacians). In chapter 2 of the Germania , written about
1494-683: A den located away from the peripheral zone of their territories, where violent encounters with other packs are less likely to occur. Germanic peoples The Germanic peoples were tribal groups who lived in Northern Europe in Classical Antiquity and the Early Middle Ages . In modern scholarship, they typically include not only the Roman-era Germani who lived in both Germania and parts of
1660-448: A family of eight wolves (two adults, juveniles, and yearlings), or sometimes two or three such families, with examples of exceptionally large packs consisting of up to 42 wolves being known. Cortisol levels in wolves rise significantly when a pack member dies, indicating the presence of stress. During times of prey abundance caused by calving or migration, different wolf packs may join together temporarily. Offspring typically stay in
1826-433: A few times and then retreating from a perceived danger. Aggressive or self-assertive wolves are characterized by their slow and deliberate movements, high body posture and raised hackles , while submissive ones carry their bodies low, flatten their fur, and lower their ears and tail. Scent marking involves urine, feces, and preputial and anal gland scents. This is more effective at advertising territory than howling and
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#17327726219601992-505: A group of mutually intelligible dialects . They share distinctive characteristics which set them apart from other Indo-European sub-families of languages, such as Grimm's and Verner's law , the conservation of the PIE ablaut system in the Germanic verb system (notably in strong verbs ), or the merger of the vowels a and o qualities ( ə , a , o > a; ā , ō > ō ). During
2158-744: A half-century later, Tacitus lists only three subgroups: the Ingvaeones (near the sea), the Herminones (in the interior of Germania), and the Istvaeones (the remainder of the tribes); Tacitus says these groups each claimed descent from the god Mannus , son of Tuisto . Tacitus also mentions a second tradition that there were four sons of either Mannus or Tuisto from whom the groups of the Marsi, Gambrivi, Suebi, and Vandili claim descent. The Herminones are also mentioned by Pomponius Mela , but otherwise, these divisions do not appear in other ancient works on
2324-573: A high degree of Celtic-Germanic shared material culture and social organization. Some evidence of linguistic convergence between Germanic and Italic languages , whose Urheimat is supposed to have been situated north of the Alps before the 1st millennium BCE, have also been highlighted by scholars. Shared changes in their grammars also suggest early contacts between Germanic and Balto-Slavic languages ; however, some of these innovations are shared with Baltic only, which may point to linguistic contacts during
2490-426: A lancehead) and linguistic cognates attested in the later Old Norse , Old Saxon and Old High German languages: fremja , fremmian and fremmen all mean 'to carry out'. In the absence of earlier evidence, it must be assumed that Proto-Germanic speakers living in Germania were members of preliterate societies. The only pre-Roman inscriptions that could be interpreted as Proto-Germanic, written in
2656-494: A law dominated by the concepts of feuding and blood compensation . The precise details, nature and origin of what is still normally called " Germanic law " are now controversial. Roman sources state that the Germanic peoples made decisions in a popular assembly (the thing ) but that they also had kings and war leaders. The ancient Germanic-speaking peoples probably shared a common poetic tradition, alliterative verse , and later Germanic peoples also shared legends originating in
2822-674: A museum specimen of a wolf from southern China collected in 1963 showed a genome that was 12–14% admixed from this unknown canid. In North America, some coyotes and wolves show varying degrees of past genetic admixture . In more recent times, some male Italian wolves originated from dog ancestry, which indicates female wolves will breed with male dogs in the wild. In the Caucasus Mountains , ten percent of dogs including livestock guardian dogs , are first generation hybrids. Although mating between golden jackals and wolves has never been observed, evidence of jackal-wolf hybridization
2988-601: A policy of trying to prevent strong leaders from emerging among the barbarians, using treachery, kidnapping, and assassination, paying off rival tribes to attack them, or by supporting internal rivals. The Migration Period is traditionally cited by historians as beginning in 375 CE, under the assumption that the appearance of the Huns prompted the Visigoths to seek shelter within the Roman Empire in 376. The end of
3154-754: A press release by the U.S. Department of Agriculture in 1921, the infamous Custer Wolf relied on coyotes to accompany him and warn him of danger. Though they fed from his kills, he never allowed them to approach him. Interactions have been observed in Eurasia between wolves and golden jackals, the latter's numbers being comparatively small in areas with high wolf densities. Wolves also kill red , Arctic and corsac foxes , usually in disputes over carcasses, sometimes eating them. Brown bears typically dominate wolf packs in disputes over carcasses, while wolf packs mostly prevail against bears when defending their den sites. Both species kill each other's young. Wolves eat
3320-587: A relatively late period, at any rate after the initial breakup of Balto-Slavic into Baltic and Slavic languages , with the similarities to Slavic being seen as remnants of Indo-European archaisms or the result of secondary contacts. According to some authors the Bastarnae , or Peucini , were the first Germani to be encountered by the Greco-Roman world and thus to be mentioned in historical records. They appear in historical sources going as far back as
3486-742: A renewed political crisis in Rome, the Rhine frontier had collapsed, and in order to restore it, the Roman magister militum Flavius Aetius engineered the destruction of the Burgundian kingdom in 435/436, possibly with Hunnic mercenaries, and launched several successful campaigns against the Visigoths. In 439, the Vandals conquered Carthage , which served as an excellent base for further raids throughout
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#17327726219603652-567: A term corresponding to Germanic-speaking peoples, this new definition—which used the Germanic language as the main criterion—presented the Germani as a people or nation ( Volk ) with a stable group identity linked to language. As a result, some scholars treat the Germani (Latin) or Germanoi (Greek) of Roman-era sources as non-Germanic if they seemingly spoke non-Germanic languages. For clarity, Germanic peoples, when defined as "speakers of
3818-494: A territorial definition ("those living in Germania ") and an ethnic definition ("having Germanic ethnic characteristics"), and the two definitions did not always align. In the 3rd century, when Romans encountered Germanic-speaking peoples living north of the Lower Danube who fought on horseback, such as Goths and Gepids, they did not call them Germani . Instead, they connected them with non-Germanic-speaking peoples such as
3984-620: A territorial sense to refer to East Francia . In modern English, the adjective Germanic is distinct from German , which is generally used when referring to modern Germans only. Germanic relates to the ancient Germani or the broader Germanic group. In modern German, the ancient Germani are referred to as Germanen and Germania as Germanien , as distinct from modern Germans ( Deutsche ) and modern Germany ( Deutschland ). The direct equivalents in English are, however, Germans for Germani and Germany for Germania although
4150-409: A variety of arthropod exoparasites, including fleas , ticks , lice , and mites . The most harmful to wolves, particularly pups, is the mange mite ( Sarcoptes scabiei ), though they rarely develop full-blown mange , unlike foxes. Endoparasites known to infect wolves include: protozoans and helminths ( flukes , tapeworms , roundworms and thorny-headed worms ). Most fluke species reside in
4316-648: A war-god or a mark of ownership engraved by its possessor. The inscription Fariarix ( * farjōn- 'ferry' + * rīk- 'ruler') carved on tetradrachms found in Bratislava (mid-1st c. BCE) may indicate the Germanic name of a Celtic ruler. By the time Germanic speakers entered written history, their linguistic territory had stretched farther south, since a Germanic dialect continuum (where neighbouring language varieties diverged only slightly between each other, but remote dialects were not necessarily mutually intelligible due to accumulated differences over
4482-411: A weight of 69–79 kg (152–174 lb). The wolf has very dense and fluffy winter fur, with a short undercoat and long, coarse guard hairs . Most of the undercoat and some guard hairs are shed in spring and grow back in autumn. The longest hairs occur on the back, particularly on the front quarters and neck. Especially long hairs grow on the shoulders and almost form a crest on the upper part of
4648-479: Is difficult and ambiguous, with some authors choosing to include C. mosbachensis (which first appeared around 1.4 million years ago) as an early subspecies of C. lupus. Considerable morphological diversity existed among wolves by the Late Pleistocene . Many Late Pleistocene wolf populations had more robust skulls and teeth than modern wolves, often with a shortened snout , a pronounced development of
4814-525: Is dominated by wild medium-sized hoofed mammals and domestic species. The wolf depends on wild species, and if these are not readily available, as in Asia, the wolf is more reliant on domestic species. Across Eurasia, wolves prey mostly on moose , red deer , roe deer and wild boar . In North America, important range-wide prey are elk , moose, caribou , white-tailed deer and mule deer . Prior to their extirpation from North America, wild horses were among
4980-478: Is equally inconsistent. Additionally, there is no linguistic or archaeological evidence for these subgroups. New archaeological finds have tended to show that the boundaries between Germanic peoples were very permeable, and scholars now assume that migration and the collapse and formation of cultural units were constant occurrences within Germania. Nevertheless, various aspects such as the alliteration of many of
5146-1220: Is itself thought to be derived from the Proto-Germanic * wulfaz . The Proto-Indo-European root * wĺ̥kʷos may also be the source of the Latin word for the animal lupus (* lúkʷos ). The name "gray wolf" refers to the grayish colour of the species. Since pre-Christian times, Germanic peoples such as the Anglo-Saxons took on wulf as a prefix or suffix in their names. Examples include Wulfhere ("Wolf Army"), Cynewulf ("Royal Wolf"), Cēnwulf ("Bold Wolf"), Wulfheard ("Wolf-hard"), Earnwulf ("Eagle Wolf"), Wulfstān ("Wolf Stone") Æðelwulf ("Noble Wolf"), Wolfhroc ("Wolf-Frock"), Wolfhetan ("Wolf Hide"), Scrutolf ("Garb Wolf"), Wolfgang ("Wolf Gait") and Wolfdregil ("Wolf Runner"). Gray wolf [REDACTED] Coyote [REDACTED] African wolf [REDACTED] Ethiopian wolf [REDACTED] Golden jackal [REDACTED] Dhole [REDACTED] African wild dog [REDACTED] Side-striped jackal [REDACTED] Black-backed jackal [REDACTED] In 1758,
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5312-399: Is large and heavy, with a wide forehead, strong jaws and a long, blunt muzzle. The skull is 230–280 mm (9–11 in) in length and 130–150 mm (5–6 in) in width. The teeth are heavy and large, making them better suited to crushing bone than those of other canids, though they are not as specialized as those found in hyenas . Its molars have a flat chewing surface, but not to
5478-541: Is largely survivable in wolves, but can be lethal to pups. Bacterial diseases carried by wolves include: brucellosis , Lyme disease , leptospirosis , tularemia , bovine tuberculosis , listeriosis and anthrax . Although lyme disease can debilitate individual wolves, it does not appear to significantly affect wolf populations. Leptospirosis can be contracted through contact with infected prey or urine, and can cause fever , anorexia , vomiting, anemia , hematuria , icterus , and death. Wolves are often infested with
5644-401: Is little evidence for a common Germanic identity. The Anglo-Saxonist Leonard Neidorf writes that historians of the continental-European Germanic peoples of the 5th and 6th centuries are "in agreement" that there was no pan-Germanic identity or solidarity. Whether a scholar favors the existence of a common Germanic identity or not is often related to their position on the nature of the end of
5810-703: Is most likely used mainly to induce vomiting to rid themselves of intestinal parasites or long guard hairs. They are known to eat the berries of mountain-ash , lily of the valley , bilberries , cowberries , European black nightshade , grain crops, and the shoots of reeds. In times of scarcity, wolves will readily eat carrion . In Eurasian areas with dense human activity, many wolf populations are forced to subsist largely on livestock and garbage. As prey in North America continue to occupy suitable habitats with low human density, North American wolves eat livestock and garbage only in dire circumstances. Cannibalism
5976-562: Is not uncommon in wolves during harsh winters, when packs often attack weak or injured wolves and may eat the bodies of dead pack members. Wolves typically dominate other canid species in areas where they both occur. In North America, incidents of wolves killing coyotes are common, particularly in winter, when coyotes feed on wolf kills. Wolves may attack coyote den sites, digging out and killing their pups, though rarely eating them. There are no records of coyotes killing wolves, though coyotes may chase wolves if they outnumber them. According to
6142-677: Is now extirpated (locally extinct) from much of its range in Western Europe, the United States and Mexico, and completely in the British Isles and Japan. In modern times, the wolf occurs mostly in wilderness and remote areas. The wolf can be found between sea level and 3,000 m (9,800 ft). Wolves live in forests, inland wetlands , shrublands , grasslands (including Arctic tundra ), pastures , deserts, and rocky peaks on mountains. Habitat use by wolves depends on
6308-458: Is often used in combination with scratch marks. Wolves increase their rate of scent marking when they encounter the marks of wolves from other packs. Lone wolves will rarely mark, but newly bonded pairs will scent mark the most. These marks are generally left every 240 m (260 yd) throughout the territory on regular travelways and junctions. Such markers can last for two to three weeks, and are typically placed near rocks, boulders, trees, or
6474-816: Is possible to refer to Germanic languages from about 500 BCE. Archaeologists usually associate the earliest clearly identifiable Germanic speaking peoples with the Jastorf culture of the Pre-Roman Iron Age in central and northern Germany and southern Denmark from the 6th to 1st centuries BCE. This existed around the same time that the First Germanic Consonant Shift is theorized to have occurred, leading to recognizably Germanic languages. Germanic languages expanded south, east, and west, coming into contact with Celtic , Iranic , Baltic , and Slavic peoples before they were noted by
6640-416: Is related to a lack of stable frontiers in this area such as were maintained by Roman armies along the Rhine and Danube. The geographer Ptolemy (2nd century CE) applied the name Germania magna ("Greater Germania", Greek : Γερμανία Μεγάλη ) to this area, contrasting it with the Roman provinces of Germania Prima and Germania Secunda (on the west bank of the Rhine). In modern scholarship, Germania magna
6806-569: Is related to the Old Irish word gair ('neighbours') or could be tied to the Celtic word for their war cries, gairm , which simplifies into 'the neighbours' or 'the screamers'. Regardless of its language of origin, the name was transmitted to the Romans via Celtic speakers. It is unclear that any people group ever referred to themselves as Germani . By late antiquity , only peoples near
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6972-421: Is sometimes also called Germania libera ("free Germania"), a name coined by Jacob Grimm around 1835. Caesar and, following him, Tacitus, depicted the Germani as sharing elements of a common culture. A small number of passages by Tacitus and other Roman authors (Caesar, Suetonius) mention Germanic tribes or individuals speaking a language distinct from Gaulish. For Tacitus ( Germania 43, 45, 46), language
7138-604: Is thought to possibly reflect a Germanic and Slavic component. The identification of the Jastorf culture with the Germani has been criticized by Sebastian Brather , who notes that it seems to be missing areas such as southern Scandinavia and the Rhine-Weser area, which linguists argue to have been Germanic, while also not according with the Roman era definition of Germani , which included Celtic-speaking peoples further south and west. A category of evidence used to locate
7304-412: Is unknown, although several proposals have been put forward. Even the language from which it derives is a subject of dispute, with proposals of Germanic, Celtic , and Latin, and Illyrian origins. Herwig Wolfram , for example, thinks Germani must be Gaulish . The historian Wolfgang Pfeifer more or less concurs with Wolfram and surmises that the name Germani is likely of Celtic etymology and
7470-678: Is unlikely that Germanic populations spoke a single dialect, and traces of early linguistic varieties have been highlighted by scholars. Sister dialects of Proto-Germanic itself certainly existed, as evidenced by the absence of the First Germanic Sound Shift (Grimm's law) in some "Para-Germanic" recorded proper names, and the reconstructed Proto-Germanic language was only one among several dialects spoken at that time by peoples identified as "Germanic" by Roman sources or archeological data. Although Roman sources name various Germanic tribes such as Suevi, Alemanni, Bauivari , etc., it
7636-498: Is unlikely that the members of these tribes all spoke the same dialect. Definite and comprehensive evidence of Germanic lexical units only occurred after Caesar 's conquest of Gaul in the 1st century BCE, after which contacts with Proto-Germanic speakers began to intensify. The Alcis , a pair of brother gods worshipped by the Nahanarvali , are given by Tacitus as a Latinized form of * alhiz (a kind of ' stag '), and
7802-595: Is usually constructed not more than 500 m (550 yd) away from a water source. It typically faces southwards where it can be better warmed by sunlight exposure, and the snow can thaw more quickly. Resting places, play areas for the pups, and food remains are commonly found around wolf dens. The odor of urine and rotting food emanating from the denning area often attracts scavenging birds like magpies and ravens . Though they mostly avoid areas within human sight, wolves have been known to nest near homes, paved roads and railways . During pregnancy, female wolves remain in
7968-435: Is usually founded by an unrelated dispersing male and female, travelling together in search of an area devoid of other hostile packs. Wolf packs rarely adopt other wolves into their fold and typically kill them. In the rare cases where other wolves are adopted, the adoptee is almost invariably an immature animal of one to three years old, and unlikely to compete for breeding rights with the mated pair. This usually occurs between
8134-665: The Urheimat ('original homeland') of the Proto-Germanic language , the ancestral idiom of all attested Germanic dialects, existed in or near the archaeological culture known as the late Jastorf culture , of the central Elbe in present day Germany, stretching north into Jutland and east into present day Poland. If the Jastorf Culture is the origin of the Germanic peoples, then the Scandinavian peninsula would have become Germanic either via migration or assimilation over
8300-414: The Anglo-Saxons of Britain converted to Christianity, but the Saxons and Scandinavians converted only much later. The Germanic peoples shared a native script—known as runes —from around the first century or before, which was gradually replaced with the Latin script , although runes continued to be used for specialized purposes thereafter. Traditionally, the Germanic peoples have been seen as possessing
8466-580: The Cimbrian War (113–101 BCE) against the Romans, in which the Teutons and Cimbri were victorious over several Roman armies but were ultimately defeated. The first century BCE was a time of the expansion of Germanic-speaking peoples at the expense of Celtic-speaking polities in modern southern Germany and the Czech Republic. Before 60 BCE, Ariovistus , described by Caesar as king of
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#17327726219608632-609: The Corded Ware culture towards modern-day Denmark, resulting in cultural mixing with the earlier Funnelbeaker culture . The subsequent culture of the Nordic Bronze Age (c. 2000/1750 – c. 500 BCE) shows definite cultural and population continuities with later Germanic peoples, and is often supposed to have been the culture in which the Germanic Parent Language , the predecessor of
8798-599: The Etruscan alphabet , have not been found in Germania but rather in the Venetic region. The inscription harikastiteiva \\\ip , engraved on the Negau helmet in the 3rd–2nd centuries BCE, possibly by a Germanic-speaking warrior involved in combat in northern Italy, has been interpreted by some scholars as Harigasti Teiwǣ ( * harja-gastiz 'army-guest' + * teiwaz 'god, deity'), which could be an invocation to
8964-601: The Eurasian lynx , which may feed on smaller prey where wolves are present and may be suppressed by large wolf populations. Wolves encounter cougars along portions of the Rocky Mountains and adjacent mountain ranges. Wolves and cougars typically avoid encountering each other by hunting at different elevations for different prey ( niche partitioning ). This is more difficult during winter. Wolves in packs usually dominate cougars and can steal their kills or even kill them, while one-to-one encounters tend to be dominated by
9130-800: The Franks , Goths , Saxons , and Alemanni . During the Migration Period (375–568), such Germanic peoples entered the Roman Empire and eventually established their own " barbarian kingdoms " within the territory of the Western Roman empire itself. Over time, the Franks became the most powerful of them, conquering many of the others. Eventually, the Frankish king Charlemagne claimed the title of Holy Roman Emperor for himself in 800. Archaeological finds suggest that Roman-era sources portrayed
9296-514: The Germani and Celtic peoples , usually identified with the archaeological La Tène culture , found in southern Germany and the modern Czech Republic. Early contacts probably occurred during the Pre-Germanic and Pre-Celtic periods, dated to the 2nd millennium BCE, and the Celts appear to have had a large amount of influence on Germanic culture from up until the first century CE, which led to
9462-516: The Germani as a large category of peoples distinct from the Gauls and Scythians was Julius Caesar , writing around 55 BCE during his governorship of Gaul. In Caesar's account, the clearest defining characteristic of the Germani people was that their homeland was east of the Rhine , opposite Gaul on the west side. Caesar sought to explain both why his legions stopped at the Rhine and also why
9628-618: The Germani were more dangerous than the Gauls to the empire. Explaining this threat he also classified the Cimbri and Teutons , who had previously invaded Italy, as Germani . Although Caesar described the Rhine as the border between Germani and Celts, he also describes the Germani cisrhenani on the west bank of the Rhine, who he believed had moved from the east. It is unclear if these Germani were actually Germanic speakers. According to
9794-636: The Germani , led a force including Suevi across the Rhine into Gaul near Besançon , successfully aiding the Sequani against their enemies the Aedui at the Battle of Magetobriga . Ariovistus was initially considered an ally of Rome. In 58 BCE, with increasing numbers of settlers crossing the Rhine to join Ariovistus, Julius Caesar went to war with them, defeating them at the Battle of Vosges . In
9960-497: The Germani , though they did not live in Germania, and they were beginning to look like Sarmatians through intermarriage. The Osi and Cotini lived in Germania, but were not Germani , because they had other languages and customs. The Aesti lived on the eastern shore of the Baltic and were like Suebi in their appearance and customs, although they spoke a different language. Ancient authors did not differentiate consistently between
10126-443: The Germani . There are a number of inconsistencies in the listing of Germanic subgroups by Tacitus and Pliny. While both Tacitus and Pliny mention some Scandinavian tribes, they are not integrated into the subdivisions. While Pliny lists the Suebi as part of the Herminones, Tacitus treats them as a separate group. Additionally, Tacitus's description of a group of tribes as united by the cult of Nerthus ( Germania 40) as well as
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#173277262196010292-418: The Huns , Sarmatians , and Alans , who shared a similar culture. Romans also called them "Gothic peoples", ( gentes Gothicae ) even if they did not speak a Germanic language, and they often referred to the Goths as " Getae ", equating them to a non-Germanic people residing in the same region. The writer Procopius described these new "Getic" peoples as sharing similar appearance, laws, Arian religion, and
10458-427: The Pre-Germanic linguistic period (2500–500 BCE), the proto-language was almost certainly influenced by an unknown non-Indo-European language , still noticeable in the Germanic phonology and lexicon . Although Proto-Germanic is reconstructed without dialects via the comparative method , it is almost certain that it never was a uniform proto-language. The late Jastorf culture occupied so much territory that it
10624-467: The Rhine , to southern Scandinavia and the Vistula in the east, and to the upper Danube in the south. Other Germanic speakers, such as the Bastarnae and Goths, lived further east in what is now Moldova and Ukraine . The term Germani is generally only used to refer to historical peoples from the 1st to 4th centuries CE. Different academic disciplines have their own definitions of what makes someone or something "Germanic". Some scholars call for
10790-468: The Saxon tribes towards modern-day England. The Germanic languages are traditionally divided between East , North and West Germanic branches. The modern prevailing view is that North and West Germanic were also encompassed in a larger subgroup called Northwest Germanic. Further internal classifications are still debated among scholars, as it is unclear whether the internal features shared by several branches are due to early common innovations or to
10956-543: The coyote and the golden jackal , to produce fertile hybrids with them. The wolf's fur is usually mottled white, brown, gray, and black, although subspecies in the arctic region may be nearly all white. Of all members of the genus Canis , the wolf is most specialized for cooperative game hunting as demonstrated by its physical adaptations to tackling large prey, its more social nature , and its highly advanced expressive behaviour , including individual or group howling . It travels in nuclear families consisting of
11122-414: The dog and dingo , though gray wolves, as popularly understood, only comprise naturally-occurring wild subspecies. The wolf is the largest wild extant member of the family Canidae , and is further distinguished from other Canis species by its less pointed ears and muzzle, as well as a shorter torso and a longer tail. The wolf is nonetheless related closely enough to smaller Canis species, such as
11288-457: The incubation period for rabies is eight to 21 days, and results in the host becoming agitated, deserting its pack, and travelling up to 80 km (50 mi) a day, thus increasing the risk of infecting other wolves. Although canine distemper is lethal in dogs, it has not been recorded to kill wolves, except in Canada and Alaska. The canine parvovirus, which causes death by dehydration , electrolyte imbalance , and endotoxic shock or sepsis ,
11454-467: The rabies virus , may infect wolves. The global wild wolf population was estimated to be 300,000 in 2003 and is considered to be of Least Concern by the International Union for Conservation of Nature (IUCN). Wolves have a long history of interactions with humans, having been despised and hunted in most pastoral communities because of their attacks on livestock, while conversely being respected in some agrarian and hunter-gatherer societies. Although
11620-610: The temporalis muscle, and robust premolars . It is proposed that these features were specialized adaptations for the processing of carcass and bone associated with the hunting and scavenging of Pleistocene megafauna . Compared with modern wolves, some Pleistocene wolves showed an increase in tooth breakage similar to that seen in the extinct dire wolf . This suggests they either often processed carcasses, or that they competed with other carnivores and needed to consume their prey quickly. The frequency and location of tooth fractures in these wolves indicates they were habitual bone crackers like
11786-463: The 2nd and 3rd centuries CE, migrations of East Germanic gentes from the Baltic Sea coast southeastwards into the hinterland led to their separation from the dialect continuum. By the late 3rd century CE, linguistic divergences like the West Germanic loss of the final consonant -z had already occurred within the "residual" Northwest dialect continuum. The latter definitely ended after the 5th- and 6th-century migrations of Angles , Jutes and part of
11952-665: The 3rd century BCE through the 4th century CE. Another eastern people known from about 200 BCE, and sometimes believed to be Germanic-speaking, are the Sciri (Greek: Skiroi ), who are recorded threatening the city of Olbia on the Black Sea. Late in the 2nd century BCE, Roman and Greek sources recount the migrations of the Cimbri, Teutones and Ambrones whom Caesar later classified as Germanic. The movements of these groups through parts of Gaul , Italy and Hispania resulted in
12118-541: The Cherusci—initially an ally of Rome—drew a large Roman force into an ambush in northern Germany, and destroyed the three legions of Publius Quinctilius Varus at the Battle of the Teutoburg Forest . Marboduus and Arminius went to war with each other in 17 CE; Arminius was victorious and Marboduus was forced to flee to the Romans. Following the Roman defeat at the Teutoburg Forest, Rome gave up on
12284-668: The Early Middle Ages no longer use it. Apart from the designation of a language family (i.e., "Germanic languages"), the application of the term "Germanic" has become controversial in scholarship since 1990, especially among archaeologists and historians. Scholars have increasingly questioned the notion of ethnically defined people groups ( Völker ) as stable basic actors of history. The connection of archaeological assemblages to ethnicity has also been increasingly questioned. This has resulted in different disciplines developing different definitions of "Germanic". Beginning with
12450-628: The Egyptian Sinai Peninsula showed admixture with Middle Eastern gray wolves and dogs. There is evidence of gene flow between golden jackals and Middle Eastern wolves, less so with European and Asian wolves, and least with North American wolves. This indicates the golden jackal ancestry found in North American wolves may have occurred before the divergence of the Eurasian and North American wolves. The common ancestor of
12616-399: The Eurasian wolf is a mixture of ochreous (yellow to orange) and rusty ochreous (orange/red/brown) colours with light gray. The muzzle is pale ochreous gray, and the area of the lips, cheeks, chin, and throat is white. The top of the head, forehead, under and between the eyes, and between the eyes and ears is gray with a reddish film. The neck is ochreous. Long, black tips on the hairs along
12782-612: The Germanic frontier carefully, meddling in cross-border politics, and constructing a long fortified border, the Limes Germanicus . From 166 to 180 CE, Rome was embroiled in a conflict against the Germanic Marcomanni and Quadi with their allies, which was known as the Marcomannic Wars . After this major disruption, new Germanic peoples appear for the first time in the historical record, such as
12948-640: The Germanic peoples divided and fractious. Rome established relationships with individual Germanic kings that are often discussed as being similar to client states ; however, the situation on the border was always unstable, with rebellions by the Frisians in 28 CE, and attacks by the Chauci and Chatti in the 60s CE. The most serious threat to the Roman order was the Revolt of the Batavi in 69 CE, during
13114-430: The Germanic way of life as more primitive than it actually was. Instead, archaeologists have unveiled evidence of a complex society and economy throughout Germania. Germanic-speaking peoples originally shared similar religious practices. Denoted by the term Germanic paganism , they varied throughout the territory occupied by Germanic-speaking peoples. Over the course of Late Antiquity , most continental Germanic peoples and
13280-470: The Greuthungi. The Goths and their allies defeated the Romans first at Marcianople , then defeated and killed emperor Valens in the Battle of Adrianople in 378, destroying two-thirds of Valens' army. Following further fighting, peace was negotiated in 382, granting the Goths considerable autonomy within the Roman Empire. However, these Goths—who would be known as the Visigoths —revolted several more times, finally coming to be ruled by Alaric . In 397,
13446-580: The Huns and the majority of the Tervingi abandoned Athanaric; they subsequently fled—accompanied by a contingent of Greuthungi—to the Danube in 376, seeking asylum in the Roman Empire. The emperor Valens chose only to admit the Tervingi, who were settled in the Roman provinces of Thrace and Moesia . Due to mistreatment by the Romans, the Tervingi revolted in 377, starting the Gothic War , joined by
13612-676: The Huns had largely conquered them by 406. One Gothic group under Hunnic domination was ruled by the Amal dynasty , who would form the core of the Ostrogoths . The situation outside the Roman empire in 410s and 420s is poorly attested, but it is clear that the Huns continued to spread their influence onto the middle Danube. In 428, the Vandal leader Geiseric moved his forces across the strait of Gibraltar into north Africa. Within two years, they had conquered most of north Africa. By 434, following
13778-522: The Huns interfered in a Frankish succession dispute, leading in 451 to an invasion of Gaul. Aetius, by uniting a coalition of Visigoths, part of the Franks, and others, was able to defeat the Hunnic army at the Battle of the Catalaunian Plains . In 453, Attila died unexpectedly, and an alliance led by Ardaric's Gepids rebelled against the rule of his sons, defeating them in the Battle of Nedao . Either before or after Attila's death, Valamer ,
13944-647: The Latin Germania is also used. To avoid ambiguity, the Germani may instead be called "ancient Germans" or Germani by using the Latin term in English. The modern definition of Germanic peoples developed in the 19th century, when the term Germanic was linked to the newly identified Germanic language family . Linguistics provided a new way of defining the Germanic peoples, which came to be used in historiography and archaeology. While Roman authors did not consistently exclude Celtic-speaking people or have
14110-568: The Marcomanni and Quadi, and Commodus forbid them to hold assemblies unless a Roman centurion was present. The period after the Marcomannic Wars saw the emergence of peoples with new names along the Roman frontiers, which were probably formed by the merger of smaller groups. These new confederacies or peoples tended to border the Roman imperial frontier. Many ethnic names from earlier periods disappear. The Alamanni emerged along
14276-771: The Mediterranean and became the basis for the Vandal Kingdom . The loss of Carthage forced Aetius to make peace with the Visigoths in 442, effectively recognizing their independence within the boundaries of the empire. During the resulting peace, Aetius resettled the Burgundians in Sapaudia in southern Gaul. In the 430s, Aetius negotiated peace with the Suevi in Spain, leading to a practical loss of Roman control in
14442-513: The Middle East, and Asia is estimated to be fairly recent at around 1,600 years ago. Among New World wolves, the Mexican wolf diverged around 5,400 years ago. In the distant past, there was gene flow between African wolves , golden jackals , and gray wolves. The African wolf is a descendant of a genetically admixed canid of 72% wolf and 28% Ethiopian wolf ancestry. One African wolf from
14608-564: The Migration Period. The publishing of Tacitus 's Germania by humanist scholars in the 1400s greatly influenced the emerging idea of "Germanic peoples". Later scholars of the Romantic period , such as Jacob and Wilhelm Grimm , developed several theories about the nature of the Germanic peoples that were highly influenced by romantic nationalism . For those scholars, the "Germanic" and modern "German" were identical. Ideas about
14774-769: The Proto-Germanic homeland is founded on traces of early linguistic contacts with neighbouring languages. Germanic loanwords in the Finnic and Sámi languages have preserved archaic forms (e.g. Finnic kuningas , from Proto-Germanic * kuningaz 'king'; rengas , from * hringaz 'ring'; etc.), with the older loan layers possibly dating back to an earlier period of intense contacts between pre-Germanic and Finno-Permic (i.e. Finno-Samic ) speakers. Shared lexical innovations between Celtic and Germanic languages, concentrated in certain semantic domains such as religion and warfare, indicates intensive contacts between
14940-457: The Proto-Germanic language, developed. However, it is unclear whether these earlier peoples possessed any ethnic continuity with the later Germanic peoples. Generally, scholars agree that it is possible to speak of Germanic-speaking peoples after 500 BCE, although the first attestation of the name Germani is not until much later. Between around 500 BCE and the beginning of the common era , archeological and linguistic evidence suggest that
15106-581: The Rhine as a border. Starting in 13 BCE, there were Roman campaigns across the Rhine for a 28-year period. First came the pacification of the Usipetes, Sicambri, and Frisians near the Rhine, then attacks increased further from the Rhine, on the Chauci , Cherusci , Chatti and Suevi (including the Marcomanni ). These campaigns eventually reached and even crossed the Elbe, and in 5 CE Tiberius
15272-793: The Rhine for an indeterminate distance, bounded by the Baltic Sea and the Hercynian Forest . Pliny the Elder and Tacitus placed the eastern border at the Vistula . The Upper Danube served as a southern border. Between there and the Vistula Tacitus sketched an unclear boundary, describing Germania as separated in the south and east from the Dacians and the Sarmatians by mutual fear or mountains. This undefined eastern border
15438-531: The Rhine, especially the Franks and sometimes the Alemanni, were called Germani or Germanoi by Latin and Greek writers respectively. Germani subsequently ceased to be used as a name for any group of people and was revived as such only by the humanists in the 16th century. Previously, scholars during the Carolingian period (8th–11th centuries) had already begun using Germania and Germanicus in
15604-473: The Roman Empire . Defenders of continued use of the term Germanic argue that the speakers of Germanic languages can be identified as Germanic people by language regardless of how they saw themselves. Linguists and philologists have generally reacted skeptically to claims that there was no Germanic identity or cultural unity, and they may view Germanic simply as a long-established and convenient term. Some archaeologists have also argued in favor of retaining
15770-629: The Roman emperor Flavius Constantius , the Visigoths were settled as Roman allies in Gaul between modern Toulouse and Bourdeaux. Other Goths, including those of Athanaric, continued to live outside the empire, with three groups crossing into the Roman territory after the Tervingi. The Huns gradually conquered Gothic groups north of the Danube, of which at least six are known, from 376 to 400. Those in Crimea may never have been conquered. The Gepids also formed an important Germanic people under Hunnic rule;
15936-466: The Roman empire, but also all Germanic speaking peoples from this era, irrespective of where they lived, most notably the Goths . Another term, ancient Germans , is considered problematic by many scholars since it suggests identity with present-day Germans . Although the first Roman descriptions of Germani involved tribes west of the Rhine, their homeland of Germania was portrayed as stretching east of
16102-641: The Roman historian Tacitus in his Germania (c. 98 CE), it was among this group, specifically the Tungri , that the name Germani first arose, before it spread to further groups. Tacitus reported that in his time many of the peoples west of the Rhine within Roman Gaul were still considered Germani . Caesar's division of the Germani from the Celts was not taken up by most writers in Greek. Caesar and authors following him regarded Germania as stretching east of
16268-609: The Roman territory. The revolt ended following several defeats, with Civilis claiming to have only supported the imperial claims of Vespasian , who was victorious in the civil war. The century after the Batavian Revolt saw mostly peace between the Germanic peoples and Rome. In 83 CE, Emperor Domitian of the Flavian dynasty attacked the Chatti north of Mainz (Mogontiacum). This war would last until 85 CE. Following
16434-614: The Romans appear to have reserved the right to choose rulers among the barbarians on the frontier. Following sixty years of quiet on the frontier, 166 CE saw a major incursion of peoples from north of the Danube during the reign of Marcus Aurelius , beginning the Marcomannic Wars . By 168 (during the Antonine plague ), barbarian hosts consisting of Marcomanni, Quadi, and Sarmatian Iazyges, attacked and pushed their way to Italy. They advanced as far as Upper Italy, destroyed Opitergium/Oderzo and besieged Aquileia. The Romans had finished
16600-429: The Romans. Roman authors first described the Germani near the Rhine in the 1st century BCE, while the Roman Empire was establishing its dominance in that region. Under Emperor Augustus (27 BCE – 14 CE), the Romans attempted to conquer a large part of Germania between the Rhine and Elbe , but withdrew after their shocking defeat at the Battle of the Teutoburg Forest in 9 CE. The Romans continued to manage
16766-585: The Saxons, a term used generically in Latin for Germanic-speaking pirates. A system of defenses on both sides of the English Channel , the Saxon Shore , was established to deal with their raids. From 250 onward, the Gothic peoples formed the "single most potent threat to the northern frontier of Rome". In 250 CE a Gothic king Cniva led Goths with Bastarnae, Carpi, Vandals, and Taifali into
16932-466: The Swedish botanist and zoologist Carl Linnaeus published in his Systema Naturae the binomial nomenclature . Canis is the Latin word meaning " dog ", and under this genus he listed the doglike carnivores including domestic dogs, wolves, and jackals . He classified the domestic dog as Canis familiaris , and the wolf as Canis lupus . Linnaeus considered the dog to be a separate species from
17098-442: The abundance of prey, snow conditions, livestock densities, road densities, human presence and topography . Like all land mammals that are pack hunters , the wolf feeds predominantly on ungulates that can be divided into large size 240–650 kg (530–1,430 lb) and medium size 23–130 kg (51–287 lb), and have a body mass similar to that of the combined mass of the pack members. The wolf specializes in preying on
17264-481: The age of the pack's pups. They tend to increase in size in areas with low prey populations, or when the pups reach the age of six months when they have the same nutritional needs as adults. Wolf packs travel constantly in search of prey, covering roughly 9% of their territory per day, on average 25 km/d (16 mi/d). The core of their territory is on average 35 km (14 sq mi) where they spend 50% of their time. Prey density tends to be much higher on
17430-414: The age of two years and sexually mature from the age of three years. The age of first breeding in wolves depends largely on environmental factors: when food is plentiful, or when wolf populations are heavily managed, wolves can rear pups at younger ages to better exploit abundant resources. Females are capable of producing pups every year, one litter annually being the average. Oestrus and rut begin in
17596-437: The back form a broad stripe, with black hair tips on the shoulders, upper chest and rear of the body. The sides of the body, tail, and outer limbs are a pale dirty ochreous colour, while the inner sides of the limbs, belly, and groin are white. Apart from those wolves which are pure white or black, these tones vary little across geographical areas, although the patterns of these colours vary between individuals. In North America,
17762-655: The brown bears they kill, while brown bears seem to eat only young wolves. Wolf interactions with American black bears are much rarer because of differences in habitat preferences. Wolves have been recorded on numerous occasions actively seeking out American black bears in their dens and killing them without eating them. Unlike brown bears, American black bears frequently lose against wolves in disputes over kills. Wolves also dominate and sometimes kill wolverines , and will chase off those that attempt to scavenge from their kills. Wolverines escape from wolves in caves or up trees. Wolves may interact and compete with felids , such as
17928-645: The cat, who likewise will kill wolves. Wolves more broadly affect cougar population dynamics and distribution by dominating territory and prey opportunities and disrupting the feline's behaviour. Wolf and Siberian tiger interactions are well-documented in the Russian Far East , where tigers significantly depress wolf numbers, sometimes to the point of localized extinction . In Israel, Palestine, Central Asia and India wolves may encounter striped hyenas , usually in disputes over carcasses. Striped hyenas feed extensively on wolf-killed carcasses in areas where
18094-621: The civil wars following the death of Nero known as the Year of the Four Emperors . The Batavi had long served as auxiliary troops in the Roman army as well as in the imperial bodyguard as the so-called Numerus Batavorum , often called the Germanic bodyguard. The uprising was led by Gaius Julius Civilis , a member of the Batavian royal family and Roman military officer, and attracted a large coalition of people both inside and outside of
18260-665: The coat colours of wolves follow Gloger's rule , wolves in the Canadian arctic being white and those in southern Canada, the U.S., and Mexico being predominantly gray. In some areas of the Rocky Mountains of Alberta and British Columbia, the coat colour is predominantly black, some being blue-gray and some with silver and black. Differences in coat colour between sexes is absent in Eurasia; females tend to have redder tones in North America. Black-coloured wolves in North America acquired their colour from wolf-dog admixture after
18426-419: The confrontation with Rome as things that could cause a sense of shared "Germanic" culture. Despite being cautious of the use of Germanic to refer to peoples, Sebastian Brather , Wilhelm Heizmann and Steffen Patzold nevertheless refer to further commonalities such as the widely attested worship of deities such as Odin , Thor and Frigg , and a shared legendary tradition . The first author to describe
18592-586: The course of the same period. Alternatively, Hermann Ament [ de ] has stressed that two other archaeological groups must have belonged to the Germani , one on either side of the Lower Rhine and reaching to the Weser , and another in Jutland and southern Scandinavia. These groups would thus show a "polycentric origin" for the Germanic peoples. The neighboring Przeworsk culture in modern Poland
18758-460: The coyote and the wolf admixed with a ghost population of an extinct unidentified canid. This canid was genetically close to the dhole and evolved after the divergence of the African hunting dog from the other canid species. The basal position of the coyote compared to the wolf is proposed to be due to the coyote retaining more of the mitochondrial genome of this unidentified canid. Similarly,
18924-441: The crisis. From the later third century onward, the Roman army relied increasingly on troops of Barbarian origin, often recruited from Germanic peoples, with some functioning as senior commanders in the Roman army. In the 4th century, warfare along the Rhine frontier between the Romans and Franks and Alemanni seems to have mostly consisted of campaigns of plunder, during which major battles were avoided. The Romans generally followed
19090-802: The cult of the Alcis controlled by the Nahanarvali ( Germania 43) and Tacitus's account of the origin myth of the Semnones ( Germania 39) all suggest different subdivisions than the three mentioned in Germania chapter 2. The subdivisions found in Pliny and Tacitus have been very influential for scholarship on Germanic history and language up until recent times. However, outside of Tacitus and Pliny there are no other textual indications that these groups were important. The subgroups mentioned by Tacitus are not used by him elsewhere in his work, contradict other parts of his work, and cannot be reconciled with Pliny, who
19256-591: The distance) covered a region roughly located between the Rhine , the Vistula , the Danube , and southern Scandinavia during the first two centuries of the Common Era . East Germanic speakers dwelled on the Baltic sea coasts and islands, while speakers of the Northwestern dialects occupied territories in present-day Denmark and bordering parts of Germany at the earliest date when they can be identified. In
19422-488: The disunited eastern Empire submitted to some of his demands, possibly giving him control over Epirus . In the aftermath of the large-scale Gothic entries into the empire, the Franks and Alemanni became more secure in their positions in 395, when Stilicho , the barbarian generalissimo who held power in the western Empire, made agreements with them. In 401, Alaric invaded Italy, coming to an understanding with Stilicho in 404/5. This agreement allowed Stilicho to fight against
19588-573: The early Germans were also highly influential among members of the nationalist and racist völkisch movement and later co-opted by the Nazis . During the second half of the 20th century, the controversial misuse of ancient Germanic history and archaeology was discredited and has since resulted in a backlash against many aspects of earlier scholarship. The etymology of the Latin word Germani , from which Latin Germania and English Germanic are derived,
19754-475: The empire, laying siege to Philippopolis . He followed his victory there with another on the marshy terrain at Abrittus , a battle which cost the life of Roman emperor Decius . In 253/254, further attacks occurred reaching Thessalonica and possibly Thrace . In 267/268 there were large raids led by the Herules in 267/268, and a mixed group of Goths and Herules in 269/270. Gothic attacks were abruptly ended in
19920-400: The end of the war with the Chatti, Domitian reduced the number of Roman soldiers on the upper Rhine and shifted the Roman military to guarding the Danube frontier, beginning the construction of the limes , the longest fortified border in the empire. The period afterwards was peaceful enough that the emperor Trajan reduced the number of soldiers on the frontier. According to Edward James ,
20086-534: The extant wolf C. lupus from the earlier C. mosbachensis (which in turn descended from C. etruscus ) is widely accepted. Among the oldest fossils of the modern grey wolf is from Ponte Galeria in Italy, dating to 406,500 ± 2,400 years ago. Remains from Cripple Creek Sump in Alaska may be considerably older, around 1 million years old, though differentiating between the remains of modern wolves and C. mosbachensis
20252-482: The fear of wolves exists in many human societies, the majority of recorded attacks on people have been attributed to animals suffering from rabies . Wolf attacks on humans are rare because wolves are relatively few, live away from people, and have developed a fear of humans because of their experiences with hunters, farmers, ranchers, and shepherds. The English "wolf" stems from the Old English wulf , which
20418-465: The first arrival of dogs across the Bering Strait 12,000 to 14,000 years ago. Research into the inheritance of white colour from dogs into wolves has yet to be undertaken. Wolves occur across Eurasia and North America. However, deliberate human persecution because of livestock predation and fear of attacks on humans has reduced the wolf's range to about one-third of its historic range; the wolf
20584-560: The first of them was Maroboduus of the Marcomanni, who had led his people away from the Roman activities into Bohemia , which was defended by forests and mountains, and had formed alliances with other peoples. In 6 CE, Rome planned an attack against him but the campaign was cut short when forces were needed for the Illyrian revolt in the Balkans. Just three years later (9 CE), the second of these Germanic figures, Arminius of
20750-450: The following years Caesar pursued a controversial campaign to conquer all of Gaul on behalf of Rome, establishing the Rhine as a border. In 55 BCE he crossed the Rhine into Germania near Cologne . Near modern Nijmegen he also massacred a large migrating group of Tencteri and Usipetes who had crossed the Rhine from the east. Throughout the reign of Augustus—from 27 BCE until 14 CE—the Roman empire expanded into Gaul, with
20916-554: The force of Radagaisus , who had crossed the Middle Danube in 405/6 and invaded Italy, only to be defeated outside Florence. That same year, a large force of Vandals, Suevi, Alans, and Burgundians crossed the Rhine , fighting the Franks but facing no Roman resistance. In 409, the Suevi, Vandals, and Alans crossing the Pyrenees into Spain, where they took possession of the northern part of the peninsula. The Burgundians seized
21082-834: The hind feet are 220–250 mm ( 8 + 5 ⁄ 8 – 9 + 7 ⁄ 8 in). The size and weight of the modern wolf increases proportionally with latitude in accordance with Bergmann's rule . The mean body mass of the wolf is 40 kg (88 lb), the smallest specimen recorded at 12 kg (26 lb) and the largest at 79.4 kg (175 lb). On average, European wolves weigh 38.5 kg (85 lb), North American wolves 36 kg (79 lb), and Indian and Arabian wolves 25 kg (55 lb). Females in any given wolf population typically weigh 2.3–4.5 kg (5–10 lb) less than males. Wolves weighing over 54 kg (119 lb) are uncommon, though exceptionally large individuals have been recorded in Alaska and Canada. In central Russia, exceptionally large males can reach
21248-523: The land around modern Speyer , Worms , and Strasbourg, territory that was recognized by the Roman Emperor Honorius . When Stilicho fell from power in 408, Alaric invaded Italy again and eventually sacked Rome in 410; Alaric died shortly thereafter. The Visigoths withdrew into Gaul where they faced a power struggle until the succession of Wallia in 415 and his son Theodoric I in 417/18. Following successful campaigns against them by
21414-480: The later diffusion of local dialectal innovations. The Germanic-speaking peoples speak an Indo-European language . The leading theory for the origin of Germanic languages, suggested by archaeological, linguistic and genetic evidence, postulates a diffusion of Indo-European languages from the Pontic–Caspian steppe towards Northern Europe during the third millennium BCE, via linguistic contacts and migrations from
21580-478: The lineage that led to dogs from other Old World wolves around 11,100–12,300 years ago. An extinct Late Pleistocene wolf may have been the ancestor of the dog, with the dog's similarity to the extant wolf being the result of genetic admixture between the two. The dingo, Basenji , Tibetan Mastiff and Chinese indigenous breeds are basal members of the domestic dog clade. The divergence time for wolves in Europe,
21746-424: The middle of the back is 60–70 mm ( 2 + 3 ⁄ 8 – 2 + 3 ⁄ 4 in), and the guard hairs on the shoulders generally do not exceed 90 mm ( 3 + 1 ⁄ 2 in), but can reach 110–130 mm ( 4 + 3 ⁄ 8 – 5 + 1 ⁄ 8 in). A wolf's coat colour is determined by its guard hairs. Wolves usually have some hairs that are white, brown, gray and black. The coat of
21912-680: The migration period is usually set at 568 when the Lombards invaded Italy. During this time period, numerous barbarian groups invaded the Roman Empire and established new kingdoms within its boundaries. These Germanic migrations traditionally mark the transition between antiquity and the beginning of the early Middle Ages . The reasons for the migrations of the period are unclear, but scholars have proposed overpopulation, climate change, bad harvests, famines, and adventurousness as possible reasons. Migrations were probably carried out by relatively small groups rather than entire peoples. The Greuthungi ,
22078-527: The modern spotted hyena . Genomic studies suggest modern wolves and dogs descend from a common ancestral wolf population. A 2021 study found that the Himalayan wolf and the Indian plains wolf are part of a lineage that is basal to other wolves and split from them 200,000 years ago. Other wolves appear to share most of their common ancestry much more recently, within the last 23,000 years (around
22244-593: The modern wolf and the dog are sister taxa , as modern wolves are not closely related to the population of wolves that was first domesticated . In 2019, a workshop hosted by the IUCN /Species Survival Commission's Canid Specialist Group considered the New Guinea singing dog and the dingo to be feral Canis familiaris , and therefore should not be assessed for the IUCN Red List . The phylogenetic descent of
22410-400: The months of February and May. Adoptee males may mate with an available pack female and then form their own pack. In some cases, a lone wolf is adopted into a pack to replace a deceased breeder. Wolves are territorial and generally establish territories far larger than they require to survive assuring a steady supply of prey. Territory size depends largely on the amount of prey available and
22576-958: The most frequently consumed prey of North American wolves. Wolves can digest their meal in a few hours and can feed several times in one day, making quick use of large quantities of meat. A well-fed wolf stores fat under the skin, around the heart, intestines, kidneys, and bone marrow, particularly during the autumn and winter. Nonetheless, wolves are not fussy eaters. Smaller-sized animals that may supplement their diet include rodents , hares , insectivores and smaller carnivores. They frequently eat waterfowl and their eggs. When such foods are insufficient, they prey on lizards , snakes , frogs , and large insects when available. Wolves in some areas may consume fish and even marine life. Wolves also consume some plant material. In Europe, they eat apples, pears, figs , melons, berries and cherries . In North America, wolves eat blueberries and raspberries . They also eat grass, which may provide some vitamins, but
22742-417: The neck. The hairs on the cheeks are elongated and form tufts. The ears are covered in short hairs and project from the fur. Short, elastic and closely adjacent hairs are present on the limbs from the elbows down to the calcaneal tendons . The winter fur is highly resistant to the cold. Wolves in northern climates can rest comfortably in open areas at −40 °C (−40 °F) by placing their muzzles between
22908-478: The number of worms and the age of the host. The wolf is a social animal . Its populations consist of packs and lone wolves, most lone wolves being temporarily alone while they disperse from packs to form their own or join another one. The wolf's basic social unit is the nuclear family consisting of a mated pair accompanied by their offspring. The average pack size in North America is eight wolves and 5.5 in Europe. The average pack across Eurasia consists of
23074-451: The pack for 10–54 months before dispersing. Triggers for dispersal include the onset of sexual maturity and competition within the pack for food. The distance travelled by dispersing wolves varies widely; some stay in the vicinity of the parental group, while other individuals may travel great distances of upwards of 206 km (128 mi), 390 km (240 mi), and 670 km (420 mi) from their natal (birth) packs. A new pack
23240-459: The pack usually before and after hunts, to pass on an alarm particularly at a den site, to locate each other during a storm, while crossing unfamiliar territory, and to communicate across great distances. Wolf howls can under certain conditions be heard over areas of up to 130 km (50 sq mi). Other vocalizations include growls , barks and whines. Wolves do not bark as loudly or continuously as dogs do in confrontations, rather barking
23406-466: The pads come in contact with ice and snow. In warm climates, the fur is coarser and scarcer than in northern wolves. Female wolves tend to have smoother furred limbs than males and generally develop the smoothest overall coats as they age. Older wolves generally have more white hairs on the tip of the tail, along the nose, and on the forehead. Winter fur is retained longest by lactating females, although with some hair loss around their teats. Hair length on
23572-406: The peak and the end of the Last Glacial Maximum ), originating from Siberia or Beringia . While some sources have suggested that this was a consequence of a population bottleneck , other studies have suggested that this a result of gene flow homogenising ancestry. A 2016 genomic study suggests that Old World and New World wolves split around 12,500 years ago followed by the divergence of
23738-453: The possibility of fully integrating this region into the empire. Rome launched successful campaigns across the Rhine between 14 and 16 CE under Tiberius and Germanicus, but the effort of integrating Germania now seemed to outweigh its benefits. In the reign of Augustus's successor, Tiberius, it became state policy to expand the empire no further than the frontier based roughly upon the Rhine and Danube, recommendations that were specified in
23904-401: The principal causes of wolf mortality, one study concluding that 14–65% of wolf deaths in Minnesota and the Denali National Park and Preserve were due to other wolves. Wolves communicate using vocalizations, body postures, scent, touch, and taste. The phases of the moon have no effect on wolf vocalization, and despite popular belief, wolves do not howl at the Moon. Wolves howl to assemble
24070-401: The province. Despite the peace, the Suevi expanded their territory by conquering Mérida in 439 and Seville in 441. By 440, Attila and the Huns had come to rule a multi-ethnic empire north of the Danube; two of the most important peoples within this empire were the Gepids and the Goths. The Gepid king Ardaric came to power around 440 and participated in various Hunnic campaigns. In 450,
24236-410: The rear legs and covering their faces with their tail. Wolf fur provides better insulation than dog fur and does not collect ice when warm breath is condensed against it. In cold climates, the wolf can reduce the flow of blood near its skin to conserve body heat. The warmth of the foot pads is regulated independently from the rest of the body and is maintained at just above tissue-freezing point where
24402-474: The same extent as the coyote, whose diet contains more vegetable matter. Females tend to have narrower muzzles and foreheads, thinner necks, slightly shorter legs, and less massive shoulders than males. Adult wolves measure 105–160 cm (41–63 in) in length and 80–85 cm (31–33 in) at shoulder height. The tail measures 29–50 cm (11–20 in) in length, the ears 90–110 mm ( 3 + 1 ⁄ 2 – 4 + 3 ⁄ 8 in) in height, and
24568-552: The second half of winter and lasts for two weeks. Dens are usually constructed for pups during the summer period. When building dens, females make use of natural shelters like fissures in rocks, cliffs overhanging riverbanks and holes thickly covered by vegetation. Sometimes, the den is the appropriated burrow of smaller animals such as foxes, badgers or marmots. An appropriated den is often widened and partly remade. On rare occasions, female wolves dig burrows themselves, which are usually small and short with one to three openings. The den
24734-433: The skeletons of large animals. Raised leg urination is considered to be one of the most important forms of scent communication in the wolf, making up 60–80% of all scent marks observed. Wolves are monogamous , mated pairs usually remaining together for life. Should one of the pair die, another mate is found quickly. With wolves in the wild, inbreeding does not occur where outbreeding is possible. Wolves become mature at
24900-444: The term Germanic due to its broad recognizability. Archaeologist Heiko Steuer defines his own work on the Germani in geographical terms (covering Germania ), rather than in ethnic terms. He nevertheless argues for some sense of shared identity between the Germani , noting the use of a common language, a common runic script , various common objects of material culture such as bracteates and gullgubber (small gold objects) and
25066-436: The term's total abandonment as a modern construct, since lumping "Germanic peoples" together implies a common group identity for which there is little evidence. Other scholars have defended the term's continued use and argue that a common Germanic language allows one to speak of "Germanic peoples", regardless of whether these ancient and medieval peoples saw themselves as having a common identity. Scholars generally agree that it
25232-551: The territory's periphery. Wolves tend to avoid hunting on the fringes of their range to avoid fatal confrontations with neighbouring packs. The smallest territory on record was held by a pack of six wolves in northeastern Minnesota, which occupied an estimated 33 km (13 sq mi), while the largest was held by an Alaskan pack of ten wolves encompassing 6,272 km (2,422 sq mi). Wolf packs are typically settled, and usually leave their accustomed ranges only during severe food shortages. Territorial fights are among
25398-410: The tribal names in Tacitus's account and the name of Mannus himself suggest that the descent from Mannus was an authentic Germanic tradition. All Germanic languages derive from the Proto-Indo-European language (PIE), which is generally thought to have been spoken between 4500 and 2500 BCE. The ancestor of Germanic languages is referred to as Proto- or Common Germanic , and likely represented
25564-423: The two species interact. One-to-one, hyenas dominate wolves, and may prey on them, but wolf packs can drive off single or outnumbered hyenas. There is at least one case in Israel of a hyena associating and cooperating with a wolf pack. Viral diseases carried by wolves include: rabies , canine distemper , canine parvovirus , infectious canine hepatitis , papillomatosis , and canine coronavirus . In wolves,
25730-595: The upper Rhine and are mentioned in Roman sources from the third century onward. The Goths begin to be mentioned along the lower Danube, where they attacked the city of Histria in 238. The Franks are first mentioned occupying territory between the Rhine and Weser. The Lombards seem to have moved their center of power to the central Elbe. Groups such as the Alamanni, Goths, and Franks were not unified polities; they formed multiple, loosely associated groups, who often fought each other and some of whom sought Roman friendship. The Romans also begin to mention seaborne attacks by
25896-404: The vulnerable individuals of large prey, with a pack of 15 able to bring down an adult moose . The variation in diet between wolves living on different continents is based on the variety of hoofed mammals and of available smaller and domesticated prey. In North America, the wolf's diet is dominated by wild large hoofed mammals (ungulates) and medium-sized mammals. In Asia and Europe, their diet
26062-400: The war by 180, through a combination of Roman military victories, the resettling of some peoples on Roman territory, and by making alliances with others. Marcus Aurelius's successor Commodus chose not to permanently occupy any territory conquered north of the Danube, and the following decades saw an increase in the defenses at the limes . The Romans renewed their right to choose the kings of
26228-423: The will of Augustus and read aloud by Tiberius himself. Roman intervention in Germania led to a shifting and unstable political situation, in which pro- and anti-Roman parties vied for power. Arminius was murdered in 21 CE by his fellow Germanic tribesmen, due in part to these tensions and for his attempt to claim supreme kingly power for himself. In the wake of Arminius's death, Roman diplomats sought to keep
26394-431: The wolf because of its "cauda recurvata" (upturning tail) which is not found in any other canid . In the third edition of Mammal Species of the World published in 2005, the mammalogist W. Christopher Wozencraft listed under C. lupus 36 wild subspecies, and proposed two additional subspecies: familiaris (Linnaeus, 1758) and dingo (Meyer, 1793). Wozencraft included hallstromi —the New Guinea singing dog —as
26560-468: The wolf's intestines. Tapeworms are commonly found in wolves, which they get though their prey, and generally cause little harm in wolves, though this depends on the number and size of the parasites, and the sensitivity of the host. Symptoms often include constipation , toxic and allergic reactions , irritation of the intestinal mucosa , and malnutrition . Wolves can carry over 30 roundworm species, though most roundworm infections appear benign, depending on
26726-462: The word sapo ('hair dye') is certainly borrowed from Proto-Germanic * saipwōn- (English soap ) , as evidenced by the parallel Finnish loanword saipio . The name of the framea , described by Tacitus as a short spear carried by Germanic warriors, most likely derives from the compound * fram-ij-an- ('forward-going one'), as suggested by comparable semantical structures found in early runes (e.g., raun-ij-az 'tester', on
26892-443: The work of the "Toronto School" around Walter Goffart , various scholars have denied that anything such as a common Germanic ethnic identity ever existed. Such scholars argue that most ideas about Germanic culture are taken from far later epochs and projected backwards to antiquity. Historians of the Vienna School, such as Walter Pohl , have also called for the term to be avoided or used with careful explanation, and argued that there
27058-423: The years after 270, after a Roman victory in which the Gothic king Cannabaudes was killed. The Roman limes largely collapsed in 259/260, during the Crisis of the Third Century (235–284), and Germanic raids penetrated as far as northern Italy. The limes on the Rhine and upper Danube was brought under control again in 270s, and by 300 the Romans had reestablished control over areas they had abandoned during
27224-423: Was a characteristic, but not defining feature of the Germanic peoples. Many of the ascribed ethnic characteristics of the Germani represented them as typically "barbarian", including the possession of stereotypical vices such as "wildness" and of virtues such as chastity. Tacitus was at times unsure whether a people were Germanic or not. He expressed uncertainty about the Peucini , who he says spoke and lived like
27390-462: Was able to show strength by having a Roman fleet enter the Elbe and meet the legions in the heart of Germania . Once Tiberius subdued the Germanic people between the Rhine and the Elbe, the region at least up to Weser —and possibly up to the Elbe —was made the Roman province Germania and provided soldiers to the Roman army. However, within this period two Germanic kings formed larger alliances. Both of them had spent some of their youth in Rome;
27556-411: Was discovered through mitochondrial DNA analysis of jackals living in the Caucasus Mountains and in Bulgaria. In 2021, a genetic study found that the dog's similarity to the extant gray wolf was the result of substantial dog-into-wolf gene flow , with little evidence of the reverse. The wolf is the largest extant member of the family Canidae , and is further distinguished from coyotes and jackals by
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