Misplaced Pages

#795204

107-494: Observations about Cetacea have been recorded since at least classical times. Ancient Greek fishermen created an artificial notch on the dorsal fin of dolphins entangled in nets so that they could tell them apart years later. Approximately 2,300 years ago, Aristotle carefully took notes on cetaceans while traveling on boats with fishermen in the Aegean Sea . In his book Historia animalium ( History of Animals ), Aristotle

214-546: A 'strongman' was often the best solution. Athens fell under a tyranny in the second half of the 6th century BC. When this tyranny was ended, the Athenians founded the world's first democracy as a radical solution to prevent the aristocracy regaining power. A citizens' assembly (the Ecclesia ), for the discussion of city policy, had existed since the reforms of Draco in 621 BC; all citizens were permitted to attend after

321-412: A build-up of gases inside the gut, prompting the sphincter to relieve the pressure and release the feces. After an animal has digested eaten material, the remains of that material are discharged from its body as waste. Although it is lower in energy than the food from which it is derived, feces may retain a large amount of energy, often 50% of that of the original food. This means that of all food eaten,

428-417: A century into the past, discussing 6th century BC historical figures such as Darius I of Persia , Cambyses II and Psamtik III , and alluding to some 8th century BC persons such as Candaules . The accuracy of Herodotus' works is debated. Herodotus was succeeded by authors such as Thucydides , Xenophon , Demosthenes , Plato and Aristotle . Most were either Athenian or pro-Athenian, which

535-500: A coalition of 31 Greek city states, including Athens and Sparta, determined to resist the Persian invaders. At the same time, Greek Sicily was invaded by a Carthaginian force. In 480 BC, the first major battle of the invasion was fought at Thermopylae , where a small rearguard of Greeks, led by three hundred Spartans, held a crucial pass guarding the heart of Greece for several days; at the same time Gelon , tyrant of Syracuse, defeated

642-609: A council of elders (the Gerousia ) and magistrates specifically appointed to watch over the kings (the Ephors ). Only free, land-owning, native born men could be citizens entitled to the full protection of the law in a city-state. In most city-states, unlike the situation in Rome , social prominence did not allow special rights. Sometimes families controlled public religious functions, but this ordinarily did not give any extra power in

749-528: A decisive victory, and in 447 lost Boeotia again. Athens and Sparta signed the Thirty Years' Peace in the winter of 446/5, ending the conflict. Despite the treaty, Athenian relations with Sparta declined again in the 430s, and in 431 BC the Peloponnesian War began. The first phase of the war saw a series of fruitless annual invasions of Attica by Sparta, while Athens successfully fought

856-433: A deterrent for humans, as consuming or touching it may result in sickness or infection. Feces are discharged through the anus or cloaca during defecation . This process requires pressures that may reach 100 millimetres of mercury (3.9 inHg) (13.3 kPa) in humans and 450 millimetres of mercury (18 inHg) (60 kPa) in penguins. The forces required to expel the feces are generated through muscular contractions and

963-436: A group of city-states allied themselves to defend Greece, the vast majority of poleis remained neutral, and after the Persian defeat, the allies quickly returned to infighting. Thus, the major peculiarities of the ancient Greek political system were its fragmented nature (and that this does not particularly seem to have tribal origin), and the particular focus on urban centers within otherwise tiny states. The peculiarities of

1070-437: A human fingerprint; no one animal's appearance exactly like another's. Once Biggs and Ellis found they could recognize certain individuals, they realised that the animals travel in stable groups called pods. Researchers use photo identification to identify specific individuals and pods. The photographic system has also worked well in humpback whale studies. Researchers use the color of the pectoral fins and color and scarring of

1177-425: A job but to become an effective citizen. Girls also learned to read, write and do simple arithmetic so they could manage the household. They almost never received education after childhood. Feces Feces ( or faeces ; sg. : faex ) are the solid or semi-solid remains of food that was not digested in the small intestine , and has been broken down by bacteria in the large intestine . Feces contain

SECTION 10

#1732797947796

1284-655: A loose collection of culturally and linguistically related city-states and other territories. Prior to the Roman period , most of these regions were officially unified once under the Kingdom of Macedon from 338 to 323 BC. In Western history , the era of classical antiquity was immediately followed by the Early Middle Ages and the Byzantine period. Three centuries after the decline of Mycenaean Greece during

1391-409: A plant in having fruit is that animals will eat the fruit and unknowingly disperse the seed in doing so. This mode of seed dispersal is highly successful, as seeds dispersed around the base of a plant are unlikely to succeed and often are subject to heavy predation . Provided the seed can withstand the pathway through the digestive system, it is not only likely to be far away from the parent plant, but

1498-414: A relatively small amount of metabolic waste products such as bacterially altered bilirubin , and dead epithelial cells from the lining of the gut. Feces are discharged through the anus or cloaca during defecation . Feces can be used as fertilizer or soil conditioner in agriculture. They can also be burned as fuel or dried and used for construction . Some medicinal uses have been found. In

1605-760: A result of Epaminondas ' liberation of Messenia from Spartan rule, the helot system there came to an end and the helots won their freedom. However, it did continue to persist in Laconia until the 2nd century BC. For most of Greek history, education was private, except in Sparta. During the Hellenistic period, some city-states established public schools . Only wealthy families could afford a teacher. Boys learned how to read, write and quote literature. They also learned to sing and play one musical instrument and were trained as athletes for military service. They studied not for

1712-477: A significant amount of energy remains for the decomposers of ecosystems. Many organisms feed on feces, from bacteria to fungi to insects such as dung beetles , who can sense odors from long distances. Some may specialize in feces, while others may eat other foods. Feces serve not only as a basic food, but also as a supplement to the usual diet of some animals. This process is known as coprophagia , and occurs in various animal species such as young elephants eating

1819-523: A special type of slaves called helots . Helots were Messenians enslaved en masse during the Messenian Wars by the state and assigned to families where they were forced to stay. Helots raised food and did household chores so that women could concentrate on raising strong children while men could devote their time to training as hoplites . Their masters treated them harshly, and helots revolted against their masters several times. In 370/69 BC, as

1926-470: A valuable purpose in paleontology because they provide direct evidence of the predation and diet of extinct organisms. Coprolites may range in size from a few millimetres to more than 60 centimetres. Palaeofeces are ancient feces , often found as part of archaeological excavations or surveys. Intact paleofeces of ancient people may be found in caves in arid climates and in other locations with suitable preservation conditions. These are studied to determine

2033-722: A war with the Roman Republic in the late 3rd century. Although the First Macedonian War was inconclusive, the Romans, in typical fashion, continued to fight Macedon until it was completely absorbed into the Roman Republic (by 149 BC). In the east, the unwieldy Seleucid Empire gradually disintegrated, although a rump survived until 64 BC, whilst the Ptolemaic Kingdom continued in Egypt until 30 BC when it too

2140-684: A year, the Thirty had been overthrown. The first half of the fourth century saw the major Greek states attempt to dominate the mainland; none were successful, and their resulting weakness led to a power vacuum which would eventually be filled by Macedon under Philip II and then Alexander the Great. In the immediate aftermath of the Peloponnesian war, Sparta attempted to extend their own power, leading Argos, Athens, Corinth, and Thebes to join against them. Aiming to prevent any single Greek state gaining

2247-512: Is among the world's most expensive coffees. Paper is also made from elephant dung in Thailand. Haathi Chaap is a brand of paper made from elephant dung. Dog feces was used in the tanning process of leather during the Victorian era . Collected dog feces, known as "pure", "puer", or "pewer", were mixed with water to form a substance known as "bate", because proteolytic enzymes in

SECTION 20

#1732797947796

2354-536: Is burned as fuel in many countries. Animals such as the giant panda and zebra possess gut bacteria capable of producing biofuel. The bacterium in question, Brocadia anammoxidans , can be used to synthesize the rocket fuel hydrazine . A coprolite is fossilized feces and is classified as a trace fossil . In paleontology they give evidence about the diet of an animal. They were first described by William Buckland in 1829. Prior to this, they were known as "fossil fir cones " and " bezoar stones". They serve

2461-494: Is called constipation . The appearance of human fecal matter varies according to diet and health. Normally it is semisolid, with a mucus coating. A combination of bile and bilirubin , which comes from dead red blood cells , gives feces the typical brown color. After the meconium , the first stool expelled, a newborn's feces contains only bile , which gives it a yellow-green color. Breast feeding babies expel soft, pale yellowish, and not quite malodorous matter; but once

2568-427: Is even provided with its own fertilizer. Organisms that subsist on dead organic matter or detritus are known as detritivores , and play an important role in ecosystems by recycling organic matter back into a simpler form that plants and other autotrophs may absorb once again. This cycling of matter is known as the biogeochemical cycle . To maintain nutrients in soil it is therefore important that feces returns to

2675-558: Is generally considered the cradle of Western civilization , the seminal culture from which the modern West derives many of its founding archetypes and ideas in politics, philosophy, science, and art. Classical antiquity in the Mediterranean region is commonly considered to have begun in the 8th century BC (around the time of the earliest recorded poetry of Homer) and ended in the 6th century AD. Classical antiquity in Greece

2782-458: Is mountainous, and as a result, ancient Greece consisted of many smaller regions, each with its own dialect, cultural peculiarities, and identity. Regionalism and regional conflicts were prominent features of ancient Greece. Cities tended to be located in valleys between mountains, or on coastal plains, and dominated a certain area around them. In the south lay the Peloponnese , consisting of

2889-535: Is no singular form, making the word a plurale tantum ; out of various major dictionaries, only one enters variation from plural agreement . "Feces" is used more in biology and medicine than in other fields (reflecting science 's tradition of classical Latin and Neo-Latin ) There are many synonyms in informal registers for feces, just like there are for urine . Many are euphemistic , colloquial , or both; some are profane (such as shit ), whereas most belong chiefly to child-directed speech (such as poo or

2996-499: Is photo-identification. This system was popularized by Michael Bigg , a pioneer in modern orca (killer whale) research. During the mid-1970s, Bigg and Graeme Ellis photographed local orcas in the British Columbian seas. After examining the photos, they realized they could recognize certain individual whales by looking at the shape and condition of the dorsal fin, and also the shape of the saddle patch. These are as unique as

3103-470: Is relatively high in paleofeces, making it more reliable than skeletal DNA retrieval. The reason this analysis is possible at all is due to the digestive system not being entirely efficient, in the sense that not everything that passes through the digestive system is destroyed. Not all of the surviving material is recognizable, but some of it is. Generally, this material is the best indicator archaeologists can use to determine ancient diets, as no other part of

3210-521: Is the scientific terminology, while the term stool is also commonly used in medical contexts. Outside of scientific contexts, these terms are less common, with the most common layman's term being poop or poo . The term shit is also in common use, although it is widely considered vulgar or offensive. There are many other terms, see below. The word faeces is the plural of the Latin word faex meaning "dregs". In most English-language usage , there

3317-408: Is through examination of dead carcasses that wash up on the shore. If properly collected and stored, these carcasses can provide important information that is difficult to obtain in field studies. In recent decades, methods of identifying individual cetaceans have enabled accurate population counts and insights into the life cycles and social structures of various species. One such successful system

Cetology - Misplaced Pages Continue

3424-479: Is unclear exactly how this change occurred. For instance, in Athens, the kingship had been reduced to a hereditary, lifelong chief magistracy ( archon ) by c. 1050 BC; by 753 BC this had become a decennial, elected archonship; and finally by 683 BC an annually elected archonship. Through each stage, more power would have been transferred to the aristocracy as a whole, and away from a single individual. Inevitably,

3531-440: Is unique in world history as the first period attested directly in comprehensive, narrative historiography , while earlier ancient history or protohistory is known from much more fragmentary documents such as annals, king lists, and pragmatic epigraphy . Herodotus is widely known as the "father of history": his Histories are eponymous of the entire field . Written between the 450s and 420s BC, Herodotus' work reaches about

3638-562: Is used as a cement to make adobe ( mudbrick ) huts, or even in throwing sports, especially with cow and camel dung. Kopi luwak , or civet coffee, is coffee made from coffee beans that have been eaten and excreted by Asian palm civets ( Paradoxurus hermaphroditus ). Giant pandas provide fertilizer for the world's most expensive green tea . In Malaysia , tea is made from the droppings of stick insects fed on guava leaves. In northern Thailand , elephants are used to digest coffee beans in order to make Black Ivory coffee , which

3745-723: Is usually counted from the Roman victory over the Corinthians at the Battle of Corinth in 146 BC to the establishment of Byzantium by Constantine as the capital of the Roman Empire in 330 AD. Finally, Late Antiquity refers to the period of Christianization during the later 4th to early 6th centuries AD, consummated by the closure of the Academy of Athens by Justinian I in 529. The historical period of ancient Greece

3852-471: Is why far more is known about the history and politics of Athens than of many other cities. Their scope is further limited by a focus on political, military and diplomatic history, ignoring economic and social history. The archaic period, lasting from approximately 800 to 500 BC, saw the culmination of political and social developments which had begun in the Greek dark age, with the polis (city-state) becoming

3959-655: The Achaemenid Empire by Alexander the Great spread Hellenistic civilization across the Middle East. The Hellenistic Period is considered to have ended in 30 BC, when the last Hellenistic kingdom, Ptolemaic Egypt , was annexed by the Roman Republic . Classical Greek culture , especially philosophy, had a powerful influence on ancient Rome , which carried a version of it throughout the Mediterranean and much of Europe. For this reason, Classical Greece

4066-710: The Archaic period , the Greek population grew beyond the capacity of the limited arable land of Greece proper, resulting in the large-scale establishment of colonies elsewhere: according to one estimate, the population of the widening area of Greek settlement increased roughly tenfold from 800 BC to 400 BC, from 800,000 to as many as 7 + 1 ⁄ 2 -10 million. This was not simply for trade, but also to found settlements. These Greek colonies were not, as Roman colonies were, dependent on their mother-city, but were independent city-states in their own right. Greeks settled outside of Greece in two distinct ways. The first

4173-647: The Battle of Aegospotami , and began to blockade Athens' harbour; driven by hunger, Athens sued for peace, agreeing to surrender their fleet and join the Spartan-led Peloponnesian League. Following the Athenian surrender, Sparta installed an oligarchic regime, the Thirty Tyrants , in Athens, one of a number of Spartan-backed oligarchies which rose to power after the Peloponnesian war. Spartan predominance did not last: after only

4280-745: The Boeotian League and finally to the League of Corinth led by Macedon . This period was shaped by the Greco-Persian Wars , the Peloponnesian War , and the Rise of Macedon . Following the Classical period was the Hellenistic period (323–146 BC), during which Greek culture and power expanded into the Near and Middle East from the death of Alexander until the Roman conquest. Roman Greece

4387-655: The Bronze Age Collapse , Greek urban poleis began to form in the 8th century BC, ushering in the Archaic period and the colonization of the Mediterranean Basin. This was followed by the age of Classical Greece , from the Greco-Persian Wars to the death of Alexander the Great in 323 BC, and which included the Golden Age of Athens and the Peloponnesian War . The unification of Greece by Macedon under Philip II and subsequent conquest of

Cetology - Misplaced Pages Continue

4494-576: The Delian League gradually transformed from a defensive alliance of Greek states into an Athenian empire, as Athens' growing naval power intimidated the other league states. Athens ended its campaigns against Persia in 450, after a disastrous defeat in Egypt in 454, and the death of Cimon in action against the Persians on Cyprus in 450. As the Athenian fight against the Persian empire waned, conflict grew between Athens and Sparta. Suspicious of

4601-740: The Paeonians due north, the Thracians to the northeast, and the Illyrians , with whom the Macedonians were frequently in conflict, to the northwest. Chalcidice was settled early on by southern Greek colonists and was considered part of the Greek world, while from the late 2nd millennium BC substantial Greek settlement also occurred on the eastern shores of the Aegean , in Anatolia . During

4708-441: The diet and health of the people who produced them through the analysis of seeds, small bones, and parasite eggs found inside. Feces may contain information about the person excreting the material as well as information about the material. They also may be analyzed chemically for more in-depth information on the individual who excreted them, using lipid analysis and ancient DNA analysis. The success rate of usable DNA extraction

4815-467: The dung of whales often floats and can be collected to tell important information about their diet and about the role they have in the environment. Often cetology involves waiting and paying close attention. Cetologists use equipment including hydrophones to listen to calls of communicating animals, binoculars and other optical devices for scanning the horizon, cameras, notes, and a few other devices and tools. An alternative method of studying cetaceans

4922-504: The palindromic word poop ) or to crude humor (such as crap , dump , load and turd .). The feces of animals often have special names (some of them are slang), for example: In all human cultures, feces elicit varying degrees of disgust in adults. Children under two years typically have no disgust response to it, suggesting it is culturally derived. Disgust toward feces appears to be strongest in cultures where flush toilets make olfactory contact with human feces minimal. Disgust

5029-449: The poleis grouped themselves into leagues, membership of which was in a constant state of flux. Later in the Classical period, the leagues would become fewer and larger, be dominated by one city (particularly Athens , Sparta and Thebes ); and often poleis would be compelled to join under threat of war (or as part of a peace treaty). Even after Philip II of Macedon conquered the heartlands of ancient Greece, he did not attempt to annex

5136-464: The 146 BC conquest of Greece after the Battle of Corinth. Macedonia became a Roman province while southern Greece came under the surveillance of Macedonia's prefect ; however, some Greek poleis managed to maintain a partial independence and avoid taxation. The Aegean Islands were added to this territory in 133 BC. Athens and other Greek cities revolted in 88 BC, and the peninsula was crushed by

5243-583: The Carthaginian invasion at the Battle of Himera . The Persians were decisively defeated at sea by a primarily Athenian naval force at the Battle of Salamis , and on land in 479 BC at the Battle of Plataea . The alliance against Persia continued, initially led by the Spartan Pausanias but from 477 by Athens, and by 460 Persia had been driven out of the Aegean. During this long campaign,

5350-525: The Corinthian empire in northwest Greece and defended its own empire, despite a plague which killed the leading Athenian statesman Pericles . The war turned after Athenian victories led by Cleon at Pylos and Sphakteria , and Sparta sued for peace, but the Athenians rejected the proposal. The Athenian failure to regain control of Boeotia at Delium and Brasidas ' successes in northern Greece in 424 improved Sparta's position after Sphakteria. After

5457-569: The Elder stated that they were fish, and it was followed by many naturalists . However, Pierre Belon (1517–1575) and G. Rondelet (1507–1566) persisted on believing they were mammals. They argued that the animals had lungs and a uterus , just like mammals. Not until 1758, when Swedish botanist Carl Linnaeus (1707–1778) published the tenth edition of Systema Naturae , were they seen as mammals. Only decades later, French zoologist and paleontologist Baron Georges Cuvier (1769–1832) described

SECTION 50

#1732797947796

5564-668: The Greek city-states. It greatly widened the horizons of the Greeks and led to a steady emigration of the young and ambitious to the new Greek empires in the east. Many Greeks migrated to Alexandria, Antioch and the many other new Hellenistic cities founded in Alexander's wake, as far away as present-day Afghanistan and Pakistan , where the Greco-Bactrian Kingdom and the Indo-Greek Kingdom survived until

5671-460: The Greek colonies Syracusae ( Συράκουσαι ), Neapolis ( Νεάπολις ), Massalia ( Μασσαλία ) and Byzantion ( Βυζάντιον ). These colonies played an important role in the spread of Greek influence throughout Europe and also aided in the establishment of long-distance trading networks between the Greek city-states, boosting the economy of ancient Greece . Ancient Greece consisted of several hundred relatively independent city-states ( poleis ). This

5778-578: The Greek colony Sybaris in southern Italy, its allies, and the Serdaioi. In 499 BC, the Ionian city states under Persian rule rebelled against their Persian-supported tyrant rulers. Supported by troops sent from Athens and Eretria , they advanced as far as Sardis and burnt the city before being driven back by a Persian counterattack. The revolt continued until 494, when the rebelling Ionians were defeated. Darius did not forget that Athens had assisted

5885-420: The Greek system are further evidenced by the colonies that they set up throughout the Mediterranean , which, though they might count a certain Greek polis as their 'mother' (and remain sympathetic to her), were completely independent of the founding city. Inevitably smaller poleis might be dominated by larger neighbors, but conquest or direct rule by another city-state appears to have been quite rare. Instead

5992-815: The Greeks began 250 years of expansion, settling colonies in all directions. To the east, the Aegean coast of Asia Minor was colonized first, followed by Cyprus and the coasts of Thrace , the Sea of Marmara and south coast of the Black Sea . Eventually, Greek colonization reached as far northeast as present-day Ukraine and Russia ( Taganrog ). To the west the coasts of Illyria , Southern Italy (called " Magna Graecia ") were settled, followed by Southern France , Corsica , and even eastern Spain . Greek colonies were also founded in Egypt and Libya . Modern Syracuse , Naples , Marseille and Istanbul had their beginnings as

6099-423: The Greeks were very aware of their tribal origins; Herodotus was able to extensively categorise the city-states by tribe. Yet, although these higher-level relationships existed, they seem to have rarely had a major role in Greek politics. The independence of the poleis was fiercely defended; unification was something rarely contemplated by the ancient Greeks. Even when, during the second Persian invasion of Greece,

6206-535: The Ionian revolt, and in 490 he assembled an armada to retaliate. Though heavily outnumbered, the Athenians—supported by their Plataean allies—defeated the Persian hordes at the Battle of Marathon , and the Persian fleet turned tail. Ten years later, a second invasion was launched by Darius' son Xerxes . The city-states of northern and central Greece submitted to the Persian forces without resistance, but

6313-483: The League of Corinth following the death of Philip, Alexander began his campaign against Persia in 334 BC. He conquered Persia, defeating Darius III at the Battle of Issus in 333 BC, and after the Battle of Gaugamela in 331 BC proclaimed himself king of Asia. From 329 BC he led expeditions to Bactria and then India; further plans to invade Arabia and North Africa were halted by his death in 323 BC. The period from

6420-634: The Roman general Sulla . The Roman civil wars devastated the land even further, until Augustus organized the peninsula as the province of Achaea in 27 BC. Greece was a key eastern province of the Roman Empire, as the Roman culture had long been in fact Greco-Roman . The Greek language served as a lingua franca in the East and in Italy , and many Greek intellectuals such as Galen would perform most of their work in Rome . The territory of Greece

6527-579: The ancient Greeks did not think in terms of race . Most families owned slaves as household servants and laborers, and even poor families might have owned a few slaves. Owners were not allowed to beat or kill their slaves. Owners often promised to free slaves in the future to encourage slaves to work hard. Unlike in Rome, freedmen did not become citizens. Instead, they were mixed into the population of metics , which included people from foreign countries or other city-states who were officially allowed to live in

SECTION 60

#1732797947796

6634-421: The animals as mammals without any hind legs. Skeletons were assembled and displayed in the first natural history museums , and on a closer look and comparisons with other extinct animal fossils, led zoologists to conclude that cetaceans came from a family of ancient land mammals. Between the 16th and 20th centuries, much of our information on cetaceans came from whalers . Whalers were the most knowledgeable about

6741-401: The animals, but their information was regarding migration routes and outer anatomy, and only little information of behavior. During the 1960s, people began studying the animals intensively, often in dedicated research institutes. The Tethys Institute of Milan, founded in 1986, compiled an extensive cetology database of the Mediterranean. This came from both concern about wild populations and also

6848-468: The archaeological record is so direct an indicator. A process that preserves feces in a way that they may be analyzed later is the Maillard reaction . This reaction creates a casing of sugar that preserves the feces from the elements. To extract and analyze the information contained within, researchers generally have to freeze the feces and grind it up into powder for analysis. Animal dung occasionally

6955-423: The area from which they came, which is not always the case in human society where food may be transported from rural areas to urban populations and then feces disposed of into a river or sea. Depending on the individual and the circumstances, human beings may defecate several times a day, every day, or once every two or three days. Extensive hardening of the feces that interrupts this routine for several days or more

7062-514: The baby begins to eat, and the body starts expelling bilirubin from dead red blood cells, its matter acquires the familiar brown color. At different times in their life, human beings will expel feces of different colors and textures. A stool that passes rapidly through the intestines will look greenish; lack of bilirubin will make the stool look like clay. The feces of animals, e.g. guano and manure , often are used as fertilizer . Dry animal dung , such as that of camel , bison and cattle ,

7169-444: The capture of larger animals such as the orca, and gaining popularity of dolphin shows in marine parks . Studying cetaceans presents numerous challenges. Cetaceans only spend 10% of their time on the surface, and all they do at the surface is breathe. There is very little behavior seen at the surface. It is also impossible to find any signs that an animal has been in an area. Cetaceans do not leave tracks that can be followed. However,

7276-431: The case of human feces , fecal transplants or fecal bacteriotherapy are in use. Urine and feces together are called excreta . The distinctive odor of feces is due to skatole , and thiols ( sulfur -containing compounds), as well as amines and carboxylic acids. Skatole is produced from tryptophan via indoleacetic acid. Decarboxylation gives skatole. The perceived bad odor of feces has been hypothesized to be

7383-666: The center, while in the east lay Boeotia , Attica , and Megaris . Northeast lay Thessaly , while Epirus lay to the northwest. Epirus stretched from the Ambracian Gulf in the south to the Ceraunian Mountains and the Aoos river in the north, and consisted of Chaonia (north), Molossia (center), and Thesprotia (south). In the northeast corner was Macedonia , originally consisting Lower Macedonia and its regions, such as Elimeia , Pieria , and Orestis . Around

7490-455: The course of the archaic period. Already in the seventh century, the right of all citizen men to attend the assembly appears to have been established. After a failed coup led by Cylon of Athens around 636 BC, Draco was appointed to establish a code of laws in 621. This failed to reduce the political tension between the poor and the elites, and in 594 Solon was given the authority to enact another set of reforms, which attempted to balance

7597-473: The course of the eighth and seventh century. According to Spartan tradition, this constitution was established by the legendary lawgiver Lycurgus . Over the course of the first and second Messenian wars , Sparta subjugated the neighbouring region of Messenia , enserfing the population. In the sixth century, Greek city-states began to develop formal relationships with one another, where previously individual rulers had relied on personal relationships with

7704-461: The death of Alexander the Great in 323 BC until the death of Cleopatra , the last Macedonian ruler of Egypt, is known as the Hellenistic period. In the early part of this period, a new form of kingship developed based on Macedonian and Near Eastern traditions. The first Hellenistic kings were previously Alexander's generals, and took power in the period following his death, though they were not part of existing royal lineages and lacked historic claims to

7811-552: The death of Alexander the Great in 323 BC. The Classical Period is characterized by a "classical" style, i.e. one which was considered exemplary by later observers, most famously in the Parthenon of Athens. Politically, the Classical Period was dominated by Athens and the Delian League during the 5th century, but displaced by Spartan hegemony during the early 4th century BC, before power shifted to Thebes and

7918-423: The deaths of Cleon and Brasidas, the strongest proponents of war on each side, a peace treaty was negoitiated in 421 by the Athenian general Nicias . The peace did not last, however. In 418 BC allied forces of Athens and Argos were defeated by Sparta at Mantinea . In 415 Athens launched an ambitious naval expedition to dominate Sicily; the expedition ended in disaster at the harbor of Syracuse , with almost

8025-508: The dog feces helped to relax the fibrous structure of the hide before the final stages of tanning. Dog feces collectors were known as pure finders . Elephants, hippos , koalas and pandas are born with sterile intestines, and require bacteria obtained from eating the feces of their mothers to digest vegetation. In India, cow dung and cow urine are major ingredients of the traditional Hindu drink Panchagavya . Politician Shankarbhai Vegad stated that they can cure cancer . Feces

8132-674: The dominance that would allow it to challenge Persia, the Persian king initially joined the alliance against Sparta, before imposing the Peace of Antalcidas ("King's Peace") which restored Persia's control over the Anatolian Greeks. By 371 BC, Thebes was in the ascendancy, defeating Sparta at the Battle of Leuctra , killing the Spartan king Cleombrotus I , and invading Laconia. Further Theban successes against Sparta in 369 led to Messenia gaining independence; Sparta never recovered from

8239-409: The domination of politics and concomitant aggregation of wealth by small groups of families was apt to cause social unrest in many poleis . In many cities a tyrant (not in the modern sense of repressive autocracies), would at some point seize control and govern according to their own will; often a populist agenda would help sustain them in power. In a system wracked with class conflict , government by

8346-726: The east to the Indian king Chandragupta Maurya in exchange for war elephants, and later lost large parts of Persia to the Parthian Empire . By the mid-third century, the kingdoms of Alexander's successors was mostly stable, though there continued to be disputes over border areas. The great capitals of Hellenistic culture were Alexandria in the Ptolemaic Kingdom and Antioch in the Seleucid Empire . The conquests of Alexander had numerous consequences for

8453-552: The elites of other cities. Towards the end of the archaic period, Sparta began to build a series of alliances, the Peloponnesian League , with cities including Corinth , Elis , and Megara , isolating Messenia and reinforcing Sparta's position against Argos , the other major power in the Peloponnese. Other alliances in the sixth century included those between Elis and Heraea in the Peloponnese; and between

8560-619: The end of the first century BC. The city-states within Greece formed themselves into two leagues; the Achaean League (including Corinth and Argos) and the Aetolian League (including Sparta and Athens). For much of the period until the Roman conquest, these leagues were at war, often participating in the conflicts between the Diadochi (the successor states to Alexander's empire). The Antigonid Kingdom became involved in

8667-609: The entire army killed, and the ships destroyed. Soon after the Athenian defeat in Syracuse, Athens' Ionian allies began to rebel against the Delian league, while Persia began to once again involve itself in Greek affairs on the Spartan side. Initially the Athenian position continued relatively strong, with important victories at Cyzicus in 410 and Arginusae in 406. However, in 405 the Spartan Lysander defeated Athens in

8774-540: The exact borders of the Hellenistic kingdoms were not settled. Antigonus attempted to expand his territory by attacking the other successor kingdoms until they joined against him, and he was killed at the Battle of Ipsus in 301 BC. His son Demetrius spent many years in Seleucid captivity, and his son, Antigonus II , only reclaimed the Macedonian throne around 276. Meanwhile, the Seleucid kingdom gave up territory in

8881-425: The feces of their mothers to gain essential gut flora , or by other animals such as dogs, rabbits, and monkeys. Feces and urine, which reflect ultraviolet light, are important to raptors such as kestrels , who can see the near ultraviolet and thus find their prey by their middens and territorial markers. Seeds also may be found in feces. Animals who eat fruit are known as frugivores . An advantage for

8988-487: The fluke to identify individuals. Scars from orca attacks found on the flukes of humpbacks are also used in identification. Ancient Greece Ancient Greece ( Ancient Greek : Ἑλλάς , romanized :  Hellás ) was a northeastern Mediterranean civilization, existing from the Greek Dark Ages of the 12th–9th centuries BC to the end of classical antiquity ( c.  600 AD ), that comprised

9095-469: The founding of Greek colonies around the Mediterranean, with Euboean settlements at Al-Mina in the east as early as 800 BC, and Ischia in the west by 775. Increasing contact with non-Greek peoples in this period, especially in the Near East, inspired developments in art and architecture, the adoption of coinage, and the development of the Greek alphabet. Athens developed its democratic system over

9202-576: The government. In Athens, the population was divided into four social classes based on wealth. People could change classes if they made more money. In Sparta, all male citizens were called homoioi , meaning "peers". However, Spartan kings, who served as the city-state's dual military and religious leaders, came from two families. Women in Ancient Greece appear to have primarily performed domestic tasks, managed households, and borne and reared children. Slaves had no power or status. Slaves had

9309-547: The increasing Athenian power funded by the Delian League, Sparta offered aid to reluctant members of the League to rebel against Athenian domination. These tensions were exacerbated in 462 BC when Athens sent a force to aid Sparta in overcoming a helot revolt, but this aid was rejected by the Spartans. In the 450s, Athens took control of Boeotia, and won victories over Aegina and Corinth. However, Athens failed to win

9416-482: The knowledge he had gained about cetaceans was lost, only to be re-discovered during the Renaissance . Many of the medieval texts on cetaceans come mainly from Scandinavia and Iceland, most came about the mid-13th century. One of the better known is Speculum Regale . This text describes various species that lived around the island of Iceland . It mentions orcs that had dog-like teeth and would demonstrate

9523-514: The loss of Messenia's fertile land and the helot workforce it provided. The rising power of Thebes led Sparta and Athens to join forces; in 362 they were defeated by Thebes at the Battle of Mantinea . In the aftermath of Mantinea, none of the major Greek states were able to dominate. Though Thebes had won the battle, their general Epaminondas was killed, and they spent the following decades embroiled in wars with their neighbours; Athens, meanwhile, saw its second naval alliance, formed in 377, collapse in

9630-495: The mid-350s. The power vacuum in Greece after the Battle of Mantinea was filled by Macedon, under Philip II . In 338 BC, he defeated a Greek alliance at the Battle of Chaeronea , and subsequently formed the League of Corinth . Philip planned to lead the League to invade Persia, but was murdered in 336 BC. His son Alexander the Great was left to fulfil his father's ambitions. After campaigns against Macedon's western and northern enemies, and those Greek states that had broken from

9737-443: The most important unit of political organisation in Greece. The absence of powerful states in Greece after the collapse of Mycenaean power, and the geography of Greece, where many settlements were separated from their neighbours by mountainous terrain, encouraged the development of small independent city-states. Several Greek states saw tyrants rise to power in this period, most famously at Corinth from 657 BC. The period also saw

9844-410: The power of the rich and the poor. In the middle of the sixth century, Pisistratus established himself as a tyrant, and after his death in 527 his son Hippias inherited his position; by the end of the sixth century he had been overthrown and Cleisthenes carried out further democratising reforms. In Sparta, a political system with two kings, a council of elders , and five ephors developed over

9951-617: The reforms of Solon (early 6th century), but the poorest citizens could not address the assembly or run for office. With the establishment of the democracy, the assembly became the de jure mechanism of government; all citizens had equal privileges in the assembly. However, non-citizens, such as metics (foreigners living in Athens) or slaves , had no political rights at all. After the rise of democracy in Athens, other city-states founded democracies. However, many retained more traditional forms of government. As so often in other matters, Sparta

10058-420: The regions of Laconia (southeast), Messenia (southwest), Elis (west), Achaia (north), Korinthia (northeast), Argolis (east), and Arcadia (center). These names survive to the present day as regional units of modern Greece , though with somewhat different boundaries. Mainland Greece to the north, nowadays known as Central Greece , consisted of Aetolia and Acarnania in the west, Locris , Doris , and Phocis in

10165-518: The right to have a family and own property, subject to their master's goodwill and permission, but they had no political rights. By 600 BC, chattel slavery had spread in Greece. By the 5th century BC, slaves made up one-third of the total population in some city-states. Between 40–80% of the population of Classical Athens were slaves. Slaves outside of Sparta almost never revolted because they were made up of too many nationalities and were too scattered to organize. However, unlike later Western culture ,

10272-589: The same kind of aggression towards other cetaceans as wild dogs would to other terrestrial animals . The text even illustrated the hunting technique of orcs , which are now called orcas. The Speculum Regale describes other cetaceans, including the sperm whale and narwhal . Many times they were seen as terrible monsters, such as killers of men, and destroyers of ships. They even bore odd names such as "pig whale", "horse whale", and "red whale". But not all creatures described were said to be fierce. Some were seen to be good, such as whales that drove shoals of herring towards

10379-538: The shore. This was seen as very helpful to fisherman. Many of the early studies were based on dead specimens and myth . The little information that was gathered was usually about length, and a rough outer body anatomy . Because these animals live in water their entire lives, early scientists did not have the technology to study these animals further. It was not until the 16th century that things would begin to change. Then cetaceans would be proved to be mammals rather than fish. Aristotle argued they were mammals. But Pliny

10486-448: The state. City-states legally owned slaves. These public slaves had a larger measure of independence than slaves owned by families, living on their own and performing specialized tasks. In Athens, public slaves were trained to look out for counterfeit coinage , while temple slaves acted as servants of the temple's deity and Scythian slaves were employed in Athens as a police force corralling citizens to political functions. Sparta had

10593-695: The territories they controlled. The most important of these rulers in the decades after Alexander's death were Antigonus I and his son Demetrius in Macedonia and the rest of Greece, Ptolemy in Egypt, and Seleucus I in Syria and the former Persian empire; smaller Hellenistic kingdoms included the Attalids in Anatolia and the Greco-Bactrian kingdom . In the early part of the Hellenistic period,

10700-515: The territory or unify it into a new province, but compelled most of the poleis to join his own Corinthian League . Initially many Greek city-states seem to have been petty kingdoms; there was often a city official carrying some residual, ceremonial functions of the king ( basileus ), e.g., the archon basileus in Athens. However, by the Archaic period and the first historical consciousness, most had already become aristocratic oligarchies . It

10807-626: The time of Alexander I of Macedon , the Argead kings of Macedon started to expand into Upper Macedonia , lands inhabited by independent Macedonian tribes like the Lyncestae , Orestae and the Elimiotae and to the west, beyond the Axius river , into Eordaia , Bottiaea , Mygdonia , and Almopia , regions settled by Thracian tribes. To the north of Macedonia lay various non-Greek peoples such as

10914-539: Was a notable exception to the rest of Greece, ruled through the whole period by not one, but two hereditary monarchs. This was a form of diarchy . The Kings of Sparta belonged to the Agiads and the Eurypontids, descendants respectively of Eurysthenes and Procles . Both dynasties' founders were believed to be twin sons of Aristodemus , a Heraclid ruler. However, the powers of these kings were held in check by both

11021-460: Was a situation unlike that in most other contemporary societies, which were either tribal or kingdoms ruling over relatively large territories. Undoubtedly, the geography of Greece —divided and sub-divided by hills, mountains, and rivers—contributed to the fragmentary nature of ancient Greece. On the one hand, the ancient Greeks had no doubt that they were "one people"; they had the same religion , same basic culture, and same language. Furthermore,

11128-430: Was careful enough to distinguish between the baleen whales and toothed whales , a taxonomical separation still used today. He also described the sperm whale and the common dolphin , stating that they can live for at least twenty-five or thirty years. His achievement was remarkable for its time, because even today it is very difficult to estimate the life-span of advanced marine animals. After Aristotle's death, much of

11235-752: Was conquered by the Romans. The Aetolian league grew wary of Roman involvement in Greece, and sided with the Seleucids in the Roman–Seleucid War ; when the Romans were victorious, the league was effectively absorbed into the Republic. Although the Achaean league outlasted both the Aetolian league and Macedon, it was also soon defeated and absorbed by the Romans in 146 BC, bringing Greek independence to an end. The Greek peninsula came under Roman rule during

11342-417: Was in permanent settlements founded by Greeks, which formed as independent poleis. The second form was in what historians refer to as emporia ; trading posts which were occupied by both Greeks and non-Greeks and which were primarily concerned with the manufacture and sale of goods. Examples of this latter type of settlement are found at Al Mina in the east and Pithekoussai in the west. From about 750 BC

11449-610: Was preceded by the Greek Dark Ages ( c.  1200 – c.  800 BC ), archaeologically characterised by the protogeometric and geometric styles of designs on pottery. Following the Dark Ages was the Archaic Period , beginning around the 8th century BC, which saw early developments in Greek culture and society leading to the Classical Period from the Persian invasion of Greece in 480 BC until

#795204