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112-556: Butchery is an ancient trade, whose duties may date back to the domestication of livestock ; its practitioners formed guilds in England as far back as 1272. Since the 20th century, many countries and local jurisdictions offer trade certifications for butchers in order to ensure quality, safety, and health standards but not all butchers have formal certification or training. Trade qualification in English-speaking countries

224-420: A "broad" definition: "a coevolutionary process that arises from a mutualism, in which one species (the domesticator) constructs an environment where it actively manages both the survival and reproduction of another species (the domesticate) in order to provide the former with resources and/or services." He comments that this adds niche construction to the activities of the domesticator. Domestication syndrome

336-569: A bill to distribute farmland to his veterans, and landless members of the urban poor. A similar measure had been rejected in 63 BC, which arguably made the Senate over confident in their ability to control popular unrest. Although Pompey could not overcome optimate opposition on his own, the situation changed when Marius' nephew Julius Caesar sought his endorsement for the consulship in 59 BC. A skilled, unscrupulous, and ambitious politician, Caesar used this alliance to harness Pompey's influence with

448-545: A centre of piracy, along with other inland areas and reorganised into six parts. These actions significantly increased Roman state income and presented Pompey with multiple opportunities to increase his personal wealth and patronage base. Before his return to Italy in 62 BC, Pompey paid his troops bonuses totalling around 16,000 talents , but despite fears he intended to follow Sulla's example, they were dismissed upon arrival at Brundisium. His journey to Rome drew huge crowds wherever he stopped, showing that although opinion in

560-544: A commander in the civil war of 83–81 BC . Pompey's success as a general while young enabled him to advance directly to his first consulship without following the traditional cursus honorum (the required steps to advance in a political career). He was elected as consul on three occasions (70, 55, 52 BC). He celebrated three triumphs , served as a commander in the Sertorian War , the Third Servile War ,

672-860: A few have been domesticated, including squid , cuttlefish and octopus , all used in research on behaviour and neurology . Terrestrial snails in the genera Helix are raised for food. Several parasitic or parasitoidal insects, including the fly Eucelatoria , the beetle Chrysolina , and the wasp Aphytis are raised for biological control. Conscious or unconscious artificial selection has many effects on species under domestication; variability can readily be lost by inbreeding, selection against undesired traits, or genetic drift, while in Drosophila , variability in eclosion time (when adults emerge) has increased. Humans foraged for wild cereals, seeds, and nuts thousands of years before they were domesticated; wild wheat and barley, for example, were gathered in

784-699: A fresh revolt in Caucasian Albania forced him to retrace his steps. Victory at the Abas enabled him to impose terms on the Albanians and agree truces with other tribes on the northern side of the Caucasus. Pompey then wintered in Armenia, settling minor border contests and raids between his allies Phraates and Tigranes. Relying on his naval blockade to wear down Mithridates, Pompey spent 64 BC annexing

896-484: A means of communication between far-flung places through the exploitation of the pigeon's homing instinct; research suggests it was domesticated as early as 10,000 years ago. Chicken fossils in China have been dated to 7,400 years ago. The chicken's wild ancestor is Gallus gallus , the red junglefowl of Southeast Asia. The species appears to have been kept initially for cockfighting rather than for food. Two insects ,

1008-468: A population bottleneck where genetic drift is enhanced by the small population size. Mutations can also be fixed in a population by a selective sweep . Mutational load can be increased by reduced selective pressure against moderately harmful traits when reproductive fitness is controlled by human management. However, there is evidence against a bottleneck in crops, such as barley, maize, and sorghum, where genetic diversity slowly declined rather than showing

1120-779: A predictable supply of a resource, resulting in mutual benefits . She noted further that it is not synonymous with agriculture since agriculture depends on domesticated organisms but does not automatically result from domestication. Michael D. Purugganan notes that domestication has been hard to define, despite the "instinctual consensus" that it means "the plants and animals found under the care of humans that provide us with benefits and which have evolved under our control." He comments that insects such as termites , ambrosia beetles , and leafcutter ants have domesticated some species of fungi , and notes further that other groups such as weeds and commensals have wrongly been called domesticated. Starting from Zeder's definition, Purugganan proposes

1232-425: A protracted coevolutionary process with multiple stages along different pathways. There are three proposed major pathways that most mammal domesticates followed into domestication: Humans did not intend to domesticate mammals from either the commensal or prey pathways, or at least they did not envision a domesticated animal would result from it. In both of those cases, humans became entangled with these species as

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1344-575: A rapid initial fall at the point of domestication. Further, the genetic diversity of these crops was regularly replenished from the natural population. Similar evidence exists for horses, pigs, cows, and goats. At least three groups of insects, namely ambrosia beetles, leafcutter ants, and fungus-growing termites, have domesticated species of fungi . Ambrosia beetles in the weevil subfamilies Scolytinae and Platypodinae excavate tunnels in dead or stressed trees into which they introduce fungal gardens, their sole source of nutrition. After landing on

1456-708: A rebellion in Gallia Narbonensis , after which his army entered winter quarters near Narbo Martius . In early 76 BC, he crossed the Col de Portet and entered the Iberian peninsula, where he would remain for the next five years. His arrival boosted the morale of Metellus' troops, while some rebels changed sides, but soon after he was defeated by Sertorius at the Battle of Lauron , losing one third of his army while inflicting next to no losses on Sertorius' army. This

1568-568: A series of measures, one of which was to appoint Pompey to a military command. While Lepidus continued south, Pompey raised troops from among his veterans in Picenum, and moved north to besiege Mutina , capital of Cisalpine Gaul. The town was held by Lepidus' ally Marcus Junius Brutus , who surrendered after a lengthy siege, and was assassinated next day, allegedly on Pompey's orders. Catulus then defeated Lepidus outside Rome, while Pompey marched against his rear, catching him near Cosa. Lepidus and

1680-418: A special senatorial decree. Plutarch suggests Pompey supported Crassus as his co-consul in order to put him under an obligation. The two men were elected consuls for 70 BC, but allegedly differed on almost every measure, rendering their term "politically barren and without achievement." However, their consulship did see the plebeian tribune recover powers removed by Sulla. One of the most significant

1792-457: A steady supply of resources, such as meat, milk, or labor. The process is gradual and geographically diffuse, based on trial and error. Domestication affected genes for behavior in animals, making them less aggressive. In plants, domestication affected genes for morphology, such as increasing seed size and stopping the shattering of cereal seedheads. Such changes both make domesticated organisms easier to handle and reduce their ability to survive in

1904-402: A suitable tree, an ambrosia beetle excavates a tunnel in which it releases its fungal symbiont . The fungus penetrates the plant's xylem tissue, extracts nutrients from it, and concentrates the nutrients on and near the surface of the beetle gallery. Ambrosia fungi are typically poor wood degraders and instead utilize less demanding nutrients. Symbiotic fungi produce and detoxify ethanol, which

2016-633: A wider variety of animal types, meat cuts and quality of cuts. Additionally, butcher shops may focus on a particular culture , or nationality , of meat production. Some butcher shops, termed "meat delis", may also include a delicatessen . In the United States and Canada, butcher shops have become less common because of the increasing popularity of supermarkets and warehouse clubs. Many remaining ones are aimed at Hispanic and other immigrants or, more recently, those looking for organic offerings. Supermarkets employ butchers for secondary butchery, but in

2128-474: A working animal. Among invertebrates , the silkworm and the western honey bee were domesticated over 5,000 years ago for silk and honey , respectively. The domestication of plants began around 13,000–11,000 years ago with cereals such as wheat and barley in the Middle East , alongside crops such as lentil , pea , chickpea , and flax . Beginning around 10,000 years ago, Indigenous peoples in

2240-536: Is fatback , which in Europe is an important primal cut of pork , but in North America is regarded as trimmings to be used in sausage or rendered into lard . The primal cuts may be sold complete or cut further. In various periods and cultures, the term "butcher" has been applied to people who act cruelly to other human beings or slaughter them. For example, Pompey , a prominent Roman general and politician of

2352-405: Is an attractant for ambrosia beetles and likely prevents the growth of antagonistic pathogens and selects for other beneficial symbionts. Ambrosia beetles mainly colonise wood of recently dead trees. The leafcutter ants are any of some 47 species of leaf-chewing ants in the genera Acromyrmex and Atta . The ants carry the discs of leaves that they have cut back to their nest, where they feed

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2464-432: Is domesticated wheat, which relies on farmers for its reproduction and dissemination. Domesticated plants differ from their wild relatives in many ways, including Plant defenses against herbivory , such as thorns, spines, and prickles , poison, protective coverings, and sturdiness may have been reduced in domesticated plants. This would make them more likely to be eaten by herbivores unless protected by humans, but there

2576-403: Is often earned through an apprenticeship although some training organisations also certify their students. In Canada, once a butcher is trade qualified, they can learn to become a master butcher (Fleishmaster). Standards and practices of butchery differ between countries, regions and ethnic groups. Variation with respect to the types of animals that are butchered as well as the cuts and parts of

2688-669: Is only weak support for most of this. Farmers did select for reduced bitterness and lower toxicity and for food quality, which likely increased crop palatability to herbivores as to humans. However, a survey of 29 plant domestications found that crops were as well-defended against two major insect pests ( beet armyworm and green peach aphid ) both chemically (e.g. with bitter substances) and morphologically (e.g. with toughness) as their wild ancestors. During domestication, crop species undergo intense artificial selection that alters their genomes, establishing core traits that define them as domesticated, such as increased grain size. Comparison of

2800-668: Is the suite of phenotypic traits that arose during the initial domestication process and which distinguish crops from their wild ancestors . It can also mean a set of differences now observed in domesticated mammals, not necessarily reflecting the initial domestication process. The changes include increased docility and tameness, coat coloration, reductions in tooth size, craniofacial morphology, ear and tail form (e.g., floppy ears), estrus cycles, levels of adrenocorticotropic hormone and neurotransmitters, prolongations in juvenile behavior, and reductions in brain size and of particular brain regions. The domestication of animals and plants

2912-725: Is today Baluchistan in Pakistan. In East Asia 8,000 years ago, pigs were domesticated from wild boar genetically different from those found in the Fertile Crescent. The cat was domesticated in the Fertile Crescent, perhaps 10,000 years ago, from European wildcats , possibly to control rodents that were damaging stored food. The domestication of vertebrate animals is the relationship between non-human vertebrates and humans who have an influence on their care and reproduction. In his 1868 book The Variation of Animals and Plants Under Domestication , Charles Darwin recognized

3024-636: Is widely grown for food. The yeast Saccharomyces cerevisiae have been used for thousands of years to ferment beer and wine , and to leaven bread . Mould fungi including Penicillium are used to mature cheeses and other dairy products, as well as to make drugs such as antibiotics . Selection of animals for visible traits may have undesired consequences for the genetics of domestic animals. A side effect of domestication has been zoonotic diseases. For example, cattle have given humanity various viral poxes , measles , and tuberculosis ; pigs and ducks have contributed influenza ; and horses have brought

3136-607: The Anti-Lebanon Mountains and capturing Pella, Jordan and Damascus . Pompey's incursion further south, into Judea , was occasioned on account of its inhabitants, under the leadership of Hyrcanus II and Aristobulus II , having ravaged Phoenicia and Pompey wanting to bring a stop to it. The initial onslaught was disrupted by the Hasmonean Civil War , in which Pompey backed Hyrcanus II over his brother Aristobulus II. When he compelled

3248-524: The Battle of Italica . Pompey faced Sertorius in the indecisive Battle of Sucro , in which Sertorius defeated Pompey's right flank and nearly captured Pompey himself, but his legate Lucius Afranius defeated the Sertorian right. Sertorius withdrew inland, then turned to fight at Saguntum , where Pompey lost 6,000 men, including his brother-in-law Memmius, reputedly his most effective subordinate. Sertorius himself suffered 3,000 casualties, one of whom

3360-524: The Battle of the Colline Gate . Pompey, who was pursuing the government forces, arrived just after the battle. By the end of 82 BC, Sulla had expelled his opponents from Italy, and engineered his nomination as Dictator by the Senate. Either through admiration of his abilities, or concern at his ambition, Sulla sought to consolidate his alliance with Pompey by persuading him to divorce Antistia, and marry his stepdaughter Aemilia. Plutarch claims she

3472-528: The First Triumvirate , an informal political alliance designed to counter-balance the optimates . Pompey's influence was based on his reputation as a military commander, and popularity with the Roman people. Crassus' wealth allowed him to construct extensive patronage networks, but he lacked the military clout essential for political success in the late Republican era . Once elected, Caesar secured

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3584-718: The Indian Ocean and those in the Pacific Ocean . The coconut experienced a founder effect , where a small number of individuals with low diversity founded the modern population, permanently losing much of the genetic variation of the wild population. Population bottlenecks which reduced variation throughout the genome at some later date after domestication are evident in crops such as pearl millet , cotton , common bean and lima bean . In wheat, domestication involved repeated hybridization and polyploidy . These steps are large and essentially instantaneous changes to

3696-634: The Levant at least 23,000 years ago. Neolithic societies in West Asia first began to cultivate and then domesticate some of these plants around 13,000 to 11,000 years ago. The founder crops of the West Asian Neolithic included cereals ( emmer , einkorn wheat , barley ), pulses ( lentil , pea , chickpea , bitter vetch ), and flax . Other plants were independently domesticated in 13 centers of origin (subdivided into 24 areas) of

3808-683: The Roman Senate . Sulla's return in 83 BC sparked a civil war within the Roman world. In the year prior to Sulla's return Pompey had raised and equipped a full legion from amongst his father's old clients and veterans in Picenum . In the spring of 83 Sulla landed in Brundusium . As he marched north-west towards Campania , Pompey led his own legion south to join him. The government in Rome sent out three separate armies in an attempt to prevent

3920-543: The Social War (91–87 BC) . Strabo died in 87 BC during the short-lived civil war known as the Bellum Octavianum , although sources differ on whether he succumbed to disease, or was murdered by his own soldiers. Prior to his death, Strabo was accused of embezzlement ; as his legal heir, Pompey was held responsible for the alleged crime and put on trial. He was acquitted, supposedly after agreeing to marry

4032-614: The Third Mithridatic War , and in various other military campaigns. Pompey's early success earned him the cognomen Magnus – "the Great" – after his boyhood hero Alexander the Great . His adversaries gave him the nickname adulescentulus carnifex ("teenage butcher") for his ruthlessness. In 60 BC, Pompey joined Crassus and Caesar in the informal political alliance known as the First Triumvirate , cemented by Pompey's marriage with Caesar's daughter, Julia . After

4144-497: The Third Servile War . Pompey returned to Italy just before Crassus defeated the main rebel army in 71 BC, arriving in time to massacre 6,000 fugitives from the battle. His claim to have ended the war by doing so was a long-standing source of resentment for Crassus. Pompey was granted a second triumph for his victory in Hispania, and nominated for the consulship. Since he was both too young and technically ineligible, this required

4256-582: The Vaccaei . Dire straits caused by this stretch of the campaign and Sertorius' guerrilla warfare led Pompey to write a letter to the Senate asking for funds and men, and scolding their lack of support for him and Metellus. Pompey's letter had the effect of galvanizing the Senate into sending him more men and funds. Reinforced by two more legions, in 74 BC he and Metellus began a war of attrition against their enemy. As his chief opponent had lost most of his Roman legionaries and could no longer match him in

4368-411: The carotid or brachial arteries to facilitate blood removal), skinning (removing the hide or pelt) or scalding and dehairing (pork), evisceration (removing the viscera ) and splitting (dividing the carcass in half longitudinally ). After the carcasses are chilled (unless "hot-boned"), primary butchery consists of selecting carcasses, sides, or quarters from which primal cuts can be produced with

4480-619: The coding DNA of chromosome 8 in rice between fragrant and non-fragrant varieties showed that aromatic and fragrant rice, including basmati and jasmine , is derived from an ancestral rice domesticate that suffered a deletion in exon 7 which altered the coding for betaine aldehyde dehydrogenase (BADH2). Comparison of the potato genome with that of other plants located genes for resistance to potato blight caused by Phytophthora infestans . In coconut , genomic analysis of 10 microsatellite loci (of noncoding DNA ) found two episodes of domestication based on differences between individuals in

4592-470: The domestication of wheat . Wild wheat shatters and falls to the ground to reseed itself when ripe, but domesticated wheat stays on the stem for easier harvesting. This change was possible because of a random mutation in the wild populations at the beginning of wheat's cultivation . Wheat with this mutation was harvested more frequently and became the seed for the next crop. Therefore, without realizing it, early farmers selected for this mutation. The result

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4704-417: The domestication syndrome , traits presumed essential in the early stages of domestication, while others represent later improvement traits. Domesticated mammals in particular tend to be smaller and less aggressive than their wild counterparts; other common traits are floppy ears, a smaller brain, and a shorter muzzle. Domestication traits are generally fixed within all domesticates, and were selected during

4816-520: The judge's daughter, Antistia . One of the main issues at stake in 87 BC was the appointment of the consul Lucius Cornelius Sulla as commander of the Roman army in the ongoing First Mithridatic War , an opportunity to amass enormous wealth. During his absence in the East, his political rivals led by Lucius Cornelius Cinna , Gnaeus Papirius Carbo and Gaius Marius the Younger regained control of

4928-437: The rhinoviruses . Many parasites , too, have their origins in domestic animals. Alongside these, the advent of domestication resulted in denser human populations, which provided ripe conditions for pathogens to reproduce, mutate, spread, and eventually find a new host in humans. Scholars have expressed widely differing viewpoints on domestication's effects on society. Anarcho-primitivism critiques domestication as destroying

5040-436: The silkworm and the western honey bee , have been domesticated for over 5,000 years, often for commercial use. The silkworm is raised for the silk threads wound around its pupal cocoon; the western honey bee, for honey , and, from the 20th century, for pollination of crops. Several other invertebrates have been domesticated, both terrestrial and aquatic, including some such as Drosophila melanogaster fruit flies and

5152-555: The Americas began to cultivate peanuts , squash , maize , potatoes , cotton , and cassava . Rice was first domesticated in China some about 9,000 years ago. In Africa, crops such as sorghum were domesticated. Agriculture developed in some 13 centres around the world, domesticating different crops and animals. Three groups of insects, namely ambrosia beetles , leafcutter ants , and fungus-growing termites have independently domesticated species of fungi, on which they feed. In

5264-592: The Americas, Africa, and Asia (the Middle East, South Asia, the Far East, and New Guinea and Wallacea); in some thirteen of these regions people began to cultivate grasses and grains. Rice was first cultivated in East Asia. Sorghum was widely cultivated in sub-Saharan Africa, while peanuts, squash, cotton, maize , potatoes , and cassava were domesticated in the Americas. Continued domestication

5376-498: The Great of Armenia . Lucullus was a skilled general who won numerous victories, but claims he was protracting the war for "power and wealth" led to a Senate investigation, while by 69 BC his troops were weary and mutinous. In 68 BC, Quintus Marcius Rex replaced Lucullus in Cicilia, while Manius Acilius Glabrio received Bithynia. He also assumed leadership of the war against Mithridates, but failed to respond decisively when

5488-524: The Great , was a general and statesman of the Roman Republic . He played a significant role in the transformation of Rome from republic to empire . Early in his career, he was a partisan and protégé of the Roman general and dictator Sulla ; later, he became the political ally, and finally the enemy, of Julius Caesar . A member of the senatorial nobility, Pompey entered into a military career while still young. He rose to prominence serving Sulla as

5600-595: The Late Pleistocene. Domestication reduces genetic diversity of the domesticated population, especially of alleles of genes targeted by selection. One reason is a population bottleneck created by artificially selecting the most desirable individuals to breed from. Most of the domesticated strain is then born from just a few ancestors, creating a situation similar to the founder effect . Domesticated populations such as of dogs, rice, sunflowers, maize, and horses have an increased mutation load , as expected in

5712-769: The Mithridatic War, notably Soli , renamed Pompeiopolis, and Dyme in Greece, with others sent to towns in Libya and Calabria . These communities retained a strong attachment to both Rome and Pompey. In 73 BC, Lucius Licinius Lucullus , formerly one of Sulla's chief lieutenants, was made proconsul of Cilicia , and commander in the Third Mithridatic War. The war began in 74 BC, when the last ruler of Bithynia died and left his kingdom to Rome, sparking an invasion by Mithridates VI of Pontus , and Tigranes

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5824-522: The Senate was divided, Pompey remained as popular as ever with the masses. He was awarded a third triumph for his achievements in Asia Minor, celebrated on his 45th birthday in 61 BC. Pompey claimed the new provinces established in the East had increased annual state income from 200 million to 340 million sesterces , plus an additional payment of 480 million sesterces to the treasury. He refused to provide details of his personal fortune, but given

5936-477: The Senate, Pompey was appointed military commander in Spain with proconsular authority in order to defeat Sertorius. This act was technically illegal as he had yet to hold public office, illustrating Pompey's preference for military glory, and disregard for traditional political constraints. Pompey recruited 30,000 infantry and 1,000 cavalry, evidence of the threat posed by Sertorius. En route to Hispania, he subdued

6048-508: The United States even that role is diminished with the advent of "case-ready" meat, where the product is packaged for retail sale at the packinghouse or specialized central processing plants. A primal cut is a piece of meat initially separated from the carcass during butchering. Different countries and cultures make these cuts in different ways, and primal cuts also differ between type of carcass. The British, American and French primal cuts all differ in some respects. One notable example with pork

6160-519: The amounts declared publicly, this must have been enormous. Some of it was used to build one of the most famous structures of Ancient Rome, the Theatre of Pompey . However, the Senate then refused to ratify the treaties agreed by Pompey as part of his settlement of the East. Opposition was led by the optimates Cato the Younger and Metellus Celer , whose sister Mucia had recently been divorced by Pompey, for reasons still disputed. They also defeated

6272-419: The animal that are sold depends on the types of foods that are prepared by the butcher's customers. Butchery is a traditional line of work. In the industrialized world, slaughterhouses use butchers to slaughter the animals, performing one or a few of the steps repeatedly as specialists on a semi-automated disassembly line. The steps include stunning (rendering the animal incapacitated), exsanguination (severing

6384-458: The beginning of the Holocene 11,700 years ago, a warmer climate and increasing human populations led to small-scale animal and plant domestication and an increased supply of food. The appearance of the domestic dog in the archaeological record, at least 15,000 years ago, was followed by domestication of livestock and of crops such as wheat and barley , the invention of agriculture , and

6496-562: The case of the termites, the relationship is a fully obligate symbiosis on both sides. Domestication (not to be confused with the taming of an individual animal ), is from the Latin domesticus , 'belonging to the house'. The term remained loosely defined until the 21st century, when the American archaeologist Melinda A. Zeder defined it as a long-term relationship in which humans take over control and care of another organism to gain

6608-717: The consul Gaius Papirius Carbo in Cisalpine Gaul . During this campaign Pompey acted as Metellus's cavalry commander. Metellus and Pompey defeated Carbo's lieutenant, the praetor Gaius Carrinas , in a six-hour battle at the river Aesis, only to be blockaded by Carbo himself. When word of Sulla's victory at the Battle of Sacriportus reached them, Carbo retreated to his base at Ariminium , severely harassed by Pompey's cavalry. Some time later Metellus defeated Gaius Marcius Censorinus , another of Carbo's lieutenants, Pompey's cavalry caught Censorinus's fleeing troops outside their base at Sena Gallica , defeating them and plundering

6720-479: The deaths of Julia and Crassus (in 54 and 53 BC), Pompey switched to the political faction known as the optimates —a conservative faction of the Roman Senate . Pompey and Caesar then began contending for leadership of the Roman state in its entirety, eventually leading to Caesar's Civil War . Pompey was defeated at the Battle of Pharsalus in 48 BC, and he sought refuge in Ptolemaic Egypt , where he

6832-556: The deposed Hiempsal to the Numidian throne. Around this time, his troops began referring to him as Magnus , or "the Great", after Alexander the Great , a figure much admired by the Romans. Shortly thereafter, Pompey formally made this part of his name . On returning to Rome, he asked for a triumph to celebrate his victories, an unprecedented demand for someone so young. Pompey refused to disband his army until Sulla agreed, although

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6944-514: The difficulties Pompey faced came from officials who resented his authority. In Gaul, Piso hampered his recruitment efforts, while in Crete , Quintus Metellus refused to comply with his instructions. Pompey spread his forces throughout the Mediterranean to prevent the pirates escaping a Roman fleet by moving elsewhere. Fifteen legates were given specific areas to patrol, while he secured

7056-509: The domestication of animals involved violence against animals and damage to the environment. This, in turn, he argues, corrupted human ethics and paved the way for "conquest, extermination, displacement, repression, coerced and enslaved servitude, gender subordination and sexual exploitation, and hunger." Domesticated ecosystems provide food, reduce predator and natural dangers, and promote commerce, but their creation has resulted in habitat alteration or loss, and multiple extinctions commencing in

7168-515: The domestication syndrome; it is predicted that domestic animals are less resistant to parasites than their wild relatives. Domesticated birds principally mean poultry , raised for meat and eggs: some Galliformes ( chicken , turkey , guineafowl ) and Anseriformes (waterfowl: ducks , geese , and swans ). Also widely domesticated are cagebirds such as songbirds and parrots ; these are kept both for pleasure and for use in research. The domestic pigeon has been used both for food and as

7280-564: The end of 66 BC. According to contemporary sources, Mithridates and a small contingent escaped the battle, outstripped their pursuers, and reached Colchis on the Black Sea . While there, he took control of the Cimmerian Bosporus from its Roman-backed ruler, his son Machares , who later committed suicide. Meanwhile, Pompey invaded Armenia supported by Tigranes the Younger , whose father quickly came to terms; in return for

7392-482: The field, Pompey, along with Metellus, gained the upper hand, conquering more and more Sertorian cities, slowly grinding down Sertorius' revolt. By now, Sertorius was being undermined by internal divisions. Discontent in Sertorius' coalition of Iberian and Roman forces came to a head in 72 or 73 BC when Perperna, leading a conspiracy with other prominent Sertorians, had Sertorius assassinated and assumed control of

7504-600: The first century BC, got the Latin nickname adulescentulus carnifex , translated as "The Teenage Butcher" or "The Butcher Boy", due to brutal treatment of political opponents in the early part of his career. More recently, the Bosnian Serb war criminal Ratko Mladić was nicknamed "the Butcher of the Balkans ". The term can also be used in a semi-humorous or metaphorical way to describe someone whose actions resemble

7616-447: The freshwater cnidarian Hydra for research into genetics and physiology. Few have a long history of domestication. Most are used for food or other products such as shellac and cochineal . The phyla involved are Cnidaria , Platyhelminthes (for biological pest control ), Annelida , Mollusca , Arthropoda (marine crustaceans as well as insects and spiders), and Echinodermata . While many marine mollusks are used for food, only

7728-810: The genome and the epigenome , enabling a rapid evolutionary response to artificial selection. Polyploidy increases the number of chromosomes, bringing new combinations of genes and alleles, which in turn enable further changes such as by chromosomal crossover . The microbiome , the collection of microorganisms inhabiting the surface and internal tissue of plants, is affected by domestication. This includes changes in microbial species composition and diversity. Plant lineage, including speciation , domestication, and breeding , have shaped plant endophytes ( phylosymbiosis ) in similar patterns as plant genes. Several species of fungi have been domesticated for use directly as food, or in fermentation to produce foods and drugs. The cultivated mushroom Agaricus bisporus

7840-627: The government forces made one final effort to march to the relief of Praeneste. They mustered 10,000 legionaries and marched to join forces with the Samnites and the Lucanians , fierce enemies of Sulla, who had campaigned against them in the Social War. Pursued by Pompey they united their forces and made for Praeneste. Unable to break through Sulla's blockade, they marched for undefended Rome, only to be caught just in time and defeated by Sulla at

7952-611: The grain route to Rome. These measures won him control of the western Mediterranean in just 40 days, after which his fleets moved to the east, forcing the pirates back to their bases in Cilicia. Pompey led the decisive assault on their stronghold in Coracaesium , winning the Battle of Korakesion and concluding the war in only three months. Most of his opponents surrendered without fighting, thanks to Pompey's reputation for clemency. They were granted lands in cities devastated during

8064-532: The homogenizing effect of gene flow from wild boars into pigs, and created domestication islands in the genome. The same process may apply to other domesticated animals. The 2023 parasite-mediated domestication hypothesis suggests that endoparasites such as helminths and protozoa could have mediated the domestication of mammals. Domestication involves taming, which has an endocrine component; and parasites can modify endocrine activity and microRNAs . Genes for resistance to parasites might be linked to those for

8176-467: The idea. When Sulla died in 78 BC, Lepidus sought to block his state funeral and roll back some of Sulla's laws, then became proconsul of Cisalpine and Transalpine Gaul in January 77 BC. When the Senate ordered him back to Rome, Lepidus refused to comply unless granted another term as consul, a proposal that was rapidly rejected. Assembling an army, he began marching on Rome; the Senate responded with

8288-558: The independent and wealthy cities of Syria , which were incorporated into a new Roman province. In the process, he acquired large amounts of money and prestige, as well as criticism from his opponents in Rome, who argued doing so exceeded his authority. Meanwhile, an ageing Mithridates had been cornered in Panticapaeum by another of his sons, Pharnaces II of Pontus . An attempt to commit suicide by taking poison allegedly failed due to his habit of taking "precautionary antidotes", and he

8400-426: The initial episode of domestication of that animal or plant, whereas improvement traits are present only in a proportion of domesticates, though they may be fixed in individual breeds or regional populations . Certain animal species, and certain individuals within those species, make better candidates for domestication because of their behavioral characteristics: The beginnings of mammal domestication involved

8512-435: The latter re-occupied much of Pontus in 67 BC, then attacked Cappadocia , a Roman ally. Seeing an opportunity, in 66 BC Pompey used the tribunate to pass the lex Manilia , giving him extensive powers throughout Asia Minor in order to defeat Mithridates, in addition to those granted by the lex Gabinia . The optimates were privately horrified that one man should hold so much influence, but fearful of his popularity allowed

8624-628: The latter to surrender Jerusalem , its defenders took refuge in the Temple , which the Romans first stormed , then looted. Judea became a client kingdom ruled by Hyrcanus, while its northern section was incorporated into the Decapolis , a league of semi-autonomous cities (see map). Both Judea and the League were made subordinate to the new province of Syria. Other organisational changes included creating

8736-543: The latter tried to offset the impact by awarding simultaneous triumphs to Lucius Licinius Murena and Gaius Valerius Flaccus . Sometime during this period, Pompey married Mucia Tertia, a member of the powerful Metellus family. They had three children before their divorce in 61 BC; Pompey the younger , usually known as Gnaeus, a daughter, Pompeia Magna , and a younger son, Sextus . Pompey supported Marcus Aemilius Lepidus as consul for 78 BC; Plutarch claims he did so against Sulla's advice, but most modern historians refute

8848-580: The leaf material to the fungi that they tend. Some of these fungi are not fully domesticated: the fungi farmed by Mycocepurus smithii constantly produce spores that are not useful to the ants, which eat fungal hyphae instead. The process of domestication by Atta ants, on the other hand, is complete; it took 30 million years. Some 330 fungus-growing termite species of the subfamily Macrotermitinae cultivate Termitomyces fungi to eat; domestication occurred exactly once, 25–40 mya . The fungi, described by Roger Heim in 1942, grow on 'combs' formed from

8960-414: The local Roman administration, showing a lack of animosity towards his former opponents, which extended his patronage throughout Hispania and into southern Gaul. Pompey and his army remained in Hispania for a few years conquering the Sertorian remnants, and then marched back to Rome. During Pompey's absence, Marcus Licinius Crassus was charged with suppressing the slave rebellion led by Spartacus known as

9072-549: The measure to pass. Incensed at being replaced, Lucullus called Pompey a "vulture" who profited from the work of others, a reference both to his new command and claim to have finished the war against Spartacus. Pompey agreed an alliance with Phraates III , king of Parthia , whom he persuaded to invade Armenia. When Mithridates offered a truce, Lucullus argued the war was over, but Pompey demanded concessions which could not be accepted. Outnumbered, Mithridates withdrew into Armenia, followed by Pompey, who defeated him at Lycus near

9184-483: The minimum of wastage; separating the primal cuts from the carcass; trimming primal cuts and preparing them for secondary butchery or sale; and storing cut meats. Secondary butchery involves boning, trimming and value-adding of primal cuts in preparation for sale. Historically, primary and secondary butchery were performed in the same establishment, but the advent of methods of preservation ( vacuum packing ) and low cost transportation has largely separated them. In parts of

9296-477: The passage of a new agrarian bill, helped by Pompey's veterans, who filled the streets of Rome and allegedly intimidated the Senate. When Bibulus opposed the measure, he was attacked in the forum , and spent the rest of his consulship under virtual house arrest. Caesar then ensured ratification of Pompey's settlements in the east, while the Lex Vatinia made him governor of Gallia Cisalpina and Illyricum . He

9408-597: The past fifty years, partly because of the decline of traditional naval powers like Rhodes , while previous attempts to subdue them had been unsuccessful. However, Romans routinely referred to their opponents as "pirates" or "brigands", and some historians argue it is more accurate to see them as a conventional enemy, rather than disorganised outlaws. Principally based in Cilicia , in 68 BC they raided as far as Ostia , Rome's port, and kidnapped two senators, to general outrage. Prompted by Pompey, Aulus Gabinius , tribune of

9520-461: The plebs in 67 BC, proposed the Lex Gabinia , giving him a mandate for their suppression. It granted him proconsular authority for three years in any province within 50 miles of the Mediterranean, along with the power to appoint legates and significant financial resources. Concerned by one man holding such wide-ranging powers, the Senate opposed the law but it was passed by the people. Most of

9632-400: The province of Bithynia and Pontus , with the rest of Mithridates' territories distributed among Roman allies. Elsewhere, Ariobarzanes I of Cappadocia was restored to his throne, while Lesser Armenia was taken from Tigranes and incorporated into Galatia , with Pompey's client Deiotarus becoming ruler of the new kingdom. Finally, Cilicia received the coastal region of Pamphylia , previously

9744-403: The rebel army. Pompey engaged Perperna in battle and defeated him swiftly at the Battle near Osca . Perperna was captured and attempted to persuade Pompey to spare him by giving over Sertorius' correspondence, allegedly containing proof of communications between the rebel leader and leading men in Rome. Pompey burned the letters unread and executed Perperna, and then spent some time restructuring

9856-514: The relationship between them intensified, and humans' role in their survival and reproduction gradually led to formalized animal husbandry . Although the directed pathway for draft and riding animals proceeded from capture to taming, the other two pathways are not as goal-oriented, and archaeological records suggest that they took place over much longer time frames. Unlike other domestic species selected primarily for production-related traits, dogs were initially selected for their behaviors. The dog

9968-627: The remnants of his army retreated to Sardinia , where he died. The Sertorian War began in 80 BC when Quintus Sertorius , a prominent proscribed Marian general, initiated a rebellion in Hispania , where he was joined by other Roman exiles like Perperna. Supported by local Iberian tribes, he took control of Hispania Ulterior and repeatedly defeated Quintus Caecilius Metellus Pius through skillful use of guerrilla warfare . Sertorius defeated other Roman generals sent to oust him and soon conquered Hispania Citerior as well. Backed by his allies in

10080-400: The restoration of Armenian territories taken by Lucullus, he paid a substantial cash indemnity and allowed Roman troops to be based on his territory. In 65 BC, Pompey set out to take Colchis, but to do so had first to subdue various local tribes and allies of Mithridrates. After winning a series of battles, he reached Phasis and linked up with Servilius, admiral of his Euxine fleet, before

10192-459: The small number of traits that made domestic species different from their wild ancestors. He was also the first to recognize the difference between conscious selective breeding in which humans directly select for desirable traits and unconscious selection, in which traits evolve as a by-product of natural selection or from selection on other traits. There is a difference between domestic and wild populations; some of these differences constitute

10304-451: The supposed primitive state of harmony with nature in hunter-gatherer societies, and replacing it, possibly violently or by enslavement, with a social hierarchy as property and power emerged. The dialectal naturalist Murray Bookchin has argued that domestication of animals, in turn, meant the domestication of humanity, both parties being unavoidably altered by their relationship with each other. The sociologist David Nibert asserts that

10416-472: The termites' excreta, dominated by tough woody fragments. The termites and the fungi are both obligate symbionts in the relationship. Pompey People Events Places Gnaeus Pompeius Magnus ( Latin: [ˈŋnae̯ʊs pɔmˈpɛjjʊs ˈmaŋnʊs] ; 29 September 106 BC – 28 September 48 BC), known in English as Pompey ( / ˈ p ɒ m p i / POM -pee ) or Pompey

10528-668: The town. While Metellus remained in the north-west, Pompey seems to have transferred to Sulla's command in the south. Pompey advanced south-west along the Via Flaminia towards Spoletium , where he joined Marcus Licinius Crassus, together they defeated Carrinas once again. Pompey laid siege to Carrinas in Spoletium but the latter managed to escape. Pompey resumed his march to join Sulla's command. Not long afterwards Pompey successfully ambushed another large force under Censorinus, which

10640-637: The transition of humans from foraging to farming in different places and times across the planet. For instance, small-scale trial cultivation of cereals began some 28,000 years ago at the Ohalo II site in Israel. In the Fertile Crescent 11,000–10,000 years ago, zooarchaeology indicates that goats, pigs, sheep, and taurine cattle were the first livestock to be domesticated. Two thousand years later, humped zebu cattle were domesticated in what

10752-588: The union between Pompey's and Sulla's army. Pompey attacked one of these armies and routed it. The three enemy commanders, unable to agree on a course of action, withdrew. Soon after Pompey arrived at Sulla's camp. He was greeted by Sulla with the official title of Imperator (General). At some point in 83 BC, it is not clear when but definitely before the onset of winter, Sulla sent Pompey back to Picenum to raise more troops. When fighting broke out once more in 82 Sulla advanced towards Rome, while Metellus (one of his lieutenants), supported by Pompey, campaigned against

10864-440: The urban electorate. With additional support from Crassus, Caesar became one of the two consuls for 59 BC, the other being the optimate Marcus Calpurnius Bibulus . This meant Caesar could help pass legislation sponsored by Pompey and Crassus, while it was in his interest to keep them aligned, an important factor given the rivalry between his two patrons. Despite appearing to be the most junior, Caesar thus became central to

10976-562: The various skills and methods of a butcher (chopping, cutting, slicing, stabbing etc.) Spanish footballer Andoni Goikoetxea was popularly ascribed the epithet "The Butcher of Bilbao" in recognition of his perceived aggressive style of play and frequent, sometimes injurious, challenges on opposing players. Domestication Domestication is a multi-generational mutualistic relationship in which an animal species, such as humans or leafcutter ants , takes over control and care of another species, such as sheep or fungi, to obtain from them

11088-408: The wild. The first animal to be domesticated by humans was the dog , as a commensal , at least 15,000 years ago. Other animals, including goats , sheep , and cows , were domesticated around 11,000 years ago. Among birds, the chicken was first domesticated in East Asia, seemingly for cockfighting, some 7,000 years ago. The horse came under domestication around 5,500 years ago in central Asia as

11200-692: The world, it is common for butchers to perform many or all of the butcher's duties. Where refrigeration is less common, these skills are required to sell the meat of slaughtered animals. Butchers sell their goods in specialized stores, commonly termed a butcher shop ( American English ), butchery ( South African English ) or butchers ( British English ). Butchers at a butcher shop may perform primary butchery, but will typically perform secondary butchery to prepare fresh cuts of meat for sale. These shops may also sell related products, such as Charcuterie , hot food (using their own meat products), food preparation supplies, baked goods and grocery items. Butcher shops can have

11312-512: Was Hirtuleius. Although Metellus defeated Perperna in a separate battle, Sertorius was able to withdraw to Clunia late in the year, where he repaired the walls to lure his opponents into a siege, while forming garrisons from other towns into a new field army. Once this was ready, he escaped from Clunia and used it to disrupt Roman logistics on land and by sea. Lack of supplies forced Metellus to quarter his troops in Gaul , while Pompey wintered among

11424-472: Was a serious blow to Pompey's prestige, who spent the rest of the year re-organising his army. Metellus' failure to dislodge Sertorius and Pompey's defeat meant the senatorial generals made no progress in the year. In 75 BC, Sertorius led the campaign against Metellus, while Pompey defeated his subordinates Perperna and Gaius Herennius outside Valencia . When Sertorius took over operations against Pompey, Metellus defeated his deputy Lucius Hirtuleius at

11536-493: Was already pregnant by her former husband, and died in childbirth soon after. The surviving Marians escaped to Sicily, where their ally Marcus Perperna was propraetor . They were supported by a fleet under Carbo, while Gnaeus Domitius Ahenobarbus occupied the Roman province of Africa . Perperna abandoned Sicily after Pompey landed on the island with a large force, while Carbo was captured and later executed. Pompey claimed this

11648-535: Was also assigned Gallia Transalpina after its governor died in office, before leaving Rome to launch the Gallic Wars in 58 BC. His alliance with Pompey was strengthened when the latter married Caesar's daughter Julia. Senatorial opposition to the triumvirate was led by Cicero , a long-standing Pompeian ally. Despite this, the latter supported the populist politician Publius Clodius Pulcher in an attack on Cicero for executing Roman citizens without trial during

11760-554: Was assassinated by the courtiers of Ptolemy XIII . Pompey was born in Picenum on 29 September 106 BC, eldest son of a provincial noble called Gnaeus Pompeius Strabo . Although the dominant family in Picenum, Strabo was the first of his branch to achieve senatorial status in Rome ; he completed the traditional cursus honorum , becoming consul in 89 BC, and acquired a reputation for greed, political duplicity, and military ruthlessness. Pompey began his career serving with his father in

11872-500: Was domesticated long before other animals, becoming established across Eurasia before the end of the Late Pleistocene era, well before agriculture . The archaeological and genetic data suggest that long-term bidirectional gene flow between wild and domestic stocks – such as in donkeys , horses , New and Old World camelids, goats, sheep, and pigs – was common. Human selection for domestic traits likely counteracted

11984-532: Was gradual and geographically diffuse – happening in many small steps and spread over a wide area – on the evidence of both archaeology and genetics. It was a process of intermittent trial and error and often resulted in diverging traits and characteristics. Whereas domestication of animals impacted most on the genes that controlled behavior, that of plants impacted most on the genes that controlled morphology (seed size, plant architecture, dispersal mechanisms) and physiology (timing of germination or ripening), as in

12096-410: Was justified by Carbo's alleged crimes against Roman citizens, but his opponents nicknamed him adulescentulus carnifex , or "young butcher", as a result. Pompey now sailed for Africa, leaving Sicily in the hands of his brother-in-law, Gaius Memmius . After defeating and killing Ahenobarbus at the Battle of Utica , Pompey subdued Numidia and executed its king Hiarbas , a Marian ally. He restored

12208-607: Was killed by the rebels. Pharnaces sent his embalmed body to Pompey, in return for which he was granted the Bosporan Kingdom and made an ally of Rome. The final collapse of the Seleucid Empire allowed Pompey to annex Syria in 64 BC, but its dissolution destabilised the region, while many of its cities had used the power vacuum to achieve independence. In early 63 BC, Pompey left Antioch and marched south, occupying coastal cities like Apamea , before crossing

12320-516: Was the ability to veto Senatorial bills, an act often seen as a turning point in the politics of the late Republic. Although popular with the people, the measure must have been opposed by the optimates , and thus passing it required support from both consuls, although most extant sources barely mention Crassus. Pirates operated throughout the Mediterranean, while their fleets often formed temporary alliances with enemies of Rome, including Sertorius and Mithridates. Their power and range had increased over

12432-557: Was triggered by the climatic and environmental changes that occurred after the peak of the Last Glacial Maximum and which continue to this present day. These changes made obtaining food by hunting and gathering difficult. The first animal to be domesticated was the dog at least 15,000 years ago. The Younger Dryas 12,900 years ago was a period of intense cold and aridity that put pressure on humans to intensify their foraging strategies but did not favour agriculture. By

12544-530: Was trying to get through to Praeneste where Carbo's consular colleague, Marius the Younger (who was the figurehead of the struggle against Sulla), was blockaded. It was the failure of these attempts to get through the Sullan blockade in Umbria and Etruria , added to Metellus's success in winning control of the north, which broke the back of the government's resistance. At the end of the campaigning season of 82,

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