Misplaced Pages

#383616

148-640: The name (in Greek) Αλιάκμονας is composite and derives from άλας (salt, sea) and άκμων (anvil). In Greek mythology Haliakmon was one of the Potamoi , who were sons of Oceanus and Tethys , according to the allegorical obsessive-anthropomorphic concept familiar to the Ancient Greeks , on geological upheaval after Deucalion's deluge . An ancient tradition says that sheep that drank water from Haliakmon would turn their colour to white. This tradition

296-579: A pederastic light . Alexandrian poets at first, then more generally literary mythographers in the early Roman Empire, often re-adapted stories of Greek mythological characters in this fashion. The achievement of epic poetry was to create story-cycles and, as a result, to develop a new sense of mythological chronology. Thus, Greek mythology unfolds as a phase in the development of the world and of humans. While self-contradictions in these stories make an absolute timeline impossible, an approximate chronology may be discerned. The resulting mythological "history of

444-677: A Christian moralizing perspective. The discovery of the Mycenaean civilization by the German amateur archaeologist Heinrich Schliemann in the nineteenth century, and the discovery of the Minoan civilization in Crete by the British archaeologist Arthur Evans in the twentieth century, helped to explain many existing questions about Homer's epics and provided archaeological evidence for many of

592-585: A collection of epic poems , starts with the events leading up to the war: Eris and the golden apple of Kallisti , the Judgement of Paris , the abduction of Helen , the sacrifice of Iphigenia at Aulis . To recover Helen, the Greeks launched a great expedition under the overall command of Menelaus 's brother, Agamemnon, king of Argos, or Mycenae , but the Trojans refused to return Helen. The Iliad , which

740-410: A collection of polyphagous heterotrophic consumers that network and cycle the flow of energy and nutrients from a productive base of self-feeding autotrophs . The base or basal species in a food web are those species without prey and can include autotrophs or saprophytic detritivores (i.e., the community of decomposers in soil , biofilms , and periphyton ). Feeding connections in

888-492: A combination of their name and epithets , that identify them by these distinctions from other manifestations of themselves (e.g., Apollo Musagetes is " Apollo , [as] leader of the Muses "). Alternatively, the epithet may identify a particular and localized aspect of the god, sometimes thought to be already ancient during the classical epoch of Greece. Most gods were associated with specific aspects of life. For example, Aphrodite

1036-525: A convenient framework into which to fit their own courtly and chivalric ideals. Twelfth-century authors, such as Benoît de Sainte-Maure ( Roman de Troie [Romance of Troy, 1154–60]) and Joseph of Exeter ( De Bello Troiano [On the Trojan War, 1183]) describe the war while rewriting the standard version they found in Dictys and Dares . They thus follow Horace 's advice and Virgil's example: they rewrite

1184-489: A flat disk afloat on the river of Oceanus and overlooked by a hemispherical sky with sun, moon, and stars. The Sun ( Helios ) traversed the heavens as a charioteer and sailed around the Earth in a golden bowl at night. Sun, earth, heaven, rivers, and winds could be addressed in prayers and called to witness oaths. Natural fissures were popularly regarded as entrances to the subterranean house of Hades and his predecessors, home of

1332-543: A food chain was described by a medieval Afro-Arab scholar named Al-Jahiz : "All animals, in short, cannot exist without food, neither can the hunting animal escape being hunted in his turn." The earliest graphical depiction of a food web was by Lorenzo Camerano in 1880, followed independently by those of Pierce and colleagues in 1912 and Victor Shelford in 1913. Two food webs about herring were produced by Victor Summerhayes and Charles Elton and Alister Hardy in 1923 and 1924. Charles Elton subsequently pioneered

1480-532: A food web has a historical foothold in the writings of Charles Darwin and his terminology, including an "entangled bank", "web of life", "web of complex relations", and in reference to the decomposition actions of earthworms he talked about "the continued movement of the particles of earth". Even earlier, in 1768 John Bruckner described nature as "one continued web of life". Interest in food webs increased after Robert Paine's experimental and descriptive study of intertidal shores suggesting that food web complexity

1628-418: A food web. In a simple predator-prey example, a deer is one step removed from the plants it eats (chain length = 1) and a wolf that eats the deer is two steps removed from the plants (chain length = 2). The relative amount or strength of influence that these parameters have on the food web address questions about: In a pyramid of numbers, the number of consumers at each level decreases significantly, so that

SECTION 10

#1732765666384

1776-469: A food web. Ecologists use these simplifications in quantitative (or mathematical representation) models of trophic or consumer-resource systems dynamics. Using these models they can measure and test for generalized patterns in the structure of real food web networks. Ecologists have identified non-random properties in the topological structure of food webs. Published examples that are used in meta analysis are of variable quality with omissions. However,

1924-460: A food web. Sometimes in food web terminology, complexity is defined as product of the number of species and connectance., though there have been criticisms of this definition and other proposed methods for measuring network complexity. Connectance is "the fraction of all possible links that are realized in a network". These concepts were derived and stimulated through the suggestion that complexity leads to stability in food webs, such as increasing

2072-487: A food-web illustrate direct trophic relations among species, but there are also indirect effects that can alter the abundance, distribution, or biomass in the trophic levels. For example, predators eating herbivores indirectly influence the control and regulation of primary production in plants. Although the predators do not eat the plants directly, they regulate the population of herbivores that are directly linked to plant trophism. The net effect of direct and indirect relations

2220-558: A god "greater than he", Zeus swallowed her. She was already pregnant with Athena , however, and she burst forth from his head—fully-grown and dressed for war. The earliest Greek thought about poetry considered the theogonies to be the prototypical poetic genre—the prototypical mythos —and imputed almost magical powers to it. Orpheus , the archetypal poet, also was the archetypal singer of theogonies, which he uses to calm seas and storms in Apollonius' Argonautica , and to move

2368-513: A limited number of gods, who were the focus of large pan-Hellenic cults. It was, however, common for individual regions and villages to devote their own cults to minor gods. Many cities also honored the more well-known gods with unusual local rites and associated strange myths with them that were unknown elsewhere. During the heroic age, the cult of heroes (or demigods) supplemented that of the gods. "The origins of humanity [were] ascribed to various figures, including Zeus and Prometheus ." Bridging

2516-539: A living system (e.g., ecosystem) sways from equilibrium, the greater its complexity. Complexity has multiple meanings in the life sciences and in the public sphere that confuse its application as a precise term for analytical purposes in science. Complexity in the life sciences (or biocomplexity ) is defined by the "properties emerging from the interplay of behavioral, biological, physical, and social interactions that affect, sustain, or are modified by living organisms, including humans". Several concepts have emerged from

2664-409: A long history in ecology. Like maps of unfamiliar ground, food webs appear bewilderingly complex. They were often published to make just that point. Yet recent studies have shown that food webs from a wide range of terrestrial, freshwater, and marine communities share a remarkable list of patterns. Links in food webs map the feeding connections (who eats whom) in an ecological community . Food cycle

2812-409: A measure mass or energy per m per unit time. Different consumers are going to have different metabolic assimilation efficiencies in their diets. Each trophic level transforms energy into biomass. Energy flow diagrams illustrate the rates and efficiency of transfer from one trophic level into another and up through the hierarchy. It is the case that the biomass of each trophic level decreases from

2960-405: A musical contest with Apollo . Ian Morris considers Prometheus' adventures as "a place between the history of the gods and that of man." An anonymous papyrus fragment, dated to the third century, vividly portrays Dionysus ' punishment of the king of Thrace , Lycurgus , whose recognition of the new god came too late, resulting in horrific penalties that extended into the afterlife. The story of

3108-458: A number of local legends became attached. The story of Medea , in particular, caught the imagination of the tragic poets. In between the Argo and the Trojan War, there was a generation known chiefly for its horrific crimes. This includes the doings of Atreus and Thyestes at Argos. Behind the myth of the house of Atreus (one of the two principal heroic dynasties with the house of Labdacus ) lies

SECTION 20

#1732765666384

3256-441: A poem of Troy instead of telling something completely new. Food web A food web is the natural interconnection of food chains and a graphical representation of what-eats-what in an ecological community . Position in the food web, or trophic level , is used in ecology to broadly classify organisms as autotrophs or heterotrophs . This is a non-binary classification; some organisms (such as carnivorous plants ) occupy

3404-412: A process called biomineralization . Bacteria that live in detrital sediments create and cycle nutrients and biominerals. Food web models and nutrient cycles have traditionally been treated separately, but there is a strong functional connection between the two in terms of stability, flux, sources, sinks, and recycling of mineral nutrients. Food webs are necessarily aggregated and only illustrate

3552-556: A result, silting has been greatly reduced and during the summer, when there is not much water, the sea enters and largely overwhelms the river bed. Flathead mullets and European seabasses are found in Haliakmon's Delta. In the shallow marine areas formed there, the fry of many Aegean Sea 's fish finds a place to reproduce, while about 90% of Greece's mussels are produced in the Haliakmon-Axios (Vardar) Delta. Aliki Kitrous

3700-400: A single top consumer , (e.g., a polar bear or a human ), will be supported by a much larger number of separate producers. There is usually a maximum of four or five links in a food chain, although food chains in aquatic ecosystems are more often longer than those on land. Eventually, all the energy in a food chain is dispersed as heat. Ecological pyramids place the primary producers at

3848-527: A single species can directly and indirectly influence many others. Microcosm studies are used to simplify food web research into semi-isolated units such as small springs, decaying logs, and laboratory experiments using organisms that reproduce quickly, such as daphnia feeding on algae grown under controlled environments in jars of water. While the complexity of real food webs connections are difficult to decipher, ecologists have found mathematical models on networks an invaluable tool for gaining insight into

3996-507: A spirit to every aspect of nature. Eventually, these vague spirits assumed human forms and entered the local mythology as gods. When tribes from the north of the Balkan Peninsula invaded, they brought with them a new pantheon of gods, based on conquest, force, prowess in battle, and violent heroism. Other older gods of the agricultural world fused with those of the more powerful invaders or else faded into insignificance. After

4144-514: A tangled web of omnivores." A central question in the trophic dynamic literature is the nature of control and regulation over resources and production. Ecologists use simplified one trophic position food chain models (producer, carnivore, decomposer). Using these models, ecologists have tested various types of ecological control mechanisms. For example, herbivores generally have an abundance of vegetative resources, which meant that their populations were largely controlled or regulated by predators. This

4292-493: A tiny portion of the complexity of real ecosystems. For example, the number of species on the planet are likely in the general order of 10 , over 95% of these species consist of microbes and invertebrates , and relatively few have been named or classified by taxonomists . It is explicitly understood that natural systems are 'sloppy' and that food web trophic positions simplify the complexity of real systems that sometimes overemphasize many rare interactions. Most studies focus on

4440-413: A top carnivore, without specifying which end." Nonetheless, real differences in structure and function have been identified when comparing different kinds of ecological food webs, such as terrestrial vs. aquatic food webs. Food webs serve as a framework to help ecologists organize the complex network of interactions among species observed in nature and around the world. One of the earliest descriptions of

4588-434: A very general sense, energy flow (E) can be defined as the sum of metabolic production (P) and respiration (R), such that E=P+R. Biomass represents stored energy. However, concentration and quality of nutrients and energy is variable. Many plant fibers, for example, are indigestible to many herbivores leaving grazer community food webs more nutrient limited than detrital food webs where bacteria are able to access and release

Haliacmon - Misplaced Pages Continue

4736-457: Is an area of 254 hectares, where 15–20.000 tons of salt are produced per year. Many kinds of birds have been observed in Haliakmon's region. They use it for accommodation and overwintering, like the ducks , or to rest on their long migratory journey. There have been observed 215 kinds of birds and about ⅓ of them nest in the region. Over 10% of the birds are endangered. They include Dalmatian pelican and curlew , that are considered to be among

4884-509: Is an obsolete term that is synonymous with food web. Ecologists can broadly group all life forms into one of two trophic layers, the autotrophs and the heterotrophs . Autotrophs produce more biomass energy, either chemically without the sun's energy or by capturing the sun's energy in photosynthesis , than they use during metabolic respiration . Heterotrophs consume rather than produce biomass energy as they metabolize, grow, and add to levels of secondary production . A food web depicts

5032-437: Is attributed to different sizes of producers. Aquatic communities are often dominated by producers that are smaller than the consumers that have high growth rates. Aquatic producers, such as planktonic algae or aquatic plants, lack the large accumulation of secondary growth as exists in the woody trees of terrestrial ecosystems. However, they are able to reproduce quickly enough to support a larger biomass of grazers. This inverts

5180-422: Is called trophic cascades. Trophic cascades are separated into species-level cascades, where only a subset of the food-web dynamic is impacted by a change in population numbers, and community-level cascades, where a change in population numbers has a dramatic effect on the entire food-web, such as the distribution of plant biomass. The field of chemical ecology has elucidated multitrophic interactions that entail

5328-691: Is confirmed by the following record of the Roman author Pliny the Elder (23-79 AD) : Similarly in Macedonia, those who want their sheep to be white go to Haliakmon, while those (who want them to be) black (go) to Axios ( Vardar ) . Ottoman Turks called the river Ince-Karasu (narrow-black water), a name still used in Turkey . The Eastern South Slavic name is Бистрица, Bistritsa (Бистрица, from Proto-Slavic *-bystr, meaning "clear, lucid, limpid") which

5476-414: Is known as the top-down hypothesis or 'green-world' hypothesis . Alternatively to the top-down hypothesis, not all plant material is edible and the nutritional quality or antiherbivore defenses of plants (structural and chemical) suggests a bottom-up form of regulation or control. Recent studies have concluded that both "top-down" and "bottom-up" forces can influence community structure and the strength of

5624-442: Is plants, then herbivores (level 2), and then carnivores (level 3). The trophic level equals one more than the chain length, which is the number of links connecting to the base. The base of the food chain (primary producers or detritivores ) is set at zero. Ecologists identify feeding relations and organize species into trophic species through extensive gut content analysis of different species. The technique has been improved through

5772-461: Is portrayed as a sacrificer, mentioned as a founder of altars, and imagined as a voracious eater himself; it is in this role that he appears in comedy. While his tragic end provided much material for tragedy— Heracles is regarded by Thalia Papadopoulou as "a play of great significance in examination of other Euripidean dramas." In art and literature, Heracles was represented as an enormously strong man of moderate height; his characteristic weapon

5920-619: Is set in the tenth year of the war, tells of the quarrel between Agamemnon and Achilles, who was the finest Greek warrior, and the consequent deaths in battle of Achilles' beloved comrade Patroclus and Priam 's eldest son, Hector . After Hector's death, the Trojans were joined by two exotic allies, Penthesilea , queen of the Amazons , and Memnon , king of the Ethiopians and son of the dawn-goddess, Eos . Achilles killed both of these, but Paris then managed to kill Achilles with an arrow in

6068-523: Is still in use in Bulgaria and North Macedonia. Before the construction of its diversion dam near the village of Aghia Varvara in the mid-1950s, Haliakmon had no permanent river bed in its lowland course. It often flooded and formed extensive marshes . Its devastating fury in December 1935 remains fresh in memory of the elder inhabitants of the region ( Veria and Alexandreia ). The Haliacmon rises in

Haliacmon - Misplaced Pages Continue

6216-528: Is that "the Greek gods are persons, not abstractions, ideas or concepts." Regardless of their underlying forms, the Ancient Greek gods have many fantastic abilities; most significantly, the gods are not affected by disease and can be wounded only under highly unusual circumstances. The Greeks considered immortality as the distinctive characteristic of their gods; this immortality, as well as unfading youth,

6364-424: Is the fraction of all possible links that are realized (L/S ) and represents a standard measure of food web complexity..." The distance (d) between every species pair in a web is averaged to compute the mean distance between all nodes in a web (D) and multiplied by the total number of links (L) to obtain link-density (LD), which is influenced by scale-dependent variables such as species richness . These formulas are

6512-510: Is the probable ancient course of the Haliacmon. It was the domain of the eponymous river god Haliacmon . [REDACTED] Media related to Haliacmon at Wikimedia Commons Greek mythology Greek mythology is the body of myths originally told by the ancient Greeks , and a genre of ancient Greek folklore , today absorbed alongside Roman mythology into the broader designation of classical mythology . These stories concern

6660-589: The Chimera and Medusa . Bellerophon's adventures are commonplace types, similar to the adventures of Heracles and Theseus. Sending a hero to his presumed death is also a recurrent theme of this early heroic tradition, used in the cases of Perseus and Bellerophon. The only surviving Hellenistic epic, the Argonautica of Apollonius of Rhodes (epic poet, scholar, and director of the Library of Alexandria ) tells

6808-476: The Derveni Papyrus now proves that at least in the fifth-century BC a theogonic-cosmogonic poem of Orpheus was in existence. The first philosophical cosmologists reacted against, or sometimes built upon, popular mythical conceptions that had existed in the Greek world for some time. Some of these popular conceptions can be gleaned from the poetry of Homer and Hesiod. In Homer, the Earth was viewed as

6956-520: The Geometric period from c.  900 BC to c.  800 BC onward. In fact, literary and archaeological sources integrate, sometimes mutually supportive and sometimes in conflict; however, in many cases, the existence of this corpus of data is an indication that many elements of Greek mythology have strong factual and historical roots. Mythical narration plays an important role in nearly every genre of Greek literature. Nevertheless,

7104-551: The Gramos mountains in northern Greece, near the border with Albania . In its upper course it flows generally towards the east, and turns southeast near Kastoria . It describes a wide curve around the Vourinos mountains, and turns northeast near the village Paliouria . It feeds the large artificial lake Polyfyto , that was created after the construction of the namesake hydroelectric dam and consists almost its entire course through

7252-539: The Hellenistic and Roman ages was primarily composed as a literary rather than cultic exercise. Nevertheless, it contains many important details that would otherwise be lost. This category includes the works of: Prose writers from the same periods who make reference to myths include Apuleius , Petronius , Lollianus , and Heliodorus . Two other important non-poetical sources are the Fabulae and Astronomica of

7400-598: The Parthenon depicting the sack of Troy); this artistic preference for themes deriving from the Trojan Cycle indicates its importance to the Ancient Greek civilization. The same mythological cycle also inspired a series of posterior European literary writings. For instance, Trojan Medieval European writers, unacquainted with Homer at first hand, found in the Troy legend a rich source of heroic and romantic storytelling and

7548-692: The Peloponnesian kingdoms of Mycenae , Sparta and Argos , claiming, according to legend, a right to rule them through their ancestor. Their rise to dominance is frequently called the " Dorian invasion ". The Lydian and later the Macedonian kings, as rulers of the same rank, also became Heracleidae. Other members of this earliest generation of heroes such as Perseus, Deucalion , Theseus and Bellerophon , have many traits in common with Heracles. Like him, their exploits are solitary, fantastic and border on fairy tale , as they slay monsters such as

SECTION 50

#1732765666384

7696-589: The Roman culture because of the story of Aeneas , a Trojan hero whose journey from Troy led to the founding of the city that would one day become Rome, as recounted in Virgil's Aeneid (Book II of Virgil's Aeneid contains the best-known account of the sack of Troy). Finally there are two pseudo-chronicles written in Latin that passed under the names of Dictys Cretensis and Dares Phrygius . The Trojan War cycle ,

7844-548: The ancient Greek religion 's view of the origin and nature of the world ; the lives and activities of deities , heroes , and mythological creatures ; and the origins and significance of the ancient Greeks' cult and ritual practices. Modern scholars study the myths to shed light on the religious and political institutions of ancient Greece, and to better understand the nature of mythmaking itself. The Greek myths were initially propagated in an oral-poetic tradition most likely by Minoan and Mycenaean singers starting in

7992-693: The 18th century BC; eventually the myths of the heroes of the Trojan War and its aftermath became part of the oral tradition of Homer 's epic poems , the Iliad and the Odyssey . Two poems by Homer's near contemporary Hesiod , the Theogony and the Works and Days , contain accounts of the genesis of the world, the succession of divine rulers, the succession of human ages, the origin of human woes, and

8140-473: The Ancient Greek pantheon, poets composed the Homeric Hymns (a group of thirty-three songs). Gregory Nagy (1992) regards "the larger Homeric Hymns as simple preludes (compared with Theogony ), each of which invokes one god." The gods of Greek mythology are described as having essentially corporeal but ideal bodies. According to Walter Burkert , the defining characteristic of Greek anthropomorphism

8288-610: The Greek leaders (including the wanderings of Odysseus and Aeneas (the Aeneid ), and the murder of Agamemnon) were told in two epics, the Returns (the lost Nostoi ) and Homer's Odyssey . The Trojan cycle also includes the adventures of the children of the Trojan generation (e.g., Orestes and Telemachus ). The Trojan War provided a variety of themes and became a main source of inspiration for Ancient Greek artists (e.g. metopes on

8436-472: The Greek world and noted the stories they heard, supplied numerous local myths and legends, often giving little-known alternative versions. Herodotus in particular, searched the various traditions he encountered and found the historical or mythological roots in the confrontation between Greece and the East. Herodotus attempted to reconcile origins and the blending of differing cultural concepts. The poetry of

8584-801: The Kozani prefecture. Over the bridge runs the Lake Polyfytos Bridge , part of the Athens-Kozani national road . Southeast of Veria , the Haliacmon enters the central Macedonian plains, an area of great importance to agriculture. It flows into the Thermaic Gulf west of the delta of the Axios ( Vardar ), northeast of the coastal town Methoni . Haliakmon's tributaries include Gramos, Ladopotamos, Pramoritsa, Grevenitikos, Venetikos, Tripotamos and Krasopoulis. The Haliacmon flows along

8732-553: The Olympians, the Greeks worshipped various gods of the countryside, the satyr-god Pan , Nymphs (spirits of rivers), Naiads (who dwelled in springs), Dryads (who were spirits of the trees), Nereids (who inhabited the sea), river gods, Satyrs , and others. In addition, there were the dark powers of the underworld, such as the Erinyes (or Furies), said to pursue those guilty of crimes against blood-relatives. In order to honor

8880-660: The Roman writer styled as Pseudo- Hyginus , the Imagines of Philostratus the Elder and Philostratus the Younger , and the Descriptions of Callistratus . Finally, several Byzantine Greek writers provide important details of myth, much derived from earlier now lost Greek works. These preservers of myth include Arnobius , Hesychius , the author of the Suda , John Tzetzes , and Eustathius . They often treat mythology from

9028-519: The Trojan cycle, as well as the adventures of Heracles. These visual representations of myths are important for two reasons. Firstly, many Greek myths are attested on vases earlier than in literary sources: of the twelve labors of Heracles, for example, only the Cerberus adventure occurs in a contemporary literary text. Secondly, visual sources sometimes represent myths or mythical scenes that are not attested in any extant literary source. In some cases,

SECTION 60

#1732765666384

9176-456: The age when gods lived alone and the age when divine interference in human affairs was limited was a transitional age in which gods and mortals moved together. These were the early days of the world when the groups mingled more freely than they did later. Most of these tales were later told by Ovid's Metamorphoses and they are often divided into two thematic groups: tales of love, and tales of punishment. Tales of love often involve incest, or

9324-481: The appropriation or invention of some important cultural artifact, as when Prometheus steals fire from the gods, when Tantalus steals nectar and ambrosia from Zeus' table and gives it to his subjects—revealing to them the secrets of the gods, when Prometheus or Lycaon invents sacrifice, when Demeter teaches agriculture and the Mysteries to Triptolemus , or when Marsyas invents the aulos and enters into

9472-623: The army of the dead." Another important difference between the hero cult and the cult of gods is that the hero becomes the centre of local group identity. The monumental events of Heracles are regarded as the dawn of the age of heroes. To the Heroic Age are also ascribed three great events: the Argonautic expedition, the Theban Cycle , and the Trojan War . Some scholars believe that behind Heracles' complicated mythology there

9620-454: The arrival of Dionysus to establish his cult in Thrace was also the subject of an Aeschylean trilogy. In another tragedy, Euripides' The Bacchae , the king of Thebes , Pentheus , is punished by Dionysus, because he disrespected the god and spied on his Maenads , the female worshippers of the god. In another story, based on an old folktale-motif, and echoing a similar theme, Demeter

9768-548: The base of the chain to the top. This is because energy is lost to the environment with each transfer as entropy increases. About eighty to ninety percent of the energy is expended for the organism's life processes or is lost as heat or waste. Only about ten to twenty percent of the organism's energy is generally passed to the next organism. The amount can be less than one percent in animals consuming less digestible plants, and it can be as high as forty percent in zooplankton consuming phytoplankton . Graphic representations of

9916-430: The base. They can depict different numerical properties of ecosystems, including numbers of individuals per unit of area, biomass (g/m ), and energy (k cal m yr ). The emergent pyramidal arrangement of trophic levels with amounts of energy transfer decreasing as species become further removed from the source of production is one of several patterns that is repeated amongst the planets ecosystems. The size of each level in

10064-422: The basis for comparing and investigating the nature of non-random patterns in the structure of food web networks among many different types of ecosystems. Scaling laws, complexity, chaos, and pattern correlates are common features attributed to food web structure. Food webs are extremely complex. Complexity is a term that conveys the mental intractability of understanding all possible higher-order effects in

10212-590: The basis for the collection; however, the "Library" discusses events that occurred long after his death, hence the name Pseudo-Apollodorus. Among the earliest literary sources are Homer 's two epic poems, the Iliad and the Odyssey . Other poets completed the Epic Cycle , but these later and lesser poems now are lost almost entirely. Despite their traditional name, the Homeric Hymns have no direct connection with Homer. The oldest are choral hymns from

10360-409: The beginnings of the universe in human language. The most widely accepted version at the time, although a philosophical account of the beginning of things, is reported by Hesiod , in his Theogony . He begins with Chaos , a yawning nothingness. Next comes Gaia (Earth), "the ever-sure foundation of all", and then Tartarus , "in the depth of the wide-pathed Earth", and Eros (Love), "fairest among

10508-442: The biomass or productivity at each tropic level are called ecological pyramids or trophic pyramids. The transfer of energy from primary producers to top consumers can also be characterized by energy flow diagrams. A common metric used to quantify food web trophic structure is food chain length. Food chain length is another way of describing food webs as a measure of the number of species encountered as energy or nutrients move from

10656-413: The butterfly larvae. Another example of this sort of multitrophic interaction in plants is the transfer of defensive alkaloids produced by endophytes living within a grass host to a hemiparasitic plant that is also using the grass as a host. The Law of Conservation of Mass dates from Antoine Lavoisier's 1789 discovery that mass is neither created nor destroyed in chemical reactions. In other words,

10804-512: The composition of the story of the Argonauts is earlier than Odyssey , which shows familiarity with the exploits of Jason (the wandering of Odysseus may have been partly founded on it). In ancient times, the expedition was regarded as a historical fact, an incident in the opening up of the Black Sea to Greek commerce and colonization. It was also extremely popular, forming a cycle to which

10952-451: The concept and ritual. The age in which the heroes lived is known as the Heroic age . The epic and genealogical poetry created cycles of stories clustered around particular heroes or events and established the family relationships between the heroes of different stories; they thus arranged the stories in sequence. According to Ken Dowden (1992), "there is even a saga effect: We can follow

11100-581: The concept of food cycles, food chains, and food size in his classical 1927 book "Animal Ecology"; Elton's 'food cycle' was replaced by 'food web' in a subsequent ecological text. After Charles Elton's use of food webs in his 1927 synthesis, they became a central concept in the field of ecology . Elton organized species into functional groups , which formed the basis for the trophic system of classification in Raymond Lindeman 's classic and landmark paper in 1942 on trophic dynamics. The notion of

11248-400: The culture, arts, and literature of Western civilization and remains part of Western heritage and language. Poets and artists from ancient times to the present have derived inspiration from Greek mythology and have discovered contemporary significance and relevance in the themes. Greek mythology is known today primarily from Greek literature and representations on visual media dating from

11396-405: The dams. For amateur fishermen the river has been enriched with introduced rainbow trout, that is not easy to reproduce, so there is no danger of disturbance of the river's ecosystem . In the place where the river flows into the sea, there has been formed over the years an extended Delta of 4.000 hectares, because of the large dam that was constructed and retains much of the brought matters. As

11544-461: The dead. Influences from other cultures always afforded new themes. According to Classical-era mythology, after the overthrow of the Titans, the new pantheon of gods and goddesses was confirmed. Among the principal Greek gods were the Olympians, residing on Mount Olympus under the eye of Zeus. (The limitation of their number to twelve seems to have been a comparatively modern idea.) Besides

11692-514: The deathless gods". Without male assistance, Gaia gave birth to Uranus (the Sky) who then fertilized her. From that union were born first the Titans —six males: Coeus , Crius , Cronus , Hyperion , Iapetus , and Oceanus ; and six females: Mnemosyne , Phoebe , Rhea , Theia , Themis , and Tethys . After Cronus was born, Gaia and Uranus decreed no more Titans were to be born. They were followed by

11840-400: The decomposition actions of earthworms he talked about "the continued movement of the particles of earth". Even earlier, in 1768 John Bruckner described nature as "one continued web of life". Food webs are limited representations of real ecosystems as they necessarily aggregate many species into trophic species , which are functional groups of species that have the same predators and prey in

11988-473: The decoration of votive gifts and many other artifacts. Geometric designs on pottery of the eighth century BC depict scenes from the Epic Cycle as well as the adventures of Heracles . In the succeeding Archaic , Classical , and Hellenistic periods, Homeric and various other mythological scenes appear, supplementing the existing literary evidence. Greek mythology has had an extensive influence on

12136-1514: The detrital web and the grazing web. Mushrooms produced by decomposers in the detrital web become a food source for deer, squirrels, and mice in the grazing web. Earthworms eaten by robins are detritivores consuming decaying leaves. "Detritus can be broadly defined as any form of non-living organic matter, including different types of plant tissue (e.g. leaf litter , dead wood, aquatic macrophytes, algae), animal tissue (carrion), dead microbes, faeces (manure, dung, faecal pellets, guano, frass), as well as products secreted, excreted or exuded from organisms (e.g. extra-cellular polymers, nectar, root exudates and leachates , dissolved organic matter, extra-cellular matrix, mucilage). The relative importance of these forms of detritus, in terms of origin, size and chemical composition, varies across ecosystems." Ecologists collect data on trophic levels and food webs to statistically model and mathematically calculate parameters, such as those used in other kinds of network analysis (e.g., graph theory), to study emergent patterns and properties shared among ecosystems. There are different ecological dimensions that can be mapped to create more complicated food webs, including: species composition (type of species), richness (number of species), biomass (the dry weight of plants and animals), productivity (rates of conversion of energy and nutrients into growth), and stability (food webs over time). A food web diagram illustrating species composition shows how change in

12284-592: The diet of the most specialized species is a subset of the diet of the next more generalized species, and its diet a subset of the next more generalized, and so on." Until recently, it was thought that food webs had little nested structure, but empirical evidence shows that many published webs have nested subwebs in their assembly. Food webs are complex networks . As networks, they exhibit similar structural properties and mathematical laws that have been used to describe other complex systems, such as small world and scale free properties . The small world attribute refers to

12432-439: The diets of smaller predators tend to be nested subsets of those of larger predators (Woodward & Warren 2007; YvonDurocher et al. 2008), and phylogenetic constraints, whereby related taxa are nested based on their common evolutionary history, are also evident (Cattin et al. 2004)." "Compartments in food webs are subgroups of taxa in which many strong interactions occur within the subgroups and few weak interactions occur between

12580-624: The divine-focused Theogony and Homeric Hymns in both size and popularity. Under the influence of Homer the "hero cult" leads to a restructuring in spiritual life, expressed in the separation of the realm of the gods from the realm of the dead (heroes), of the Chthonic from the Olympian. In the Works and Days , Hesiod makes use of a scheme of Four Ages of Man (or Races): Golden, Silver, Bronze, and Iron. These races or ages are separate creations of

12728-480: The earlier part of the so-called Lyric age . Hesiod , a possible contemporary with Homer, offers in his Theogony ( Origin of the Gods ) the fullest account of the earliest Greek myths, dealing with the creation of the world, the origin of the gods, Titans , and Giants , as well as elaborate genealogies, folktales, and aetiological myths. Hesiod's Works and Days , a didactic poem about farming life, also includes

12876-458: The ecosystem concept, which assumes that the phenomena under investigation (interactions and feedback loops) are sufficient to explain patterns within boundaries, such as the edge of a forest, an island, a shoreline, or some other pronounced physical characteristic. In a detrital web, plant and animal matter is broken down by decomposers, e.g., bacteria and fungi, and moves to detritivores and then carnivores. There are often relationships between

13024-450: The ecosystem to another. The trophic dynamic concept has served as a useful quantitative heuristic, but it has several major limitations including the precision by which an organism can be allocated to a specific trophic level. Omnivores, for example, are not restricted to any single level. Nonetheless, recent research has found that discrete trophic levels do exist, but "above the herbivore trophic level, food webs are better characterized as

13172-476: The evolution of their culture, of which mythology, both overtly and in its unspoken assumptions, is an index of the changes. In Greek mythology's surviving literary forms, as found mostly at the end of the progressive changes, it is inherently political, as Gilbert Cuthbertson (1975) has argued. The earlier inhabitants of the Balkan Peninsula were an agricultural people who, using animism , assigned

13320-405: The fates of some families in successive generations." After the rise of the hero cult, gods and heroes constitute the sacral sphere and are invoked together in oaths and prayers which are addressed to them. Burkert (2002) notes that "the roster of heroes, again in contrast to the gods, is never given fixed and final form. Great gods are no longer born, but new heroes can always be raised up from

13468-509: The first known representation of a myth in geometric art predates its first known representation in late archaic poetry, by several centuries. In the Archaic ( c.  750  – c.  500 BC ), Classical ( c.  480 –323 BC), and Hellenistic (323–146 BC) periods, Homeric and various other mythological scenes appear, supplementing the existing literary evidence. Greek mythology has changed over time to accommodate

13616-472: The gods, the Golden Age belonging to the reign of Cronos, the subsequent races to the creation of Zeus . The presence of evil was explained by the myth of Pandora , when all of the best of human capabilities, save hope, had been spilled out of her overturned jar. In Metamorphoses , Ovid follows Hesiod's concept of the four ages. "Myths of origin" or " creation myths " represent an attempt to explain

13764-541: The heel. Achilles' heel was the only part of his body which was not invulnerable to damage by human weaponry. Before they could take Troy, the Greeks had to steal from the citadel the wooden image of Pallas Athena (the Palladium ). Finally, with Athena's help, they built the Trojan Horse . Despite the warnings of Priam's daughter Cassandra , the Trojans were persuaded by Sinon , a Greek who feigned desertion, to take

13912-729: The highest social prestige through his appointment as official ancestor of the Dorian kings. This probably served as a legitimation for the Dorian migrations into the Peloponnese . Hyllus , the eponymous hero of one Dorian phyle , became the son of Heracles and one of the Heracleidae or Heraclids (the numerous descendants of Heracles, especially the descendants of Hyllus —other Heracleidae included Macaria , Lamos, Manto , Bianor , Tlepolemus , and Telephus ). These Heraclids conquered

14060-420: The horse inside the walls of Troy as an offering to Athena; the priest Laocoon, who tried to have the horse destroyed, was killed by sea-serpents. At night the Greek fleet returned, and the Greeks from the horse opened the gates of Troy. In the total sack that followed, Priam and his remaining sons were slaughtered; the Trojan women passed into slavery in various cities of Greece. The adventurous homeward voyages of

14208-496: The influence is environmentally context dependent. These complex multitrophic interactions involve more than two trophic levels in a food web. For example, such interactions have been discovered in the context of arbuscular mycorrhizal fungi and aphid herbivores that utilize the same plant species . Another example of a multitrophic interaction is a trophic cascade , in which predators help to increase plant growth and prevent overgrazing by suppressing herbivores. Links in

14356-431: The inverted pyramidal pattern. Population structure, migration rates, and environmental refuge for prey are other possible causes for pyramids with biomass inverted. Energy pyramids, however, will always have an upright pyramid shape if all sources of food energy are included and this is dictated by the second law of thermodynamics . Many of the Earth's elements and minerals (or mineral nutrients) are contained within

14504-618: The larger influences where the bulk of energy transfer occurs. "These omissions and problems are causes for concern, but on present evidence do not present insurmountable difficulties." There are different kinds or categories of food webs: Within these categories, food webs can be further organized according to the different kinds of ecosystems being investigated. For example, human food webs, agricultural food webs, detrital food webs, marine food webs , aquatic food webs, soil food webs , Arctic (or polar) food webs, terrestrial food webs, and microbial food webs . These characterizations stem from

14652-594: The many loosely connected nodes, non-random dense clustering of a few nodes (i.e., trophic or keystone species in ecology), and small path length compared to a regular lattice. "Ecological networks, especially mutualistic networks, are generally very heterogeneous, consisting of areas with sparse links among species and distinct areas of tightly linked species. These regions of high link density are often referred to as cliques, hubs, compartments, cohesive sub-groups, or modules...Within food webs, especially in aquatic systems, nestedness appears to be related to body size because

14800-436: The mass of any one element at the beginning of a reaction will equal the mass of that element at the end of the reaction. Food webs depict energy flow via trophic linkages. Energy flow is directional, which contrasts against the cyclic flows of material through the food web systems. Energy flow "typically includes production, consumption, assimilation, non-assimilation losses (feces), and respiration (maintenance costs)." In

14948-576: The middle of the Archaic period, myths about relationships between male gods and male heroes became more and more frequent, indicating the parallel development of pedagogic pederasty ( παιδικὸς ἔρως , eros paidikos ), thought to have been introduced around 630 BC. By the end of the fifth-century  BC, poets had assigned at least one eromenos , an adolescent boy who was their sexual companion, to every important god except Ares and many legendary figures. Previously existing myths, such as those of Achilles and Patroclus , also then were cast in

15096-577: The myth of the voyage of Jason and the Argonauts to retrieve the Golden Fleece from the mythical land of Colchis . In the Argonautica , Jason is impelled on his quest by king Pelias , who receives a prophecy that a man with one sandal would be his nemesis . Jason loses a sandal in a river, arrives at the court of Pelias, and the epic is set in motion. Nearly every member of the next generation of heroes, as well as Heracles, went with Jason in

15244-479: The mythological details about gods and heroes. The evidence about myths and rituals at Mycenaean and Minoan sites is entirely monumental, as the Linear B script (an ancient form of Greek found in both Crete and mainland Greece) was used mainly to record inventories, although certain names of gods and heroes have been tentatively identified. Geometric designs on pottery of the eighth-century  BC depict scenes from

15392-530: The myths of Prometheus , Pandora , and the Five Ages . The poet advises on the best way to succeed in a dangerous world, rendered yet more dangerous by its gods. Lyrical poets often took their subjects from myth, but their treatment became gradually less narrative and more allusive. Greek lyric poets, including Pindar , Bacchylides and Simonides , and bucolic poets such as Theocritus and Bion , relate individual mythological incidents. Additionally, myth

15540-598: The northeast, formed the boundary between Pieria , Eordaea , and Imathia . In the time of Herodotus the Haliacmon was apparently joined by the Loudias 7, 127 , the discharge of the lake of Pella ; but a change has taken place in the course of the Loudias, which no longer joins the Haliacmon, but flows directly into the Aegean Sea . The image below shows a wind gap between the Haliacmon and Loudias watersheds that

15688-403: The number of empirical studies on community webs is on the rise and the mathematical treatment of food webs using network theory had identified patterns that are common to all. Scaling laws , for example, predict a relationship between the topology of food web predator-prey linkages and levels of species richness . Food webs are the road-maps through Darwin's famous 'entangled bank' and have

15836-414: The number of trophic levels in more species rich ecosystems. This hypothesis was challenged through mathematical models suggesting otherwise, but subsequent studies have shown that the premise holds in real systems. At different levels in the hierarchy of life, such as the stability of a food web, "the same overall structure is maintained in spite of an ongoing flow and change of components." The farther

15984-426: The nutrient and energy stores. "Organisms usually extract energy in the form of carbohydrates, lipids, and proteins. These polymers have a dual role as supplies of energy as well as building blocks; the part that functions as energy supply results in the production of nutrients (and carbon dioxide, water, and heat). Excretion of nutrients is, therefore, basic to metabolism." The units in energy flow webs are typically

16132-492: The one-eyed Cyclopes and the Hecatoncheires or Hundred-Handed Ones, who were both thrown into Tartarus by Uranus. This made Gaia furious. Cronus ("the wily, youngest and most terrible of Gaia 's children") was convinced by Gaia to castrate his father. He did this and became the ruler of the Titans with his sister-wife, Rhea, as his consort, and the other Titans became his court. A motif of father-against-son conflict

16280-473: The only general mythographical handbook to survive from Greek antiquity was the Library of Pseudo-Apollodorus. This work attempts to reconcile the contradictory tales of the poets and provides a grand summary of traditional Greek mythology and heroic legends. Apollodorus of Athens lived from c.  180 BC to c.  125 BC and wrote on many of these topics. His writings may have formed

16428-408: The organic matter eaten by heterotrophs, such as sugars , provides energy. Autotrophs and heterotrophs come in all sizes, from microscopic to many tonnes - from cyanobacteria to giant redwoods , and from viruses and bdellovibrio to blue whales . Charles Elton pioneered the concept of food cycles, food chains, and food size in his classical 1927 book "Animal Ecology"; Elton's 'food cycle'

16576-697: The origin of sacrificial practices. Myths are also preserved in the Homeric Hymns , in fragments of epic poems of the Epic Cycle , in lyric poems , in the works of the tragedians and comedians of the fifth century BC, in writings of scholars and poets of the Hellenistic Age , and in texts from the time of the Roman Empire by writers such as Plutarch and Pausanias . Aside from this narrative deposit in ancient Greek literature , pictorial representations of gods, heroes, and mythic episodes featured prominently in ancient vase paintings and

16724-417: The plants to top predators. There are different ways of calculating food chain length depending on what parameters of the food web dynamic are being considered: connectance, energy, or interaction. In its simplest form, the length of a chain is the number of links between a trophic consumer and the base of the web. The mean chain length of an entire web is the arithmetic average of the lengths of all chains in

16872-459: The problem of the devolution of power and of the mode of accession to sovereignty. The twins Atreus and Thyestes with their descendants played the leading role in the tragedy of the devolution of power in Mycenae. The Theban Cycle deals with events associated especially with Cadmus , the city's founder, and later with the doings of Laius and Oedipus at Thebes; a series of stories that lead to

17020-437: The pyramid generally represents biomass, which can be measured as the dry weight of an organism. Autotrophs may have the highest global proportion of biomass, but they are closely rivaled or surpassed by microbes. Pyramid structure can vary across ecosystems and across time. In some instances biomass pyramids can be inverted. This pattern is often identified in aquatic and coral reef ecosystems. The pattern of biomass inversion

17168-424: The pyramid. Primary consumers have longer lifespans and slower growth rates that accumulates more biomass than the producers they consume. Phytoplankton live just a few days, whereas the zooplankton eating the phytoplankton live for several weeks and the fish eating the zooplankton live for several consecutive years. Aquatic predators also tend to have a lower death rate than the smaller consumers, which contributes to

17316-534: The rarest birds in the world. 27 kinds of rare and protected kinds of birds nest in the region, like purple heron . In antiquity, Claudius Ptolemy called the chain of mountains in which the river rises (northern Pindus ) the Canalovii . According to Julius Caesar , the Haliacmon formed the line of demarcation between Macedon and Thessaly . In the upper part of its course it took a southeast direction through Elimiotis , which it watered; and then, continuing to

17464-439: The rest were introduced by human intervention. Many of them are considered scarce and one, mavrotsironi , lives nowhere else in the world, i.e. it is endemic . Μost of these kinds of fish have no commercial value but only biological, since they support the food web . Fish found in Haliakmon, like carp and rainbow trout , indicate that its water is still pure. Some eels are found in its estuaries, that cannot migrate, hindered by

17612-564: The role of mixotrophs , or autotrophs that additionally obtain organic matter from non-atmospheric sources. The linkages in a food web illustrate the feeding pathways, such as where heterotrophs obtain organic matter by feeding on autotrophs and other heterotrophs. The food web is a simplified illustration of the various methods of feeding that link an ecosystem into a unified system of exchange. There are different kinds of consumer–resource interactions that can be roughly divided into herbivory , carnivory , scavenging , and parasitism . Some of

17760-466: The same predators and prey in a food web. Common examples of an aggregated node in a food web might include parasites , microbes, decomposers , saprotrophs , consumers , or predators , each containing many species in a web that can otherwise be connected to other trophic species. Food webs have trophic levels and positions. Basal species, such as plants, form the first level and are the resource-limited species that feed on no other living creature in

17908-510: The seduction or rape of a mortal woman by a male god, resulting in heroic offspring. The stories generally suggest that relationships between gods and mortals are something to avoid; even consenting relationships rarely have happy endings. In a few cases, a female divinity mates with a mortal man, as in the Homeric Hymn to Aphrodite , where the goddess lies with Anchises to produce Aeneas . The second type (tales of punishment) involves

18056-593: The ship Argo to fetch the Golden Fleece. This generation also included Theseus , who went to Crete to slay the Minotaur ; Atalanta , the female heroine, and Meleager , who once had an epic cycle of his own to rival the Iliad and Odyssey . Pindar , Apollonius and the Bibliotheca endeavor to give full lists of the Argonauts. Although Apollonius wrote his poem in the 3rd century BC,

18204-483: The society while the beliefs were held. After they ceased to become religious beliefs, few would have known the rites and rituals. Allusions often existed, however, to aspects that were quite public. Images existed on pottery and religious artwork that were interpreted and more likely, misinterpreted in many diverse myths and tales. A few fragments of these works survive in quotations by Neoplatonist philosophers and recently unearthed papyrus scraps. One of these scraps,

18352-615: The stone, which had been sitting in Cronus's stomach all this time. Zeus then challenged Cronus to war for the kingship of the gods. At last, with the help of the Cyclopes (whom Zeus freed from Tartarus), Zeus and his siblings were victorious, while Cronus and the Titans were hurled down to imprisonment in Tartarus . Zeus was plagued by the same concern, and after a prophecy that the offspring of his first wife, Metis , would give birth to

18500-406: The stony hearts of the underworld gods in his descent to Hades . When Hermes invents the lyre in the Homeric Hymn to Hermes , the first thing he does is sing about the birth of the gods. Hesiod's Theogony is not only the fullest surviving account of the gods but also the fullest surviving account of the archaic poet's function, with its long preliminary invocation to the Muses . Theogony also

18648-406: The structure, stability, and laws of food web behaviours relative to observable outcomes. "Food web theory centers around the idea of connectance." Quantitative formulas simplify the complexity of food web structure. The number of trophic links (t L ), for example, is converted into a connectance value: where, S(S-1)/2 is the maximum number of binary connections among S species. "Connectance (C)

18796-405: The study of complexity in food webs. Complexity explains many principals pertaining to self-organization, non-linearity, interaction, cybernetic feedback, discontinuity, emergence, and stability in food webs. Nestedness, for example, is defined as "a pattern of interaction in which specialists interact with species that form perfect subsets of the species with which generalists interact", "—that is,

18944-508: The subgroups. Theoretically, compartments increase the stability in networks, such as food webs." Food webs are also complex in the way that they change in scale, seasonally, and geographically. The components of food webs, including organisms and mineral nutrients, cross the thresholds of ecosystem boundaries. This has led to the concept or area of study known as cross-boundary subsidy . "This leads to anomalies, such as food web calculations determining that an ecosystem can support one half of

19092-423: The sulfur bacterium Thiobacillus , which lives in hot sulfur springs . The top level has top (or apex) predators that no other species kills directly for their food resource needs. The intermediate levels are filled with omnivores that feed on more than one trophic level and cause energy to flow through several food pathways starting from a basal species. In the simplest scheme, the first trophic level (level 1)

19240-473: The tale known to us through tragedy (e.g. Sophocles' Oedipus Rex ) and later mythological accounts. Greek mythology culminates in the Trojan War, fought between Greece and Troy , and its aftermath. In Homer's works, such as the Iliad , the chief stories have already taken shape and substance, and individual themes were elaborated later, especially in Greek drama. The Trojan War also elicited great interest in

19388-427: The tissues and diets of organisms. Hence, mineral and nutrient cycles trace food web energy pathways. Ecologists employ stoichiometry to analyze the ratios of the main elements found in all organisms: carbon (C), nitrogen (N), phosphorus (P). There is a large transitional difference between many terrestrial and aquatic systems as C:P and C:N ratios are much higher in terrestrial systems while N:P ratios are equal between

19536-587: The towns (in downstream order) Nestorio , Argos Orestiko , Neapoli , Paliouria , Velventos and Alexandreia . Haliakmon contains 33 kinds of fish. These include brown trout , gilt-head bream , sardine , carp , pike, eel , european anchovy , nursehound , bogue , garfish , red porgy , saddled seabream , angler , bream, mediterranean sand smelt , sand steenbras , mullus barbatus , freshwater bass, tuna , salmon , Mediterranean moray , weever , longfin gurnard , dogfish , dusky grouper , school shark and turbot . About 30 of them are indigenous, while

19684-620: The transfer of defensive compounds across multiple trophic levels. For example, certain plant species in the Castilleja and Plantago genera have been found to produce defensive compounds called iridoid glycosides that are sequestered in the tissues of the Taylor's checkerspot butterfly larvae that have developed a tolerance for these compounds and are able to consume the foliage of these plants. These sequestered iridoid glycosides then confer chemical protection against bird predators to

19832-422: The two systems. Mineral nutrients are the material resources that organisms need for growth, development, and vitality. Food webs depict the pathways of mineral nutrient cycling as they flow through organisms. Most of the primary production in an ecosystem is not consumed, but is recycled by detritus back into useful nutrients. Many of the Earth's microorganisms are involved in the formation of minerals in

19980-422: The use of stable isotopes to better trace energy flow through the web. It was once thought that omnivory was rare, but recent evidence suggests otherwise. This realization has made trophic classifications more complex. The trophic level concept was introduced in a historical landmark paper on trophic dynamics in 1942 by Raymond L. Lindeman . The basis of trophic dynamics is the transfer of energy from one part of

20128-577: The war of the Seven against Thebes and the eventual pillage of that city at the hands of the Epigoni . (It is not known whether the Seven figured in early epic.) As far as Oedipus is concerned, early epic accounts seem to have him continuing to rule at Thebes after the revelation that Iokaste was his mother, and subsequently marrying a second wife who becomes the mother of his children—markedly different from

20276-520: The web are called trophic links. The number of trophic links per consumer is a measure of food web connectance . Food chains are nested within the trophic links of food webs. Food chains are linear (noncyclic) feeding pathways that trace monophagous consumers from a base species up to the top consumer , which is usually a larger predatory carnivore. Linkages connect to nodes in a food web, which are aggregates of biological taxa called trophic species . Trophic species are functional groups that have

20424-467: The web. Basal species can be autotrophs or detritivores , including "decomposing organic material and its associated microorganisms which we defined as detritus, micro-inorganic material and associated microorganisms (MIP), and vascular plant material." Most autotrophs capture the sun's energy in chlorophyll , but some autotrophs (the chemolithotrophs ) obtain energy by the chemical oxidation of inorganic compounds and can grow in dark environments, such as

20572-434: The world" may be divided into three or four broader periods: While the age of gods often has been of more interest to contemporary students of myth, the Greek authors of the archaic and classical eras had a clear preference for the age of heroes, establishing a chronology and record of human accomplishments after the questions of how the world came into being were explained. For example, the heroic Iliad and Odyssey dwarfed

20720-598: Was central to classical Athenian drama . The tragic playwrights Aeschylus , Sophocles , and Euripides took most of their plots from myths of the age of heroes and the Trojan War. Many of the great tragic stories (e.g. Agamemnon and his children, Oedipus , Jason , Medea , etc.) took on their classic form in these tragedies. The comic playwright Aristophanes also used myths, in The Birds and The Frogs . Historians Herodotus and Diodorus Siculus , and geographers Pausanias and Strabo , who traveled throughout

20868-478: Was insured by the constant use of nectar and ambrosia , by which the divine blood was renewed in their veins. Each god descends from his or her own genealogy, pursues differing interests, has a certain area of expertise, and is governed by a unique personality; however, these descriptions arise from a multiplicity of archaic local variants, which do not always agree with one another. When these gods are called upon in poetry, prayer, or cult, they are referred to by

21016-592: Was probably a real man, perhaps a chieftain-vassal of the kingdom of Argos . Some scholars suggest the story of Heracles is an allegory for the sun's yearly passage through the twelve constellations of the zodiac. Others point to earlier myths from other cultures, showing the story of Heracles as a local adaptation of hero myths already well established. Traditionally, Heracles was the son of Zeus and Alcmene , granddaughter of Perseus . His fantastic solitary exploits, with their many folk-tale themes, provided much material for popular legend. According to Burkert (2002), "He

21164-511: Was repeated when Cronus was confronted by his son, Zeus . Because Cronus had betrayed his father, he feared that his offspring would do the same, and so each time Rhea gave birth, he snatched up the child and ate it. Rhea hated this and tricked him by hiding Zeus and wrapping a stone in a baby's blanket, which Cronus ate. When Zeus was full-grown, he fed Cronus a drugged drink which caused him to vomit, throwing up Rhea's other children, including Poseidon , Hades , Hestia , Demeter , and Hera , and

21312-520: Was replaced by 'food web' in a subsequent ecological text. Elton organized species into functional groups , which was the basis for Raymond Lindeman 's classic and landmark paper in 1942 on trophic dynamics. Lindeman emphasized the important role of decomposer organisms in a trophic system of classification . The notion of a food web has a historical foothold in the writings of Charles Darwin and his terminology, including an "entangled bank", "web of life", "web of complex relations", and in reference to

21460-591: Was searching for her daughter, Persephone , having taken the form of an old woman called Doso, and received a hospitable welcome from Celeus , the King of Eleusis in Attica . As a gift to Celeus, because of his hospitality, Demeter planned to make his son Demophon a god, but she was unable to complete the ritual because his mother Metanira walked in and saw her son in the fire and screamed in fright, which angered Demeter, who lamented that foolish mortals do not understand

21608-575: Was the bow but frequently also the club. Vase paintings demonstrate the unparalleled popularity of Heracles, his fight with the lion being depicted many hundreds of times. Heracles also entered Etruscan and Roman mythology and cult, and the exclamation "mehercule" became as familiar to the Romans as "Herakleis" was to the Greeks. In Italy he was worshipped as a god of merchants and traders, although others also prayed to him for his characteristic gifts of good luck or rescue from danger. Heracles attained

21756-431: Was the goddess of love and beauty, Ares was the god of war, Hades the ruler of the underworld, and Athena the goddess of wisdom and courage. Some gods, such as Apollo and Dionysus , revealed complex personalities and mixtures of functions, while others, such as Hestia (literally "hearth") and Helios (literally "sun"), were little more than personifications. The most impressive temples tended to be dedicated to

21904-503: Was the subject of many lost poems, including those attributed to Orpheus, Musaeus , Epimenides , Abaris , and other legendary seers, which were used in private ritual purifications and mystery-rites . There are indications that Plato was familiar with some version of the Orphic theogony. A silence would have been expected about religious rites and beliefs, however, and that nature of the culture would not have been reported by members of

#383616