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122-460: It is located in the state's desert region (Región Desierto) . Cuatro Ciénegas is Spanish for "four marshes "; the name was chosen by the first settlers because of the natural springs in the vicinity that create extensive areas of wetland and lakes. Archeological excavations indicate settlement in the area from approximately 5000 years ago, as well as ritual use of peyote by the inhabitants. Several failed settlements were founded here prior to

244-409: A cell wall . Newly dead animals may be covered by an exoskeleton . Fragmentation processes, which break through these protective layers, accelerate the rate of microbial decomposition. Animals fragment detritus as they hunt for food, as does passage through the gut. Freeze-thaw cycles and cycles of wetting and drying also fragment dead material. The chemical alteration of the dead organic matter

366-495: A food chain . Real systems are much more complex than this—organisms will generally feed on more than one form of food, and may feed at more than one trophic level. Carnivores may capture some prey that is part of a plant-based trophic system and others that are part of a detritus-based trophic system (a bird that feeds both on herbivorous grasshoppers and earthworms, which consume detritus). Real systems, with all these complexities, form food webs rather than food chains which present

488-619: A habitat . Ecosystem ecology is the "study of the interactions between organisms and their environment as an integrated system". The size of ecosystems can range up to ten orders of magnitude , from the surface layers of rocks to the surface of the planet. The Hubbard Brook Ecosystem Study started in 1963 to study the White Mountains in New Hampshire . It was the first successful attempt to study an entire watershed as an ecosystem. The study used stream chemistry as

610-524: A 19th-century Italian immigrant near the town of Cuatro Ciénegas, is the second largest producer of wine in Coahuila. The site attracts many visitors who are immersed in the process of making wine and it offers guide tours around the Cuatro Ciénegas valley . Scientists working in the basin and some local residents claim to have observed reduced spring discharge and a decrease in surface water in

732-615: A Spanish-based creole language called Chavacano developed in the southern Philippines. However, it is not mutually intelligible with Spanish. The number of Chavacano-speakers was estimated at 1.2 million in 1996. The local languages of the Philippines also retain significant Spanish influence, with many words derived from Mexican Spanish , owing to the administration of the islands by Spain through New Spain until 1821, until direct governance from Madrid afterwards to 1898. Ecosystem An ecosystem (or ecological system )

854-609: A central role over a wide range, for example, in the slow development of soil from bare rock and the faster recovery of a community from disturbance . Disturbance also plays an important role in ecological processes. F. Stuart Chapin and coauthors define disturbance as "a relatively discrete event in time that removes plant biomass". This can range from herbivore outbreaks, treefalls, fires, hurricanes, floods, glacial advances , to volcanic eruptions . Such disturbances can cause large changes in plant, animal and microbe populations, as well as soil organic matter content. Disturbance

976-603: A considerable number of words from Arabic , as well as a minor influence from the Germanic Gothic language through the period of Visigoth rule in Iberia. In addition, many more words were borrowed from Latin through the influence of written language and the liturgical language of the Church. The loanwords were taken from both Classical Latin and Renaissance Latin , the form of Latin in use at that time. According to

1098-507: A critical role in global nutrient cycling and ecosystem function. Phosphorus enters ecosystems through weathering . As ecosystems age this supply diminishes, making phosphorus-limitation more common in older landscapes (especially in the tropics). Calcium and sulfur are also produced by weathering, but acid deposition is an important source of sulfur in many ecosystems. Although magnesium and manganese are produced by weathering, exchanges between soil organic matter and living cells account for

1220-522: A dramatic change in the pronunciation of its sibilant consonants , known in Spanish as the reajuste de las sibilantes , which resulted in the distinctive velar [x] pronunciation of the letter ⟨j⟩ and—in a large part of Spain—the characteristic interdental [θ] ("th-sound") for the letter ⟨z⟩ (and for ⟨c⟩ before ⟨e⟩ or ⟨i⟩ ). See History of Spanish (Modern development of

1342-517: A faster recovery. More severe and more frequent disturbance result in longer recovery times. From one year to another, ecosystems experience variation in their biotic and abiotic environments. A drought , a colder than usual winter, and a pest outbreak all are short-term variability in environmental conditions. Animal populations vary from year to year, building up during resource-rich periods and crashing as they overshoot their food supply. Longer-term changes also shape ecosystem processes. For example,

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1464-482: A first language by Spaniards and educated Filipinos ( Ilustrados ). Despite a public education system set up by the colonial government, by the end of Spanish rule in 1898, only about 10% of the population had knowledge of Spanish, mostly those of Spanish descent or elite standing. Spanish continued to be official and used in Philippine literature and press during the early years of American administration after

1586-427: A function-based typology has been proposed to leverage the strengths of these different approaches into a unified system. Human activities are important in almost all ecosystems. Although humans exist and operate within ecosystems, their cumulative effects are large enough to influence external factors like climate. Ecosystems provide a variety of goods and services upon which people depend. Ecosystem goods include

1708-698: A general level, for example, tropical forests , temperate grasslands , and arctic tundra . There can be any degree of subcategories among ecosystem types that comprise a biome, e.g., needle-leafed boreal forests or wet tropical forests. Although ecosystems are most commonly categorized by their structure and geography, there are also other ways to categorize and classify ecosystems such as by their level of human impact (see anthropogenic biome ), or by their integration with social processes or technological processes or their novelty (e.g. novel ecosystem ). Each of these taxonomies of ecosystems tends to emphasize different structural or functional properties. None of these

1830-478: A long history in the territory of the current-day United States dating back to the 16th century. In the wake of the 1848 Guadalupe Hidalgo Treaty , hundreds of thousands of Spanish speakers became a minoritized community in the United States. The 20th century saw further massive growth of Spanish speakers in areas where they had been hitherto scarce. According to the 2020 census, over 60 million people of

1952-672: A means of monitoring ecosystem properties, and developed a detailed biogeochemical model of the ecosystem. Long-term research at the site led to the discovery of acid rain in North America in 1972. Researchers documented the depletion of soil cations (especially calcium) over the next several decades. Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Studies can be carried out at

2074-528: A more important role in moving nutrients around. This can be especially important as the soil thaws in the spring, creating a pulse of nutrients that become available. Decomposition rates are low under very wet or very dry conditions. Decomposition rates are highest in wet, moist conditions with adequate levels of oxygen. Wet soils tend to become deficient in oxygen (this is especially true in wetlands ), which slows microbial growth. In dry soils, decomposition slows as well, but bacteria continue to grow (albeit at

2196-406: A number of common, non random properties in the topology of their network. The carbon and nutrients in dead organic matter are broken down by a group of processes known as decomposition. This releases nutrients that can then be re-used for plant and microbial production and returns carbon dioxide to the atmosphere (or water) where it can be used for photosynthesis. In the absence of decomposition,

2318-508: A primarily Hassaniya Arabic -speaking territory, Spanish was officially spoken as the language of the colonial administration during the late 19th and 20th centuries. Today, Spanish is present in the partially-recognized Sahrawi Arab Democratic Republic as its secondary official language, and in the Sahrawi refugee camps in Tindouf ( Algeria ), where the Spanish language is still taught as

2440-421: A process known as denitrification . Mycorrhizal fungi which are symbiotic with plant roots, use carbohydrates supplied by the plants and in return transfer phosphorus and nitrogen compounds back to the plant roots. This is an important pathway of organic nitrogen transfer from dead organic matter to plants. This mechanism may contribute to more than 70 Tg of annually assimilated plant nitrogen, thereby playing

2562-617: A second language features characteristics involving the variability of the vowel system. While far from its heyday during the Spanish protectorate in Morocco , the Spanish language has some presence in northern Morocco , stemming for example from the availability of certain Spanish-language media. According to a 2012 survey by Morocco's Royal Institute for Strategic Studies (IRES), penetration of Spanish in Morocco reaches 4.6% of

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2684-531: A second language, largely by Cuban educators. The number of Spanish speakers is unknown. Spanish is also an official language of the African Union . Spanish was an official language of the Philippines from the beginning of Spanish administration in 1565 to a constitutional change in 1973. During Spanish colonization , it was the language of government, trade, and education, and was spoken as

2806-696: A significant portion of ecosystem fluxes. Potassium is primarily cycled between living cells and soil organic matter. Biodiversity plays an important role in ecosystem functioning. Ecosystem processes are driven by the species in an ecosystem, the nature of the individual species, and the relative abundance of organisms among these species. Ecosystem processes are the net effect of the actions of individual organisms as they interact with their environment. Ecological theory suggests that in order to coexist, species must have some level of limiting similarity —they must be different from one another in some fundamental way, otherwise, one species would competitively exclude

2928-408: A slower rate) even after soils become too dry to support plant growth. Ecosystems are dynamic entities. They are subject to periodic disturbances and are always in the process of recovering from past disturbances. When a perturbation occurs, an ecosystem responds by moving away from its initial state. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance,

3050-464: A small effect on ecosystem function. Ecologically distinct species, on the other hand, have a much larger effect. Similarly, dominant species have a large effect on ecosystem function, while rare species tend to have a small effect. Keystone species tend to have an effect on ecosystem function that is disproportionate to their abundance in an ecosystem. An ecosystem engineer is any organism that creates, significantly modifies, maintains or destroys

3172-468: A variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Biomes are general classes or categories of ecosystems. However, there is no clear distinction between biomes and ecosystems. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of

3294-443: A variety of goods and services upon which people depend, and may be part of. Ecosystem goods include the "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like the maintenance of hydrological cycles , cleaning air and water,

3416-522: A variety of scales, ranging from whole-ecosystem studies to studying microcosms or mesocosms (simplified representations of ecosystems). American ecologist Stephen R. Carpenter has argued that microcosm experiments can be "irrelevant and diversionary" if they are not carried out in conjunction with field studies done at the ecosystem scale. In such cases, microcosm experiments may fail to accurately predict ecosystem-level dynamics. Biomes are general classes or categories of ecosystems. However, there

3538-546: Is Mexico . Spanish is part of the Ibero-Romance language group , in which the language is also known as Castilian ( castellano ). The group evolved from several dialects of Vulgar Latin in Iberia after the collapse of the Western Roman Empire in the 5th century. The oldest Latin texts with traces of Spanish come from mid-northern Iberia in the 9th century, and the first systematic written use of

3660-416: Is cognate to the root word of satisfacer ("to satisfy"), and hecho ("made") is similarly cognate to the root word of satisfecho ("satisfied"). Compare the examples in the following table: Some consonant clusters of Latin also produced characteristically different results in these languages, as shown in the examples in the following table: In the 15th and 16th centuries, Spanish underwent

3782-431: Is a system that environments and their organisms form through their interaction. The biotic and abiotic components are linked together through nutrient cycles and energy flows. Ecosystems are controlled by external and internal factors . External factors such as climate , parent material which forms the soil and topography , control the overall structure of an ecosystem but are not themselves influenced by

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3904-413: Is also one of the most taught foreign languages throughout the world. Spanish is well represented in the humanities and social sciences . Spanish is also the third most used language on the internet by number of users after English and Chinese and the second most used language by number of websites after English. Spanish is used as an official language by many international organizations , including

4026-673: Is an accepted version of this page Spanish ( español ) or Castilian ( castellano ) is a Romance language of the Indo-European language family that evolved from the Vulgar Latin spoken on the Iberian Peninsula of Europe . Today, it is a global language with about 500 million native speakers, mainly in the Americas and Spain , and about 600 million when including second language speakers. Spanish

4148-458: Is an international synthesis by over 1000 of the world's leading biological scientists that analyzes the state of the Earth's ecosystems and provides summaries and guidelines for decision-makers. The report identified four major categories of ecosystem services: provisioning, regulating, cultural and supporting services. It concludes that human activity is having a significant and escalating impact on

4270-442: Is consumed by animals while still alive and enters the plant-based trophic system. After plants and animals die, the organic matter contained in them enters the detritus-based trophic system. Ecosystem respiration is the sum of respiration by all living organisms (plants, animals, and decomposers) in the ecosystem. Net ecosystem production is the difference between gross primary production (GPP) and ecosystem respiration. In

4392-454: Is estimated that about 486 million people speak Spanish as a native language , making it the second most spoken language by number of native speakers . An additional 75 million speak Spanish as a second or foreign language , making it the fourth most spoken language in the world overall after English, Mandarin Chinese, and Hindi with a total number of 538 million speakers. Spanish is also

4514-569: Is followed by succession, a "directional change in ecosystem structure and functioning resulting from biotically driven changes in resource supply." The frequency and severity of disturbance determine the way it affects ecosystem function. A major disturbance like a volcanic eruption or glacial advance and retreat leave behind soils that lack plants, animals or organic matter. Ecosystems that experience such disturbances undergo primary succession . A less severe disturbance like forest fires, hurricanes or cultivation result in secondary succession and

4636-554: Is governed by three sets of factors—the physical environment (temperature, moisture, and soil properties), the quantity and quality of the dead material available to decomposers, and the nature of the microbial community itself. Temperature controls the rate of microbial respiration; the higher the temperature, the faster the microbial decomposition occurs. Temperature also affects soil moisture, which affects decomposition. Freeze-thaw cycles also affect decomposition—freezing temperatures kill soil microorganisms, which allows leaching to play

4758-761: Is heavily influenced by Venezuelan Spanish. In addition to sharing most of its borders with Spanish-speaking countries, the creation of Mercosur in the early 1990s induced a favorable situation for the promotion of Spanish language teaching in Brazil . In 2005, the National Congress of Brazil approved a bill, signed into law by the President , making it mandatory for schools to offer Spanish as an alternative foreign language course in both public and private secondary schools in Brazil. In September 2016 this law

4880-405: Is no clear distinction between biomes and ecosystems. Biomes are always defined at a very general level. Ecosystems can be described at levels that range from very general (in which case the names are sometimes the same as those of biomes) to very specific, such as "wet coastal needle-leafed forests". Biomes vary due to global variations in climate . Biomes are often defined by their structure: at

5002-542: Is one of the best known wetlands in Cuatrociénegas. There is a photographic exhibition hall of the flora and fauna of the reserve, viewing platforms, and self-guided trails. La Poza de la Becerra (Spanish: The Pool of the Calf) is located 16 km (9.9 mi) from the county seat, on road number 30 Cuatrociénegas-Torreon. It is part of the biosphere reserve of Cuatrociénegas. The area contains many species endemic to

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5124-531: Is primarily achieved through bacterial and fungal action. Fungal hyphae produce enzymes that can break through the tough outer structures surrounding dead plant material. They also produce enzymes that break down lignin , which allows them access to both cell contents and the nitrogen in the lignin. Fungi can transfer carbon and nitrogen through their hyphal networks and thus, unlike bacteria, are not dependent solely on locally available resources. Decomposition rates vary among ecosystems. The rate of decomposition

5246-844: Is spoken by very small communities in Angola due to Cuban influence from the Cold War and in South Sudan among South Sudanese natives that relocated to Cuba during the Sudanese wars and returned for their country's independence. Spanish is also spoken in the integral territories of Spain in Africa, namely the cities of Ceuta and Melilla and the Canary Islands , located in the Atlantic Ocean some 100 km (62 mi) off

5368-485: Is termed its resistance . The capacity of a system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Resilience thinking also includes humanity as an integral part of the biosphere where we are dependent on ecosystem services for our survival and must build and maintain their natural capacities to withstand shocks and disturbances. Time plays

5490-422: Is the "best" classification. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of the definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Different approaches to ecological classifications have been developed in terrestrial, freshwater and marine disciplines, and

5612-813: Is the de facto national language of the country, Spanish is often used in public services and notices at the federal and state levels. Spanish is also used in administration in the state of New Mexico . The language has a strong influence in major metropolitan areas such as those of Los Angeles , Miami , San Antonio , New York , San Francisco , Dallas , Tucson and Phoenix of the Arizona Sun Corridor , as well as more recently, Chicago , Las Vegas , Boston , Denver , Houston , Indianapolis , Philadelphia , Cleveland , Salt Lake City , Atlanta , Nashville , Orlando , Tampa , Raleigh and Baltimore-Washington, D.C. due to 20th- and 21st-century immigration. Although Spanish has no official recognition in

5734-498: Is the official language of 20 countries , as well as one of the six official languages of the United Nations . Spanish is the world's second-most spoken native language after Mandarin Chinese ; the world's fourth-most spoken language overall after English , Mandarin Chinese, and Hindustani ( Hindi - Urdu ); and the world's most widely spoken Romance language. The country with the largest population of native speakers

5856-578: Is usually assumed to be derived from castillo ('castle'). In the Middle Ages , the language spoken in Castile was generically referred to as Romance and later also as Lengua vulgar . Later in the period, it gained geographical specification as Romance castellano ( romanz castellano , romanz de Castiella ), lenguaje de Castiella , and ultimately simply as castellano (noun). Different etymologies have been suggested for

5978-568: The Spanish as a First Foreign Language (SAFFL) initiative in March 2005. Spanish has historically had a significant presence on the Dutch Caribbean islands of Aruba , Bonaire and Curaçao ( ABC Islands ) throughout the centuries and in present times. The majority of the populations of each island (especially Aruba) speaking Spanish at varying although often high degrees of fluency. The local language Papiamentu (Papiamento on Aruba)

6100-508: The Spanish–American War but was eventually replaced by English as the primary language of administration and education by the 1920s. Nevertheless, despite a significant decrease in influence and speakers, Spanish remained an official language of the Philippines upon independence in 1946, alongside English and Filipino , a standardized version of Tagalog . Spanish was briefly removed from official status in 1973 but reimplemented under

6222-820: The United Nations , European Union , Organization of American States , Union of South American Nations , Community of Latin American and Caribbean States , African Union , among others. In Spain and some other parts of the Spanish-speaking world, Spanish is called not only español but also castellano (Castilian), the language from the Kingdom of Castile , contrasting it with other languages spoken in Spain such as Galician , Basque , Asturian , Catalan/Valencian , Aragonese , Occitan and other minor languages. The Spanish Constitution of 1978 uses

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6344-555: The resource inputs are generally controlled by external processes like climate and parent material, the availability of these resources within the ecosystem is controlled by internal factors like decomposition, root competition or shading. Other factors like disturbance, succession or the types of species present are also internal factors. Primary production is the production of organic matter from inorganic carbon sources. This mainly occurs through photosynthesis . The energy incorporated through this process supports life on earth, while

6466-435: The "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . They also include less tangible items like tourism and recreation, and genes from wild plants and animals that can be used to improve domestic species. Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like

6588-404: The 2021–2022 school year alone. The local business process outsourcing industry has also helped boost the language's economic prospects. Today, while the actual number of proficient Spanish speakers is around 400,000, or under 0.5% of the population, a new generation of Spanish speakers in the Philippines has likewise emerged, though speaker estimates vary widely. Aside from standard Spanish,

6710-595: The 20th century, Spanish was introduced to Equatorial Guinea and the Western Sahara , and to areas of the United States that had not been part of the Spanish Empire, such as Spanish Harlem in New York City . For details on borrowed words and other external influences upon Spanish, see Influences on the Spanish language . Spanish is the primary language in 20 countries worldwide. As of 2023, it

6832-630: The CCB and surrounding valleys, the aquifer extends far beyond the CCB and includes adjacent valleys. She published her results in the Proceedings of the National Academy of Sciences in April 2006. Regarding the source of drying out, she wrote: "Similar to situations occurring with increasing frequency in various arid regions of the world, agricultural development and associated water extraction in

6954-629: The Cuatro Ciénegas Basin (CCB) in recent years, although these changes have not been well documented in the scientific literature. Some have blamed the possible drying out on changes in climate and others have ascribed it to the introduction of large scale agriculture in adjacent valleys over the past two decades. Valeria Souza , an ecology professor and researcher at the National Autonomous University of Mexico, found that based on genetic studies of microbes in

7076-654: The Equatoguinean education system and is the primary language used in government and business. Whereas it is not the mother tongue of virtually any of its speakers, the vast majority of the population is proficient in Spanish. The Instituto Cervantes estimates that 87.7% of the population is fluent in Spanish. The proportion of proficient Spanish speakers in Equatorial Guinea exceeds the proportion of proficient speakers in other West and Central African nations of their respective colonial languages. Spanish

7198-705: The Iberian Peninsula. These languages included Proto-Basque , Iberian , Lusitanian , Celtiberian and Gallaecian . The first documents to show traces of what is today regarded as the precursor of modern Spanish are from the 9th century. Throughout the Middle Ages and into the modern era , the most important influences on the Spanish lexicon came from neighboring Romance languages — Mozarabic ( Andalusi Romance ), Navarro-Aragonese , Leonese , Catalan/Valencian , Portuguese , Galician , Occitan , and later, French and Italian . Spanish also borrowed

7320-540: The Old Spanish sibilants) for details. The Gramática de la lengua castellana , written in Salamanca in 1492 by Elio Antonio de Nebrija , was the first grammar written for a modern European language. According to a popular anecdote, when Nebrija presented it to Queen Isabella I , she asked him what was the use of such a work, and he answered that language is the instrument of empire. In his introduction to

7442-589: The Royal Spanish Academy) states that, although the Royal Spanish Academy prefers to use the term español in its publications when referring to the Spanish language, both terms— español and castellano —are regarded as synonymous and equally valid. The term castellano is related to Castile ( Castilla or archaically Castiella ), the kingdom where the language was originally spoken. The name Castile , in turn,

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7564-452: The State. ... The other Spanish languages shall also be official in their respective Autonomous Communities... The Royal Spanish Academy ( Real Academia Española ), on the other hand, currently uses the term español in its publications. However, from 1713 to 1923, it called the language castellano . The Diccionario panhispánico de dudas (a language guide published by

7686-502: The U.S. population were of Hispanic or Hispanic American by origin. In turn, 41.8 million people in the United States aged five or older speak Spanish at home, or about 13% of the population. Spanish predominates in the unincorporated territory of Puerto Rico , where it is also an official language along with English. Spanish is by far the most common second language in the country, with over 50 million total speakers if non-native or second-language speakers are included. While English

7808-481: The absence of disturbance, net ecosystem production is equivalent to the net carbon accumulation in the ecosystem. Energy can also be released from an ecosystem through disturbances such as wildfire or transferred to other ecosystems (e.g., from a forest to a stream to a lake) by erosion . In aquatic systems , the proportion of plant biomass that gets consumed by herbivores is much higher than in terrestrial systems. In trophic systems, photosynthetic organisms are

7930-457: The administration of Ferdinand Marcos two months later. It remained an official language until the ratification of the present constitution in 1987, in which it was re-designated as a voluntary and optional auxiliary language. Additionally, the constitution, in its Article XIV, stipulates that the Government shall provide the people of the Philippines with a Spanish-language translation of

8052-402: The amount of energy available to the ecosystem. Parent material determines the nature of the soil in an ecosystem, and influences the supply of mineral nutrients. Topography also controls ecosystem processes by affecting things like microclimate , soil development and the movement of water through a system. For example, ecosystems can be quite different if situated in a small depression on

8174-504: The amount of light available, the amount of leaf area a plant has to capture light (shading by other plants is a major limitation of photosynthesis), the rate at which carbon dioxide can be supplied to the chloroplasts to support photosynthesis, the availability of water, and the availability of suitable temperatures for carrying out photosynthesis. Energy and carbon enter ecosystems through photosynthesis, are incorporated into living tissue, transferred to other organisms that feed on

8296-408: The carbon makes up much of the organic matter in living and dead biomass, soil carbon and fossil fuels . It also drives the carbon cycle , which influences global climate via the greenhouse effect . Through the process of photosynthesis, plants capture energy from light and use it to combine carbon dioxide and water to produce carbohydrates and oxygen . The photosynthesis carried out by all

8418-684: The classic hispanus or hispanicus took the suffix -one from Vulgar Latin , as happened with other words such as bretón (Breton) or sajón (Saxon). Like the other Romance languages , the Spanish language evolved from Vulgar Latin , which was brought to the Iberian Peninsula by the Romans during the Second Punic War , beginning in 210 BC. Several pre-Roman languages (also called Paleohispanic languages )—some distantly related to Latin as Indo-European languages , and some that are not related at all—were previously spoken in

8540-432: The combustion of fossil fuels, ammonia gas which evaporates from agricultural fields which have had fertilizers applied to them, and dust. Anthropogenic nitrogen inputs account for about 80% of all nitrogen fluxes in ecosystems. When plant tissues are shed or are eaten, the nitrogen in those tissues becomes available to animals and microbes. Microbial decomposition releases nitrogen compounds from dead organic matter in

8662-418: The concept to draw attention to the importance of transfers of materials between organisms and their environment. He later refined the term, describing it as "The whole system, ... including not only the organism-complex, but also the whole complex of physical factors forming what we call the environment". Tansley regarded ecosystems not simply as natural units, but as "mental isolates". Tansley later defined

8784-500: The consonant written b (a bilabial with plosive and fricative allophones). In modern Spanish, there is no difference between the pronunciation of orthographic b and v . Typical of Spanish (as also of neighboring Gascon extending as far north as the Gironde estuary , and found in a small area of Calabria ), attributed by some scholars to a Basque substratum was the mutation of Latin initial f into h- whenever it

8906-404: The country's constitution. In recent years changing attitudes among non-Spanish speaking Filipinos have helped spur a revival of the language, and starting in 2009 Spanish was reintroduced as part of the basic education curriculum in a number of public high schools, becoming the largest foreign language program offered by the public school system, with over 7,000 students studying the language in

9028-403: The dead organic matter would accumulate in an ecosystem, and nutrients and atmospheric carbon dioxide would be depleted. Decomposition processes can be separated into three categories— leaching , fragmentation and chemical alteration of dead material. As water moves through dead organic matter, it dissolves and carries with it the water-soluble components. These are then taken up by organisms in

9150-447: The definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Biotic factors of the ecosystem are living things; such as plants, animals, and bacteria, while abiotic are non-living components; such as water, soil and atmosphere. Plants allow energy to enter the system through photosynthesis , building up plant tissue. Animals play an important role in

9272-655: The ecosystem or to gradual disruption of biotic processes and degradation of abiotic conditions of the ecosystem. Once the original ecosystem has lost its defining features, it is considered "collapsed ". Ecosystem restoration can contribute to achieving the Sustainable Development Goals . An ecosystem (or ecological system) consists of all the organisms and the abiotic pools (or physical environment) with which they interact. The biotic and abiotic components are linked together through nutrient cycles and energy flows. "Ecosystem processes" are

9394-528: The ecosystem. Internal factors are controlled, for example, by decomposition , root competition, shading, disturbance, succession, and the types of species present. While the resource inputs are generally controlled by external processes, the availability of these resources within the ecosystem is controlled by internal factors. Therefore, internal factors not only control ecosystem processes but are also controlled by them. Ecosystems are dynamic entities—they are subject to periodic disturbances and are always in

9516-412: The eradication of invasive flora and fauna, as well as community development and water efficient agriculture combined with organic techniques. Cuatro Ciénegas has several natural tourist attractions, including white dunes and many wetlands which have unique ecosystems . The most famous natural attractions are: Poza Azul is a protected wetland, located 9 km (5.6 mi) from Cuatrociénegas seat. It

9638-421: The flow of energy through a lake was the primary driver of the ecosystem. Hutchinson's students, brothers Howard T. Odum and Eugene P. Odum , further developed a "systems approach" to the study of ecosystems. This allowed them to study the flow of energy and material through ecological systems. Ecosystems are controlled by both external and internal factors. External factors, also called state factors, control

9760-683: The following table: Spanish is marked by palatalization of the Latin double consonants ( geminates ) nn and ll (thus Latin annum > Spanish año , and Latin anellum > Spanish anillo ). The consonant written u or v in Latin and pronounced [w] in Classical Latin had probably " fortified " to a bilabial fricative /β/ in Vulgar Latin. In early Spanish (but not in Catalan or Portuguese) it merged with

9882-458: The forests of eastern North America still show legacies of cultivation which ceased in 1850 when large areas were reverted to forests. Another example is the methane production in eastern Siberian lakes that is controlled by organic matter which accumulated during the Pleistocene . Ecosystems continually exchange energy and carbon with the wider environment . Mineral nutrients, on

10004-466: The former British colony of Belize (known until 1973 as British Honduras ) where English is the sole official language, according to the 2022 census, 54% of the total population are able to speak the language. Due to its proximity to Spanish-speaking countries and small existing native Spanish speaking minority, Trinidad and Tobago has implemented Spanish language teaching into its education system. The Trinidadian and Tobagonian government launched

10126-409: The genes necessary to partially replace its membrane phospholipids with sulfolipids through horizontal gene transfer . Several environmental conservation leaders are working to protect the valley, including Pronatura Noreste . The organization owns a private reserve, called Pozas Azules , and has several ongoing projects that include the protection of native species, including stromatolites and

10248-604: The grammar, dated 18 August 1492, Nebrija wrote that "... language was always the companion of empire." From the 16th century onwards, the language was taken to the Spanish-discovered America and the Spanish East Indies via Spanish colonization of America . Miguel de Cervantes , author of Don Quixote , is such a well-known reference in the world that Spanish is often called la lengua de Cervantes ("the language of Cervantes"). In

10370-691: The landscape, versus one present on an adjacent steep hillside. Other external factors that play an important role in ecosystem functioning include time and potential biota , the organisms that are present in a region and could potentially occupy a particular site. Ecosystems in similar environments that are located in different parts of the world can end up doing things very differently simply because they have different pools of species present. The introduction of non-native species can cause substantial shifts in ecosystem function. Unlike external factors, internal factors in ecosystems not only control ecosystem processes but are also controlled by them. While

10492-556: The language happened in Toledo , a prominent city of the Kingdom of Castile , in the 13th century. Spanish colonialism in the early modern period spurred the introduction of the language to overseas locations, most notably to the Americas. As a Romance language, Spanish is a descendant of Latin. Around 75% of modern Spanish vocabulary is Latin in origin, including Latin borrowings from Ancient Greek. Alongside English and French , it

10614-549: The language's hegemony in an intensely centralising Spanish state were established from the 18th century onward. Other European territories in which it is also widely spoken include Gibraltar and Andorra . Spanish is also spoken by immigrant communities in other European countries, such as the United Kingdom , France , Italy , and Germany . Spanish is an official language of the European Union . Today,

10736-413: The largest in the country, and the third largest in the Americas . Other featured wetlands include "Nogalito" and the ecotouristic park "La Ilusión,” as well as El Entronque and El Mesquite resorts. The Sierra de la Campana , a mountain range with a huge crater called El hundido, is also a tourist attraction. Cuatro Cienégas is also an important wine-making region; the winery Bodegas Ferrino , founded by

10858-423: The living and dead plant matter, and eventually released through respiration. The carbon and energy incorporated into plant tissues (net primary production) is either consumed by animals while the plant is alive, or it remains uneaten when the plant tissue dies and becomes detritus . In terrestrial ecosystems , the vast majority of the net primary production ends up being broken down by decomposers . The remainder

10980-408: The maintenance of hydrological cycles, cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. While material from the ecosystem had traditionally been recognized as being the basis for things of economic value, ecosystem services tend to be taken for granted. The Millennium Ecosystem Assessment

11102-405: The maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. Many ecosystems become degraded through human impacts, such as soil loss , air and water pollution , habitat fragmentation , water diversion , fire suppression , and introduced species and invasive species . These threats can lead to abrupt transformation of

11224-694: The majority of the Spanish speakers live in Hispanic America . Nationally, Spanish is the official language—either de facto or de jure —of Argentina , Bolivia (co-official with 36 indigenous languages), Chile , Colombia , Costa Rica , Cuba , Dominican Republic , Ecuador , El Salvador , Guatemala , Honduras , Mexico (co-official with 63 indigenous languages), Nicaragua , Panama , Paraguay (co-official with Guaraní ), Peru (co-official with Quechua , Aymara , and "the other indigenous languages"), Puerto Rico (co-official with English), Uruguay , and Venezuela . Spanish language has

11346-433: The movement of matter and energy through the system, by feeding on plants and on one another. They also influence the quantity of plant and microbial biomass present. By breaking down dead organic matter , decomposers release carbon back to the atmosphere and facilitate nutrient cycling by converting nutrients stored in dead biomass back to a form that can be readily used by plants and microbes. Ecosystems provide

11468-734: The northwest of the African mainland. The Spanish spoken in the Canary Islands traces its origins back to the Castilian conquest in the 15th century , and, in addition to a resemblance to Western Andalusian speech patterns, it also features strong influence from the Spanish varieties spoken in the Americas, which in turn have also been influenced historically by Canarian Spanish. The Spanish spoken in North Africa by native bilingual speakers of Arabic or Berber who also speak Spanish as

11590-537: The other hand, are mostly cycled back and forth between plants, animals, microbes and the soil. Most nitrogen enters ecosystems through biological nitrogen fixation , is deposited through precipitation, dust, gases or is applied as fertilizer . Most terrestrial ecosystems are nitrogen-limited in the short term making nitrogen cycling an important control on ecosystem production. Over the long term, phosphorus availability can also be critical. Macronutrients which are required by all plants in large quantities include

11712-509: The other. Despite this, the cumulative effect of additional species in an ecosystem is not linear: additional species may enhance nitrogen retention, for example. However, beyond some level of species richness, additional species may have little additive effect unless they differ substantially from species already present. This is the case for example for exotic species . The addition (or loss) of species that are ecologically similar to those already present in an ecosystem tends to only have

11834-403: The overall structure of an ecosystem and the way things work within it, but are not themselves influenced by the ecosystem. On broad geographic scales, climate is the factor that "most strongly determines ecosystem processes and structure". Climate determines the biome in which the ecosystem is embedded. Rainfall patterns and seasonal temperatures influence photosynthesis and thereby determine

11956-405: The plants in an ecosystem is called the gross primary production (GPP). About half of the gross GPP is respired by plants in order to provide the energy that supports their growth and maintenance. The remainder, that portion of GPP that is not used up by respiration, is known as the net primary production (NPP). Total photosynthesis is limited by a range of environmental factors. These include

12078-533: The population. Many northern Moroccans have rudimentary knowledge of Spanish, with Spanish being particularly significant in areas adjacent to Ceuta and Melilla. Spanish also has a presence in the education system of the country (through either selected education centers implementing Spain's education system, primarily located in the North, or the availability of Spanish as foreign language subject in secondary education). In Western Sahara , formerly Spanish Sahara ,

12200-585: The primary nutrients (which are most limiting as they are used in largest amounts): Nitrogen, phosphorus, potassium. Secondary major nutrients (less often limiting) include: Calcium, magnesium, sulfur. Micronutrients required by all plants in small quantities include boron, chloride, copper, iron, manganese, molybdenum, zinc. Finally, there are also beneficial nutrients which may be required by certain plants or by plants under specific environmental conditions: aluminum, cobalt, iodine, nickel, selenium, silicon, sodium, vanadium. Until modern times, nitrogen fixation

12322-413: The primary producers. The organisms that consume their tissues are called primary consumers or secondary producers — herbivores . Organisms which feed on microbes ( bacteria and fungi ) are termed microbivores . Animals that feed on primary consumers— carnivores —are secondary consumers. Each of these constitutes a trophic level. The sequence of consumption—from plant to herbivore, to carnivore—forms

12444-419: The process of recovering from some past disturbance. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance, is termed its resistance . The capacity of a system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Ecosystems can be studied through

12566-544: The region have placed new pressures on the ecological integrity of the unique ecosystems of Cuatro Ciénegas." Research by hydrogeologist Brad Wolaver at the University of Texas at Austin, now at Flinders University, also found evidence that the aquifer supplying the water that emerges at the surface of the Cuatro Ciénegas Basin extends far beyond the basin and thus is potentially impacted by agricultural water extraction in adjacent valleys. Spanish language This

12688-432: The region. Poza Churince is located 18 km (11 mi) from the Cuatrociénegas seat by the federal highway 30 Cuatrociénegas-Torreon. It is an area of spring waters and varying temperatures. Las Playitas is a large extension of wetlands located close to the municipal seat. Also known in Spanish as "Las Dunas de Yeso,” white dunes cover a large area of white sand that consists of calcium sulfate . They are considered

12810-611: The soil, react with mineral soil, or are transported beyond the confines of the ecosystem (and are considered lost to it). Newly shed leaves and newly dead animals have high concentrations of water-soluble components and include sugars , amino acids and mineral nutrients. Leaching is more important in wet environments and less important in dry ones. Fragmentation processes break organic material into smaller pieces, exposing new surfaces for colonization by microbes. Freshly shed leaf litter may be inaccessible due to an outer layer of cuticle or bark , and cell contents are protected by

12932-511: The soil, where plants, fungi, and bacteria compete for it. Some soil bacteria use organic nitrogen-containing compounds as a source of carbon, and release ammonium ions into the soil. This process is known as nitrogen mineralization . Others convert ammonium to nitrite and nitrate ions, a process known as nitrification . Nitric oxide and nitrous oxide are also produced during nitrification. Under nitrogen-rich and oxygen-poor conditions, nitrates and nitrites are converted to nitrogen gas ,

13054-483: The spatial extent of ecosystems using the term " ecotope ". G. Evelyn Hutchinson , a limnologist who was a contemporary of Tansley's, combined Charles Elton 's ideas about trophic ecology with those of Russian geochemist Vladimir Vernadsky . As a result, he suggested that mineral nutrient availability in a lake limited algal production . This would, in turn, limit the abundance of animals that feed on algae. Raymond Lindeman took these ideas further to suggest that

13176-439: The successful establishment of a town by Antonio Cordero y Bustamante on 24 May 1800. The settlement's original name was Nuestra Señora de los Dolores y Cuatro Ciénegas, which was later changed to Villa Venustiano Carranza, before finally settling on its current name. The city is formally known as Cuatro Ciénegas de Carranza , in honour of its most famous son: Venustiano Carranza , President of Mexico from 1915 to 1920, who

13298-415: The term castellano to define the official language of the whole of Spain, in contrast to las demás lenguas españolas (lit. "the other Spanish languages "). Article III reads as follows: El castellano es la lengua española oficial del Estado. ... Las demás lenguas españolas serán también oficiales en las respectivas Comunidades Autónomas... Castilian is the official Spanish language of

13420-537: The term español (Spanish). According to the Royal Spanish Academy, español derives from the Occitan word espaignol and that, in turn, derives from the Vulgar Latin * hispaniolus ('of Hispania'). Hispania was the Roman name for the entire Iberian Peninsula . There are other hypotheses apart from the one suggested by the Royal Spanish Academy. Spanish philologist Ramón Menéndez Pidal suggested that

13542-444: The theories of Ramón Menéndez Pidal , local sociolects of Vulgar Latin evolved into Spanish, in the north of Iberia, in an area centered in the city of Burgos , and this dialect was later brought to the city of Toledo , where the written standard of Spanish was first developed, in the 13th century. In this formative stage, Spanish developed a strongly differing variant from its close cousin, Leonese , and, according to some authors,

13664-640: The third most used language on the Internet , after English and Chinese. Spanish is the official language of Spain . Upon the emergence of the Castilian Crown as the dominant power in the Iberian Peninsula by the end of the Middle Ages, the Romance vernacular associated with this polity became increasingly used in instances of prestige and influence, and the distinction between "Castilian" and "Spanish" started to become blurred. Hard policies imposing

13786-421: The transfers of energy and materials from one pool to another. Ecosystem processes are known to "take place at a wide range of scales". Therefore, the correct scale of study depends on the question asked. The term "ecosystem" was first used in 1935 in a publication by British ecologist Arthur Tansley . The term was coined by Arthur Roy Clapham , who came up with the word at Tansley's request. Tansley devised

13908-717: The valley and its surrounding mountains (e.g., Fouquieria shrevei ). Among the many aquatic species in the Reserve are three endemic turtles ( Coahuilan box turtle , Cuatro Ciénegas slider and Cuatro Cienegas softshell ), eight endemic fish ( Minckley's cichlid , Cuatro Cienegas shiner , Tufa darter , Bolson pupfish , Cuatro Cienegas pupfish , Cuatrociengas gambusia , Cuatrocienegas killifish and northern platyfish ), and several endemic crustaceans and gastropods , especially hydrobiid freshwater snails. Live stromatolites inhabit Cuatro Ciénegas' pools. These are colonies of certain types of cyanobacteria , extinct in most of

14030-403: The world, linked to the origin of an oxygen rich atmosphere over 3 billion years ago. A tiny copepod crustacean, Leptocaris stromatolicolus , is known only from the interstices of these stromatolites and bottom sediments in the saline pools. The pools are an oligotrophic environment with little available phosphate , leading one local bacterial species, Bacillus coahuilensis , to acquire

14152-452: Was born there in 1859. The municipality reported 12,154 inhabitants in the year 2000 census. The Cuatro Ciénegas Basin (CCB) is an official nature reserve in Mexico. The basin has inflowing rivers and streams from the near region, but it is endorheic (no natural outflow); a number of artificial channels leading water out of it have been made for irrigation and in prehistoric times it

14274-634: Was developed in the cities of Toledo , in the 13th to 16th centuries, and Madrid , from the 1570s. The development of the Spanish sound system from that of Vulgar Latin exhibits most of the changes that are typical of Western Romance languages , including lenition of intervocalic consonants (thus Latin vīta > Spanish vida ). The diphthongization of Latin stressed short e and o —which occurred in open syllables in French and Italian, but not at all in Catalan or Portuguese—is found in both open and closed syllables in Spanish, as shown in

14396-622: Was distinguished by a heavy Basque influence (see Iberian Romance languages ). This distinctive dialect spread to southern Spain with the advance of the Reconquista , and meanwhile gathered a sizable lexical influence from the Arabic of Al-Andalus , much of it indirectly, through the Romance Mozarabic dialects (some 4,000 Arabic -derived words, make up around 8% of the language today). The written standard for this new language

14518-713: Was followed by a vowel that did not diphthongize. The h- , still preserved in spelling, is now silent in most varieties of the language, although in some Andalusian and Caribbean dialects, it is still aspirated in some words. Because of borrowings from Latin and neighboring Romance languages, there are many f -/ h - doublets in modern Spanish: Fernando and Hernando (both Spanish for "Ferdinand"), ferrero and herrero (both Spanish for "smith"), fierro and hierro (both Spanish for "iron"), and fondo and hondo (both words pertaining to depth in Spanish, though fondo means "bottom", while hondo means "deep"); additionally, hacer ("to make")

14640-470: Was part of the Rio Grande (Rio Bravo) basin. The nature reserves are small ecosystems with unique fauna and flora that are highly protected by government authorities. Recently, NASA stated that the Cuatro Ciénegas Basin could have strong links to discovering life on Mars , since the adaptability of bioforms in the region was unique in the world. There are some 150 different plants and animals endemic to

14762-462: Was revoked by Michel Temer after the impeachment of Dilma Rousseff . In many border towns and villages along Paraguay and Uruguay, a mixed language known as Portuñol is spoken. Equatorial Guinea is the only Spanish-speaking country located entirely in Africa, with the language introduced during the Spanish colonial period . Enshrined in the constitution as an official language (alongside French and Portuguese), Spanish features prominently in

14884-679: Was the major source of nitrogen for ecosystems. Nitrogen-fixing bacteria either live symbiotically with plants or live freely in the soil. The energetic cost is high for plants that support nitrogen-fixing symbionts—as much as 25% of gross primary production when measured in controlled conditions. Many members of the legume plant family support nitrogen-fixing symbionts. Some cyanobacteria are also capable of nitrogen fixation. These are phototrophs , which carry out photosynthesis. Like other nitrogen-fixing bacteria, they can either be free-living or have symbiotic relationships with plants. Other sources of nitrogen include acid deposition produced through

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