156-798: Panthera leo melanochaita is a lion subspecies in Southern and East Africa . In this part of Africa , lion populations are regionally extinct in Lesotho , Djibouti and Eritrea , and are threatened by loss of habitat and prey base, killing by local people in retaliation for loss of livestock, and in several countries also by trophy hunting . Since the turn of the 21st century, lion populations in intensively managed protected areas in Botswana, Namibia, South Africa and Zimbabwe have increased, but declined in East African range countries. In 2005,
312-609: A man-eating lion shot in 1936 in eastern Transvaal weighed about 313 kg (690 lb), and was considered to have been one of the heaviest wild lions. In 1963, two lions in Tanzania weighed 320 and 360 kg (700 and 800 lb) after killing several livestock . In the 19th and 20th centuries, lion type specimens were described on the basis of their mane size and colour. Mane colour varies from sandy, tawny, isabelline, light reddish yellow to dark brown and black. Mane length varies from short to extending to knee joints and under
468-834: A Lion Conservation Strategy was developed for East and Southern Africa. Results of a phylogeographic study indicate that lion populations in southern and eastern Africa form a major clade distinct from lion populations in West Africa, Central Africa and Asia. In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group subsumed lion populations according to the major clades into two subspecies, namely P. l. leo and P. l. melanochaita . Within P. l. melanochaita three subclades are clearly distinguishable. One from northeastern Africa, another one from southwestern Africa and
624-417: A buildup of scent attracting the attention of predators, the lioness moves her cubs to a new den site several times a month, carrying them one-by-one by the nape of the neck. Usually, the mother does not integrate herself and her cubs back into the pride until the cubs are six to eight weeks old. Sometimes the introduction to pride life occurs earlier, particularly if other lionesses have given birth at about
780-549: A closer relationship with P. leo melanochaita . Lion samples from some parts of the Ethiopian Highlands cluster genetically with those from Cameroon and Chad, while lions from other areas of Ethiopia cluster with samples from East Africa. Researchers, therefore, assume Ethiopia is a contact zone between the two subspecies. Genome -wide data of a wild-born historical lion sample from Sudan showed that it clustered with P. l. leo in mtDNA-based phylogenies, but with
936-471: A constant lookout for circling vultures, which indicate the death or distress of an animal. Most carrion on which both hyenas and lions feed upon are killed by hyenas rather than lions. Carrion is thought to provide a large part of lion diet. Lions and spotted hyenas occupy a similar ecological niche and compete for prey and carrion; a review of data across several studies indicates a dietary overlap of 58.6%. Lions typically ignore hyenas unless they are on
1092-425: A favoured position in the group, either stalking prey on the "wing", then attacking, or moving a smaller distance in the centre of the group and capturing prey fleeing from other lionesses. Males attached to prides do not usually participate in group hunting. Some evidence suggests, however, that males are just as successful as females; they are typically solo hunters who ambush prey in small bushland. They may join in
1248-470: A few reported cases of old and wounded lions falling prey to wild dogs. Most lionesses reproduce by the time they are four years of age. Lions do not mate at a specific time of year and the females are polyestrous . Like those of other cats, the male lion's penis has spines that point backward. During withdrawal of the penis, the spines rake the walls of the female's vagina, which may cause ovulation . A lioness may mate with more than one male when she
1404-468: A global rise in sea level. Sea level also increases in the Indian Ocean, except in the south tropical Indian Ocean where it decreases, a pattern most likely caused by rising levels of greenhouse gases . Among the tropical oceans, the western Indian Ocean hosts one of the largest concentrations of phytoplankton blooms in summer, due to the strong monsoon winds. The monsoonal wind forcing leads to
1560-548: A heavier mane. On average, Asiatic lions have sparser manes than African lions. This feature likely evolved to signal the fitness of males to females. Males with darker manes appear to have greater reproductive success and are more likely to remain in a pride for longer. They have longer and thicker hair and higher testosterone levels, but they are also more vulnerable to heat stress. The core body temperature does apparently not increase regardless of sex, season, feeding time, length and colour of mane, but only surface temperature
1716-602: A high affinity to P. l. melanochaita . This result suggested that the taxonomic position of lions in Central Africa may require revision. Other lion subspecies or sister species to the modern lion existed in prehistoric times: The Panthera lineage is estimated to have genetically diverged from the common ancestor of the Felidae around 9.32 to 4.47 million years ago to 11.75 to 0.97 million years ago . Results of analyses differ in
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#17327760098021872-480: A key species sought for exhibition in zoological gardens across the world since the late 18th century. Cultural depictions of lions were prominent in Ancient Egypt , and depictions have occurred in virtually all ancient and medieval cultures in the lion's historic and current range. The English word lion is derived via Anglo-Norman liun from Latin leōnem (nominative: leō ), which in turn
2028-501: A kill or are being harassed, while the latter tend to visibly react to the presence of lions with or without the presence of food. In the Ngorongoro crater , lions subsist largely on kills stolen from hyenas, causing them to increase their kill rate. In Botswana's Chobe National Park, the situation is reversed as hyenas there frequently challenge lions and steal their kills, obtaining food from 63% of all lion kills. When confronted on
2184-514: A kill, hyenas may either leave or wait patiently at a distance of 30–100 m (98–328 ft) until the lions have finished. Hyenas may feed alongside lions and force them off a kill. The two species attack one another even when there is no food involved for no apparent reason. Lions can account for up to 71% of hyena deaths in Etosha National Park . Hyenas have adapted by frequently mobbing lions that enter their home ranges. When
2340-519: A kill. Crippled lions and cubs may fall victim to hyenas and leopards or be trampled by buffalo or elephants. Careless lions may be maimed when hunting prey. Nile crocodiles may also kill and eat lions, evidenced by the occasional lion claw found in crocodile stomachs. Ticks commonly infest the ears, neck and groin regions of the lions. Adult forms of several tapeworm species of the genus Taenia have been isolated from lion intestines, having been ingested as larvae in antelope meat. Lions in
2496-457: A larger extent. Hence, dispersing lions are more vulnerable to coming into conflict with humans than adult lions. In the semi-arid savanna of Zimbabwe's Hwange National Park, 19 lions were radio-collared and tracked between 2002 and 2007. Both female and male lions moved foremost within 2 km (1.2 mi) of waterholes in all seasons. Lions living near ranches in the vicinity of Tsavo East National Park consisted of three prides, two pairs and
2652-600: A larger hybridization zone to the southwestern lion clade in the Kruger National Park area (Great Limpopo Transfrontier Park). The classification of lions in Uganda is not clarified, since there were no samples included in the most comprehensive recent study. Lions, which can be grouped into the southwestern clade are found in Angola, Northern Namibia, Northern Botswana and western Zimbabwe, extending southeast into
2808-558: A lion makes when sniffing chemical signals and involves an open mouth with bared teeth, raised muzzle, wrinkled nose, closed eyes and relaxed ears. Lions also use chemical and visual marking; males spray urine and scrape plots of ground and objects within the territory. The lion's repertoire of vocalisations is large; variations in intensity and pitch appear to be central to communication. Most lion vocalisations are variations of growling , snarling , meowing and roaring. Other sounds produced include puffing, bleating and humming. Roaring
2964-473: A longer evolutionary history than genetically less diverse lion samples from Asia and West and Central Africa. A whole genome-wide sequence of lion samples showed that samples from West Africa shared alleles with samples from Southern Africa, and samples from Central Africa shared alleles with samples from Asia. This phenomenon indicates that Central Africa was a melting pot of lion populations after they had become isolated, possibly migrating through corridors in
3120-399: A loose pride while females associate with up to 12 other females, forming a stronger pride together with their cubs. Female and male lions associate only when mating. Coalitions of males hold territory for a longer time than single lions. Males in coalitions of three or four individuals exhibit a pronounced hierarchy, in which one male dominates the others and mates more frequently. The lion
3276-747: A maximum depth of 7,290 m (23,920 ft). All of the Indian Ocean is in the Eastern Hemisphere and the centre of the Eastern Hemisphere, the 90th meridian east , passes through the Ninety East Ridge . Within these waters are a number of islands. These include those controlled by surrounding countries, and independent island states and territories. Of the non-coastal islands, there are two broad clusters: one around Madagascar, and one south of India. A few other oceanic islands are scattered elsewhere. In contrast to
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#17327760098023432-433: A nomad is a "range". Males associated with a pride patrol the fringes. Both males and females defend the pride against intruders, but the male lion is better-suited for this purpose due to its stockier, more powerful build. Some individuals consistently lead the defense against intruders, while others lag behind. Lions tend to assume specific roles in the pride; slower-moving individuals may provide other valuable services to
3588-554: A nomadic phase before gaining residence in a pride. A study undertaken in the Serengeti National Park revealed that nomadic coalitions gain residency at between 3.5 and 7.3 years of age. In Kruger National Park, dispersing male lions move more than 25 km (16 mi) away from their natal pride in search of their own territory. Female lions stay closer to their natal pride. Therefore, female lions in an area are more closely related to each other than male lions in
3744-470: A period of six years, except for debris that gets indefinitely stuck in the centre of the gyre. The garbage patch in the Indian Ocean will, according to a 2012 study, decrease in size after several decades to vanish completely over centuries. Over several millennia, however, the global system of garbage patches will accumulate in the North Pacific. There are two amphidromes of opposite rotation in
3900-492: A pride of up to 30 lions killed eight African bush elephants that were between four and eleven years old. Several cases of lion attacking people have been documented: In Africa, lions are threatened by pre-emptive killing or in retaliation for preying on livestock. Prey base depletion, loss and conversion of habitat have led to a number of subpopulations becoming small and isolated. Trophy hunting has contributed to population declines in Botswana, Namibia, Zimbabwe and Zambia. It
4056-790: A process of aridification began in the southwest Indian Ocean began around 4,000 years ago. Mammalian megafauna once widespread in the MPA was driven to near extinction in the early 20th century. Some species have been successfully recovered since then — the population of white rhinoceros ( Ceratotherium simum simum ) increased from less than 20 individuals in 1895 to more than 17,000 as of 2013. Other species still depend on fenced areas and management programs, including black rhinoceros ( Diceros bicornis minor ), African wild dog ( Lycaon pictus ), cheetah ( Acynonix jubatus ), elephant ( Loxodonta africana ), and lion ( Panthera leo ). This biodiversity hotspot (and namesake ecoregion and "Endemic Bird Area")
4212-425: A reed-bed, a cave, or some other sheltered area, usually away from the pride. She will often hunt alone while the cubs are still helpless, staying relatively close to the den. Lion cubs are born blind, their eyes opening around seven days after birth. They weigh 1.2–2.1 kg (2.6–4.6 lb) at birth and are almost helpless, beginning to crawl a day or two after birth and walking around three weeks of age. To avoid
4368-565: A short, rounded head, a reduced neck, and round ears; males have broader heads. The fur varies in colour from light buff to silvery grey, yellowish red, and dark brown. The colours of the underparts are generally lighter. A new-born lion has dark spots , which fade as the cub reaches adulthood, although faint spots may still be seen on the legs and underparts. The tail of all lions ends in a dark, hairy tuft that, in some lions, conceals an approximately 5 mm (0.20 in)-long, hard "spine" or "spur" composed of dermal papillae. The functions of
4524-504: A significant portion of the world's tuna catch and is home to endangered marine species. It faces challenges like overfishing and pollution , including a significant garbage patch . Historically, the Indian Ocean has been a hub of cultural and commercial exchange since ancient times. It played a key role in early human migrations and the spread of civilizations. In modern times, it remains crucial for global trade, especially in oil and hydrocarbons. Environmental and geopolitical concerns in
4680-726: A single lion in 2002. Lions usually hunt in groups and prey foremost on ungulates such as gemsbok ( Oryx gazella ), Cape buffalo ( Syncerus caffer ), blue wildebeest ( Connochaetes taurinus ), giraffe ( Giraffa camelopardalis ), common eland ( Tragelaphus oryx ), greater kudu ( T. strepsiceros ), nyala ( T. angasii ), roan antelope ( Hippotragus equinus ), sable antelope ( H. niger ), plains zebra ( Equus quagga ), bushpig ( Potamochoerus larvatus ), common warthog ( Phacochoerus africanus ), hartebeest ( Alcephalus buselaphus ), common tsessebe ( Damaliscus lunatus ), waterbuck ( Kobus ellipsiprymnus ), kob ( K. kob ) and Thomson's gazelle ( Eudorcas thomsonii ). Their prey
4836-464: A strong coastal and open ocean upwelling , which introduces nutrients into the upper zones where sufficient light is available for photosynthesis and phytoplankton production. These phytoplankton blooms support the marine ecosystem, as the base of the marine food web, and eventually the larger fish species. The Indian Ocean accounts for the second-largest share of the most economically valuable tuna catch. Its fish are of great and growing importance to
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4992-616: A sufficient wild prey base, make lion-human coexistence sustainable and reduce factors that lead to further fragmentation of populations. Local communities in several Southern African lion range countries generate significant income through wildlife tourism , which is a strong incentive for their support of conservation measures. Establishing corridors between protected areas is important for facilitating dispersal of lions. Makgadikgadi Pans National Park and Central Kalahari Game Reserve are key dispersal areas in Southern Africa. At
5148-615: A third one from southeastern Africa. The type specimen for P. l. melanochaita was a black-maned lion from the Cape of Good Hope , known as the Cape lion . Phylogeographic analysis of lion samples from Gabon and the Republic of the Congo indicate their close genetic relation to P. l. melanochaita samples from Namibia and Botswana . It has been referred to as the Southern lion , Southern African lion , East-Southern African lion and
5304-428: A wild-born historical lion sample from Sudan clustered with P. l. leo in mtDNA-based phylogenies, but with a high affinity to P. l. melanochaita . This result indicates that the taxonomic position of lions in Central Africa may require revision. Since the beginning of the 21st century, several phylogenetic studies were conducted to aid clarifying the taxonomic status of lion samples kept in museums and collected in
5460-429: A year old and begin to hunt effectively when nearing the age of two. Single lions are capable of bringing down zebra and wildebeest, while larger prey like buffalo and giraffe are riskier. In Chobe National Park , large prides have been observed hunting African bush elephants up to around 15 years old in exceptional cases, with the victims being calves, juveniles, and even subadults. In typical hunts, each lioness has
5616-404: Is in heat . Lions of both sexes may be involved in group homosexual and courtship activities. Males will also head-rub and roll around with each other before mounting each other. Generation length of the lion is about seven years. The average gestation period is around 110 days; the female gives birth to a litter of between one and four cubs in a secluded den, which may be a thicket,
5772-905: Is a generalist hypercarnivore and is considered to be both an apex and keystone predator due to its wide prey spectrum. Its prey consists mainly of medium-sized to large ungulates , particularly blue wildebeest , plains zebra , African buffalo , gemsbok and giraffe . It also frequently takes common warthog despite it being much smaller. In India, chital and sambar deer are the most common wild prey, while livestock contributes significantly to lion kills outside protected areas. It usually avoids fully grown adult elephants , rhinoceros and hippopotamus and small prey like dik-dik , hyraxes , hares and monkeys . Unusual prey include porcupines and small reptiles. Lions kill other predators but seldom consume them. Young lions first display stalking behaviour at around three months of age, although they do not participate in hunting until they are almost
5928-532: Is a large cat of the genus Panthera , native to Africa and India . It has a muscular, broad-chested body ; a short, rounded head; round ears; and a dark, hairy tuft at the tip of its tail. It is sexually dimorphic ; adult male lions are larger than females and have a prominent mane. It is a social species , forming groups called prides. A lion's pride consists of a few adult males, related females, and cubs. Groups of female lions usually hunt together, preying mostly on medium-sized and large ungulates . The lion
6084-553: Is a mixture of dry savannah forest and very dry, deciduous scrub forest . In Africa, the range of the lion originally spanned most of the central African rainforest zone and the Sahara desert. In the 1960s, it became extinct in North Africa, except in the southern part of Sudan. During the mid- Holocene , around 8,000-6,000 years ago, the range of lions expanded into Southeastern and Eastern Europe, partially re-occupying
6240-403: Is a patchwork of small forested areas, often with a unique assemblage of species within each, located within 200 km (120 mi) from the coast and covering a total area of c. 6,200 km (2,400 sq mi). It also encompasses coastal islands, including Zanzibar and Pemba, and Mafia. This area, one of the only two hotspots that are entirely arid, includes the Ethiopian Highlands ,
6396-953: Is affected. Unlike in other felid species, female lions consistently interact with multiple males at once. Another hypothesis suggests that the mane also serves to protect the neck in fights, but this is disputed. During fights, including those involving maneless females and adolescents, the neck is not targeted as much as the face, back, and hindquarters. Injured lions also begin to lose their manes. Almost all male lions in Pendjari National Park are either maneless or have very short manes. Maneless lions have also been reported in Senegal, in Sudan 's Dinder National Park and in Tsavo East National Park , Kenya. Castrated lions often have little to no mane because
Panthera leo melanochaita - Misplaced Pages Continue
6552-470: Is an apex and keystone predator ; although some lions scavenge when opportunities occur and have been known to hunt humans , lions typically do not actively seek out and prey on humans. The lion inhabits grasslands , savannahs , and shrublands . It is usually more diurnal than other wild cats, but when persecuted, it adapts to being active at night and at twilight . During the Neolithic period,
6708-886: Is an animal symbol in shamanistic rituals of the Nuer people . In other East African cultures, it symbolizes laziness. Scars inflicted by lions are regarded as a sign of courage among the Masai people . The name ' Simba ' is a Swahili word for the lion, which also means 'aggressive', 'king' and 'strong'. Lion populations in Southern and East Africa were referred to by several regional names, including Katanga lion, Transvaal lion, Kalahari lion, Southeast African lion, and Southwest African lion, Masai lion, Serengeti lion, Tsavo lion and Uganda lion. It has also been referred to as 'Eastern-Southern African lion', 'Southern lion', and as 'southern subspecies'. Lion The lion ( Panthera leo )
6864-643: Is approximately 30° north in the Persian Gulf . The Indian Ocean covers 70,560,000 km (27,240,000 sq mi), including the Red Sea and the Persian Gulf but excluding the Southern Ocean, or 19.5% of the world's oceans; its volume is 264,000,000 km (63,000,000 cu mi) or 19.8% of the world's oceans' volume; it has an average depth of 3,741 m (12,274 ft) and
7020-825: Is artificially connected to the Mediterranean Sea without ship lock through the Suez Canal , which is accessible via the Red Sea. The Arabian Sea is connected to the Persian Gulf by the Gulf of Oman and the Strait of Hormuz . In the Persian Gulf, the Gulf of Bahrain separates Qatar from the Arabic Peninsula. Along the west coast of India, the Gulf of Kutch and Gulf of Khambat are located in Gujarat in
7176-399: Is generally mutual and the recipient appears to express pleasure. The head and neck are the most common parts of the body licked; this behaviour may have arisen out of utility because lions cannot lick these areas themselves. Lions have an array of facial expressions and body postures that serve as visual gestures. A common facial expression is the "grimace face" or flehmen response , which
7332-533: Is no evidence for gene flow between the two lineages, indicating that they did not share the same geographic area. The Eurasian and American cave lions became extinct at the end of the last glacial period without mitochondrial descendants on other continents. The modern lion was probably widely distributed in Africa during the Middle Pleistocene and started to diverge in sub-Saharan Africa during
7488-469: Is projected to push the tropical Indian Ocean into a basin-wide near-permanent heatwave state by the end of the 21st century, where marine heatwaves are projected to increase from 20 days per year (during 1970–2000) to 220–250 days per year. South of the Equator (20–5°S), the Indian Ocean is gaining heat from June to October, during the austral winter, while it is losing heat from November to March, during
7644-415: Is short and powerful; it attempts to catch prey with a fast rush and final leap, usually pulls it down by the rump, and kills with a clamping bite to the throat or muzzle . It can hold the prey's throat for up to 13 minutes, until the prey stops moving. It has a bite force from 1593.8 to 1768 Newtons at the canine tip and up 4167.6 Newtons at the carnassial notch. Lions typically consume prey at
7800-590: Is the Tsavo lion pride that always has just one adult male. Prides act as fission–fusion societies , and members will split into subgroups that keep in contact with roars . Nomadic lions range widely and move around sporadically, either in pairs or alone. Pairs are more frequent among related males. A lion may switch lifestyles; nomads can become residents and vice versa. Interactions between prides and nomads tend to be hostile, although pride females in estrus allow nomadic males to approach them. Males spend years in
7956-615: Is the most social of all wild felid species, living in groups of related individuals with their offspring. Such a group is called a " pride ". Groups of male lions are called "coalitions". Females form the stable social unit in a pride and do not tolerate outside females. The majority of females remain in their birth prides while all males and some females will disperse . The average pride consists of around 15 lions, including several adult females and up to four males and their cubs of both sexes. Large prides, consisting of up to 30 individuals, have been observed. The sole exception to this pattern
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#17327760098028112-756: Is the primary cause for a decline of lion populations in Tanzania's Selous Game Reserve and Katavi National Park . Although lions and their prey are officially protected in Tsavo National Parks, they are regularly killed by local people, with over 100 known lion killings between 2001 and 2006. Between 2008 and 2013, bones and body parts from at least 2621 individual lions were exported from South Africa to Southeast Asia , and another 3437 lion skeletons between 2014 and 2016. Lion bones are used to replace tiger bones in traditional Asian medicines . Private game ranches in South Africa also breed lions for
8268-497: Is the third-largest of the world's five oceanic divisions, covering 70,560,000 km (27,240,000 sq mi) or approximately 20% of the water area of Earth's surface . It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by the Southern Ocean , or Antarctica , depending on the definition in use. The Indian Ocean has large marginal, or regional seas, such as
8424-480: Is typically brownish and tinged with yellow, rust, and black hairs. Mutations in the genes microphthalmia-associated transcription factor and tyrosinase are possibly responsible for the colour of manes. It starts growing when lions enter adolescence, when testosterone levels increase, and reach their full size at around four years old. Cool ambient temperatures in European and North American zoos may result in
8580-426: Is unable to consume all of the kill, it rests for a few hours before continuing to eat. On hot days, the pride retreats to shade with one or two males standing guard. Lions defend their kills from scavengers such as vultures and hyenas. Lions scavenge on carrion when the opportunity arises, scavenging animals dead from natural causes such as disease or those that were killed by other predators. Scavenging lions keep
8736-579: Is used to advertise its presence. Lions most often roar at night, a sound that can be heard from a distance of 8 kilometres (5 mi). They tend to roar in a very characteristic manner starting with a few deep, long roars that subside into grunts. The lion is listed as Vulnerable on the IUCN Red List . The Indian population is listed on CITES Appendix I and the African population on CITES Appendix II . Indian Ocean The Indian Ocean
8892-911: Is usually in the range of 190–550 kg (420–1,210 pounds). In the Serengeti National Park, lions were observed to also scavenge on carrion of animals that were killed by other predators, or died from natural causes. They kept a constant lookout for circling vultures, apparently being aware that vultures indicate a dead animal. Faeces of lions collected near waterholes in Hwange National Park also contained remains of climbing mice ( Dendromus ) and common mice ( Mus ). In Botswana's Chobe National Park , lions also prey on young and subadult African bush elephants ( Loxodonta africana ). They successfully attacked 74 elephants between 1993 and 1996, of which 26 were older than nine years, and one bull over 15 years old. In October 2005,
9048-748: The Andaman Sea , the Arabian Sea , the Bay of Bengal , and the Laccadive Sea . Once called the Eastern Ocean, it is now named after India , which protrudes into it, and has been known by its current name since at least 1515. It is the only ocean named after a country. It has an average depth of 3,741 m. All of the Indian Ocean is in the Eastern Hemisphere . Unlike the Atlantic and Pacific,
9204-748: The Arabian Peninsula in the Northern Hemisphere and north of the trade winds in the Southern Hemisphere. The Indonesian Throughflow is a unique Equatorial connection to the Pacific. The climate north of the equator is affected by a monsoon climate. Strong north-east winds blow from October until April; from May until October south and west winds prevail. In the Arabian Sea, the violent Monsoon brings rain to
9360-693: The DNA sequencing analysis using 194 lion samples from 22 countries. Four of these samples clustered with samples from Central Africa, and four with samples from East Africa, indicating that the Great Rift Valley, Ethiopia was not a complete barrier to gene flow . Southeastern Ethiopia is therefore considered a genetic admixture zone between Central and East African lions. In East and Southern Africa, lion populations declined in: Contemporary lion distribution and habitat quality in East and Southern Africa
9516-737: The East African Rift constituted major barriers between the two groups. Lion samples from Gabon's Batéké Plateau National Park and Odzala-Kokoua National Park in Republic of the Congo were found to be genetically closely related to lion samples from Namibia and Botswana. A phylogenetic analysis of lion samples from Africa and Asia showed that they shared a common ancestor probably between 98,000 and 52,000 years ago. Samples from West Africa shared alleles with samples from Southern Africa, and samples from Central Africa shared alleles with samples from Asia. This indicates that Central Africa
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#17327760098029672-510: The East African Rift valley , the Socotra islands, as well as some small islands in the Red Sea and areas on the southern Arabic Peninsula. Endemic and threatened mammals include the dibatag ( Ammodorcas clarkei ) and Speke's gazelle ( Gazella spekei ); the Somali wild ass ( Equus africanus somaliensis ) and hamadryas baboon ( Papio hamadryas ). It also contains many reptiles. In Somalia,
9828-629: The Malacca , Sunda and Torres Straits . The Gulf of Carpentaria is located on the Australian north coast while the Great Australian Bight constitutes a large part of its southern coast. Several features make the Indian Ocean unique. It constitutes the core of the large-scale Tropical Warm Pool which, when interacting with the atmosphere, affects the climate both regionally and globally. Asia blocks heat export and prevents
9984-818: The Mascarene Basin the CDW becomes a deep western boundary current before it is met by a re-circulated branch of itself, the North Indian Deep Water. This mixed water partly flows north into the Somali Basin whilst most of it flows clockwise in the Mascarene Basin where an oscillating flow is produced by Rossby waves . Water circulation in the Indian Ocean is dominated by the Subtropical Anticyclonic Gyre,
10140-512: The Narmada River in central India. Lions spend much of their time resting; they are inactive for about twenty hours per day. Although lions can be active at any time, their activity generally peaks after dusk with a period of socialising, grooming, and defecating. Intermittent bursts of activity continue until dawn, when hunting most often takes place. They spend an average of two hours a day walking and fifty minutes eating. The lion
10296-484: The Nile Basin during the early Holocene . In zoos, lions have been bred with tigers to create hybrids for the curiosity of visitors or for scientific purpose. The liger is bigger than a lion and a tiger, whereas most tigons are relatively small compared to their parents because of reciprocal gene effects. The leopon is a hybrid between a lion and leopard. The lion is a muscular, broad-chested cat with
10452-695: The Sea of Zanj is located north of Madagascar. On the northern coast of the Arabian Sea , Gulf of Aden is connected to the Red Sea by the strait of Bab-el-Mandeb . In the Gulf of Aden, the Gulf of Tadjoura is located in Djibouti and the Guardafui Channel separates Socotra island from the Horn of Africa. The northern end of the Red Sea terminates in the Gulf of Aqaba and Gulf of Suez . The Indian Ocean
10608-1171: The Tuli block . a There is a larger hybridization zone to the southeastern clade around the Kruger National Park . The lion's fur varies in colour from light buff to dark brown. It has rounded ears and a black tail tuft. Average head-to-body length of male lions is 2.47–2.84 m (8 ft 1 in – 9 ft 4 in) with a weight ranging from 150–225 kg (331–496 lb) averaging 187.5 kg (413 lb) in Southern Africa and 145.4–204.7 kg (321–451 lb) averaging 174.9 kg (386 lb) in East Africa. Females average 83–165 kg (183–364 lb) in Southern Africa and 90–167.8 kg (198–370 lb) in East Africa. Males in northern Kruger National Park weighed 200.01 kg (440.9 lb) on average, whereas females weighed 143.52 kg (316.4 lb) on average, and males in southern Kruger National Park weighed 186.55 kg (411.3 lb) on average and females weighed 118.37 kg (261.0 lb), though there
10764-819: The canned hunting industry. In 2014, seven lions in Ikona Wildlife Management Area were reportedly poisoned by a herdsman for attacking his cattle . In February 2018, the carcasses of two male and four female lions were found dead in Ruaha National Park, and were suspected to have died of poisoning. In 2015 and 2017, two male lions, Cecil and his son Xanda, were killed by trophy hunters in Zimbabwe's Hwange National Park. Uncontrolled bushfires and hunting of lions and prey species in Zambia's Kafue National Park make it difficult for
10920-432: The runoff water to the Indian Ocean. Mainly in summer, this runoff flows into the Arabian Sea but also south across the Equator where it mixes with fresher seawater from the Indonesian Throughflow . This mixed freshwater joins the South Equatorial Current in the southern tropical Indian Ocean. Sea surface salinity is highest (more than 36 PSU ) in the Arabian Sea because evaporation exceeds precipitation there. In
11076-484: The " southern subspecies ". Felis (Leo) melanochaitus was the scientific name proposed by Charles Hamilton Smith in 1842 who described a lion specimen from South Africa's Cape Province. In the 19th and 20th centuries, several naturalists described zoological specimens from Southern and East Africa and proposed the following subspecies: Dispute over the validity of these purported subspecies continued among naturalists and curators of natural history museums until
11232-423: The "Hinge zone may represent the relict of continental and proto-oceanic crustal boundary formed during the rifting of India from Antarctica ." Australia, Indonesia, and India are the three countries with the longest shorelines and exclusive economic zones . The continental shelf makes up 15% of the Indian Ocean. More than two billion people live in countries bordering the Indian Ocean, compared to 1.7 billion for
11388-768: The 15th century called it the Western Oceans. In Ancient Greek geography , the Indian Ocean region known to the Greeks was called the Erythraean Sea . The borders of the Indian Ocean , as delineated by the International Hydrographic Organization in 1953 included the Southern Ocean but not the marginal seas along the northern rim but in 2002 the IHO delimited the Southern Ocean separately, which removed waters south of 60°s from
11544-516: The 1970s, the scientific name P. l. vernayi was considered synonymous with P. l. krugeri . In 1975, Vratislav Mazák hypothesized that the Cape lion evolved geographically isolated from other populations by the Great Escarpment . In the early 21st century, Mazák's hypothesis about a geographically isolated evolution of the Cape lion was challenged. Genetic exchanges between populations in
11700-727: The Arabian Peninsula) and Limpopo rivers. After the breakup of East Gondwana and the formation of the Himalayas, the Ganges-Brahmaputra rivers flow into the world's largest delta known as the Bengal delta or Sunderbans . Marginal seas , gulfs, bays and straits of the Indian Ocean include: Along the east coast of Africa, the Mozambique Channel separates Madagascar from mainland Africa, while
11856-691: The Asiatic lion population. In 2017, the Cat Classification Task Force of the Cat Specialist Group revised lion taxonomy , and recognises two subspecies based on results of several phylogeographic studies on lion evolution , namely: However, there seems to be some degree of overlap between both groups in northern Central Africa. DNA analysis from a more recent study indicates that Central African lions are derived from both northern and southern lions, as they cluster with P. leo leo in mtDNA-based phylogenies whereas their genomic DNA indicates
12012-541: The Atlantic and 2.7 billion for the Pacific (some countries border more than one ocean). The Indian Ocean drainage basin covers 21,100,000 km (8,100,000 sq mi), virtually identical to that of the Pacific Ocean and half that of the Atlantic basin, or 30% of its ocean surface (compared to 15% for the Pacific). The Indian Ocean drainage basin is divided into roughly 800 individual basins, half that of
12168-549: The Atlantic and Pacific, the Indian Ocean is enclosed by major landmasses and an archipelago on three sides and does not stretch from pole to pole, and can be likened to an embayed ocean. It is centered on the Indian Peninsula. Although this subcontinent has played a significant role in its history, the Indian Ocean has foremostly been a cosmopolitan stage, interlinking diverse regions by innovations, trade, and religion since early in human history. The active margins of
12324-707: The Cape, Kalahari and Transvaal Province regions and farther east are considered having been possible through a corridor between the Great Escarpment and the Indian Ocean . In 2005, the authors of Mammal Species of the World recognized P. l. bleyenberghi , P. l. krugeri , P. l. vernayi , P. l. massaica , P. l. hollisteri and P. l. nyanzae as valid taxa . In 2016, IUCN Red List assessors subsumed all African lion populations to P. l. leo . Two lion subspecies are now recognised: Genome -wide data of
12480-581: The Comoros. Although both species represent an order of lobe-finned fishes known from the Early Devonian (410 mya ) and though extinct 66 mya, they are morphologically distinct from their Devonian ancestors. Over millions of years, coelacanths evolved to inhabit different environments — lungs adapted for shallow, brackish waters evolved into gills adapted for deep marine waters. Of Earth's 36 biodiversity hotspots nine (or 25%) are located on
12636-483: The East India Coastal Current to the Arabian Sea from January to April. An Indian Ocean garbage patch was discovered in 2010 covering at least 5 million square kilometres (1.9 million square miles). Riding the southern Indian Ocean Gyre , this vortex of plastic garbage constantly circulates the ocean from Australia to Africa, down the Mozambique Channel , and back to Australia in
12792-532: The Indian Ocean but included the northern marginal seas. Meridionally , the Indian Ocean is delimited from the Atlantic Ocean by the 20° east meridian , running south from Cape Agulhas , South Africa, and from the Pacific Ocean by the meridian of 146°49'E, running south from South East Cape on the island of Tasmania in Australia. The northernmost extent of the Indian Ocean (including marginal seas)
12948-405: The Indian Ocean have an average width (horizontal distance from land to shelf break ) of 19 ± 0.61 km (11.81 ± 0.38 mi) with a maximum width of 175 km (109 mi). The passive margins have an average width of 47.6 ± 0.8 km (29.58 ± 0.50 mi). The average width of the slopes (horizontal distance from shelf break to foot of slope) of
13104-509: The Indian Ocean includes beaches and intertidal zones covering 3,000 km (1,200 sq mi) and 246 larger estuaries . Upwelling areas are small but important. The hypersaline salterns in India covers between 5,000–10,000 km (1,900–3,900 sq mi) and species adapted for this environment, such as Artemia salina and Dunaliella salina , are important to bird life. Coral reefs, sea grass beds, and mangrove forests are
13260-445: The Indian Ocean is bordered by landmasses and an archipelago on three sides, making it more like an embayed ocean centered on the Indian Peninsula. Its coasts and shelves differ from other oceans, with distinct features, such as a narrower continental shelf . In terms of geology, the Indian Ocean is the youngest of the major oceans, with active spreading ridges and features like seamounts and ridges formed by hotspots . The climate of
13416-503: The Indian Ocean is characterized by monsoons . It is the warmest ocean, with a significant impact on global climate due to its interaction with the atmosphere. Its waters are affected by the Indian Ocean Walker circulation , resulting in unique oceanic currents and upwelling patterns. The Indian Ocean is ecologically diverse, with important marine life and ecosystems like coral reefs, mangroves, and sea grass beds. It hosts
13572-481: The Indian Ocean region, or almost half of the world's mangrove habitat, of which 42,500 km (16,400 sq mi) is located in Indonesia, or 50% of mangroves in the Indian Ocean. Mangroves originated in the Indian Ocean region and have adapted to a wide range of its habitats but it is also where it suffers its biggest loss of habitat. In 2016, six new animal species were identified at hydrothermal vents in
13728-730: The Indian Ocean, during the past six decades. The tuna catch rates have also declined 50–90% during the past half-century, mostly due to increased industrial fisheries, with the ocean warming adding further stress to the fish species. Endangered and vulnerable marine mammals and turtles: 80% of the Indian Ocean is open ocean and includes nine large marine ecosystems : the Agulhas Current , Somali Coastal Current , Red Sea , Arabian Sea , Bay of Bengal , Gulf of Thailand , West Central Australian Shelf , Northwest Australian Shelf and Southwest Australian Shelf . Coral reefs cover c. 200,000 km (77,000 sq mi). The coasts of
13884-432: The Indian Ocean, probably caused by Rossby wave propagation. Icebergs drift as far north as 55° south latitude , similar to the Pacific but less than in the Atlantic where icebergs reach up to 45°S. The volume of iceberg loss in the Indian Ocean between 2004 and 2012 was 24 Gt . Since the 1960s, anthropogenic warming of the global ocean combined with contributions of freshwater from retreating land ice causes
14040-466: The Indian Ocean. While the Indian Ocean warmed at a rate of 1.2°C per century during 1950–2020, climate models predict accelerated warming, at a rate of 1.7 °C–3.8 °C per century during 2020–2100. Though the warming is basin-wide, maximum warming is in the northwestern Indian Ocean including the Arabian Sea, and reduced warming off the Sumatra and Java coasts in the southeast Indian Ocean. Global warming
14196-454: The Indian subcontinent. In the southern hemisphere, the winds are generally milder, but summer storms near Mauritius can be severe. When the monsoon winds change, cyclones sometimes strike the shores of the Arabian Sea and the Bay of Bengal . Some 80% of the total annual rainfall in India occurs during summer and the region is so dependent on this rainfall that many civilisations perished when
14352-578: The Late Pleistocene. Lion populations in East and Southern Africa became separated from populations in West and North Africa when the equatorial rainforest expanded 183,500 to 81,800 years ago. They shared a common ancestor probably between 98,000 and 52,000 years ago. Due to the expansion of the Sahara between 83,100 and 26,600 years ago, lion populations in West and North Africa became separated. As
14508-536: The Monsoon failed in the past. The huge variability in the Indian Summer Monsoon has also occurred pre-historically, with a strong, wet phase 33,500–32,500 BP; a weak, dry phase 26,000–23,500 BC; and a very weak phase 17,000–15,000 BP, corresponding to a series of dramatic global events: Bølling–Allerød warming , Heinrich , and Younger Dryas . The Indian Ocean is the warmest ocean in
14664-566: The Ngorongoro Crater were afflicted by an outbreak of stable fly ( Stomoxys calcitrans ) in 1962, resulting in lions becoming emaciated and covered in bloody, bare patches. Lions sought unsuccessfully to evade the biting flies by climbing trees or crawling into hyena burrows; many died or migrated and the local population dropped from 70 to 15 individuals. A more recent outbreak in 2001 killed six lions. Captive lions have been infected with canine distemper virus (CDV) since at least
14820-615: The Pacific, of which 50% are located in Asia, 30% in Africa, and 20% in Australasia. The rivers of the Indian Ocean are shorter on average (740 km (460 mi)) than those of the other major oceans. The largest rivers are ( order 5 ) the Zambezi , Ganges - Brahmaputra , Indus , Jubba , and Murray rivers and (order 4) the Shatt al-Arab , Wadi Ad Dawasir (a dried-out river system on
14976-619: The Southeast Arabian Sea salinity drops to less than 34 PSU. It is the lowest (c. 33 PSU) in the Bay of Bengal because of river runoff and precipitation. The Indonesian Throughflow and precipitation results in lower salinity (34 PSU) along the Sumatran west coast. Monsoonal variation results in eastward transportation of saltier water from the Arabian Sea to the Bay of Bengal from June to September and in westerly transport by
15132-486: The Southwest Indian Ridge, from where it continues into the Mozambique Channel and Prince Edward Fracture Zone . North of 20° south latitude the minimum surface temperature is 22 °C (72 °F), exceeding 28 °C (82 °F) to the east. Southward of 40° south latitude , temperatures drop quickly. The Bay of Bengal contributes more than half (2,950 km or 710 cu mi) of
15288-532: The Southwest Indian Ridge: a "Hoff" crab, a "giant peltospirid" snail, a whelk-like snail, a limpet, a scaleworm and a polychaete worm. The West Indian Ocean coelacanth was discovered in the Indian Ocean off South Africa in the 1930s and in the late 1990s another species, the Indonesian coelacanth , was discovered off Sulawesi Island , Indonesia. Most extant coelacanths have been found in
15444-636: The adjacent Timbavati Private Game Reserve in eastern South Africa. They were removed from the wild in the 1970s, thus decreasing the white lion gene pool . Nevertheless, 17 births have been recorded in five prides between 2007 and 2015. White lions are selected for breeding in captivity. They have reportedly been bred in camps in South Africa for use as trophies to be killed during canned hunts . African lions live in scattered populations across sub-Saharan Africa. The lion prefers grassy plains and savannahs , scrub bordering rivers, and open woodlands with bushes. It rarely enters closed forests. On Mount Elgon ,
15600-1290: The age of two years. Cubs died due to starvation in months when large prey was not available, or following take-over of the prides by new males. Male lions in coalitions are closely related. Between 1974 and 2012, 471 coalitions comprising 796 male lions entered a study area of 2,000 km (770 sq mi). Of these, 35 coalitions included male lions that were born in the area but had left and returned after about two years of absence. Nomadic coalitions became resident at between 3.5 and 7.3 years of age. The lion population of Selous Game Reserve has been surveyed since 1996. Lion prides avoided acacia woodlands and preferred habitats near water courses with short grasses, where also prey species gathered. Two or more prides shared home ranges. In Kavango–Zambezi Transfrontier Conservation Area , lions have been monitored since 1999. In 2003, 50 lions were radio-collared in Hwange National Park and tracked until 2012. Results show that adult male and female lions preferred grassland and shrubland habitat, but avoided woodlands and areas with high human density. By contrast, subadult dispersing male lions avoided grasslands and shrublands, but moved in human-dominated areas to
15756-424: The age of two. When a new male lion takes over a pride, adolescents both male and female may be evicted. Lions may live 12–17 years in the wild. Although adult lions have no natural predators, evidence suggests most die violently from attacks by humans or other lions. Lions often inflict serious injuries on members of other prides they encounter in territorial disputes or members of the home pride when fighting at
15912-544: The animal's expressive movements are highly developed. The most common peaceful, tactile gestures are head rubbing and social licking , which have been compared with the role of allogrooming among primates. Head rubbing, nuzzling the forehead, face and neck against another lion appears to be a form of greeting and is seen often after an animal has been apart from others or after a fight or confrontation. Males tend to rub other males, while cubs and females rub females. Social licking often occurs in tandem with head rubbing; it
16068-499: The attack. They take advantage of factors that reduce visibility; many kills take place near some form of cover or at night. One study in 2018 recorded a lion running at a top speed of 74.1 km/h (46.0 mph). The lion accelerates at the start of the chase by 9.5 m/s² , whereas zebras, wildebeest and Thomson's gazelle accelerate by 5 m/s², 5.6 m/s² and 4.5 m/s², respectively; acceleration appears to be more important than steady displacement speed in lion hunts. The lion's attack
16224-756: The austral summer. In 1999, the Indian Ocean Experiment showed that fossil fuel and biomass burning in South and Southeast Asia caused air pollution (also known as the Asian brown cloud ) that reach as far as the Intertropical Convergence Zone . This pollution has implications on both a local and global scale. Forty percent of the sediment of the Indian Ocean is found in the Indus and Ganges fans. The oceanic basins adjacent to
16380-886: The beginning of the 21st century, the Addis Ababa Zoo kept 16 adult lions. It is assumed that their ancestors, five males and two females, were caught in southwestern Ethiopia as part of a zoological collection for Emperor Haile Selassie I . In 2006, eight captive lions were registered under the name P. l. massaicus , and 23 as P. l. nubicus from Tanzania by the International Species Information System ; about 100 captive lions were registered as P. l. krugeri , which derived from lions captured in South Africa. In 2012, samples of lions kept at Sana'a Zoo in Yemen were found to cluster with those of lions from East and Southern Africa. The lion
16536-623: The belly. Lions without a mane were observed in the Tsavo area. Mane development is related to age: older males have more extensive manes than younger ones; manes continue to grow up to the age of four to five years, long after lions become sexually mature . Males living in the Kenyan highlands above elevations of 800 m (2,600 ft) develop heavier manes than lions in the more humid and warmer lowlands of eastern and northern Kenya. Average ambient temperature , nutrition and testosterone influence
16692-400: The bordering countries for domestic consumption and export. Fishing fleets from Russia, Japan, South Korea , and Taiwan also exploit the Indian Ocean, mainly for shrimp and tuna. Research indicates that increasing ocean temperatures are taking a toll on the marine ecosystem. A study on the phytoplankton changes in the Indian Ocean indicates a decline of up to 20% in the marine plankton in
16848-460: The cause of the decline is not fully understood, habitat loss and conflicts with humans are the greatest causes for concern. One of the most widely recognised animal symbols in human culture, the lion has been extensively depicted in sculptures and paintings, on national flags, and in literature and films. Lions have been kept in menageries since the time of the Roman Empire and have been
17004-701: The centre of the 1,500,000 km (580,000 sq mi) hotspot, the landscape is dominated by Acacia - Commiphora deciduous bushland, but also includes the Yeheb nut ( Cordeauxia edulus ) and species discovered more recently such as the Somali cyclamen ( Cyclamen somalense ), the only cyclamen outside the Mediterranean. Warsangli linnet ( Carduelis johannis ) is an endemic bird found only in northern Somalia. An unstable political situation and mismanagement has resulted in overgrazing which has produced one of
17160-488: The colour and size of the mane. Its length is an indicator for age and fighting ability of the lion. In Serengeti National Park, female lions favour males with dense and dark manes as mates. White lions have occasionally been encountered in and around South Africa's Kruger National Park and the adjacent Timbavati Private Game Reserve . Their whitish fur is a rare morph caused by a double recessive allele . It has normal pigmentation in eyes and skin. They were removed from
17316-505: The continental shelves are 50.4–52.4 km (31.3–32.6 mi) for active and passive margins respectively, with a maximum width of 205.3–255.2 km (127.6–158.6 mi). In correspondence of the Shelf break , also known as Hinge zone, the Bouguer gravity ranges from 0 to 30 mGals that is unusual for a continental region of around 16 km thick sediments. It has been hypothesized that
17472-551: The continental slopes mostly contain terrigenous sediments. The ocean south of the polar front (roughly 50° south latitude ) is high in biologic productivity and dominated by non-stratified sediment composed mostly of siliceous oozes . Near the three major mid-ocean ridges the ocean floor is relatively young and therefore bare of sediment, except for the Southwest Indian Ridge due to its ultra-slow spreading rate. The ocean's currents are mainly controlled by
17628-411: The cubs varies—one male could patiently let the cubs play with his tail or his mane, while another may snarl and bat the cubs away. Pride lionesses often synchronise their reproductive cycles and communal rearing and suckling of the young, which suckle indiscriminately from any or all of the nursing females in the pride. The synchronisation of births is advantageous because the cubs grow to being roughly
17784-455: The early 21st century. In the 20th century, some authors supported the view of the Cape lion being a distinct subspecies. In 1939, the American zoologist Allen also recognized F. l. bleyenberghi , F. l. krugeri and F. l. vernayi as valid subspecies in Southern Africa, and F. l. hollisteri , F. l. nyanzae and F. l. massaica as valid subspecies in East Africa. Pocock subordinated
17940-612: The eastern extension of which is blocked by the Southeast Indian Ridge and the 90°E Ridge. Madagascar and the Southwest Indian Ridge separate three cells south of Madagascar and off South Africa. North Atlantic Deep Water reaches into the Indian Ocean south of Africa at a depth of 2,000–3,000 m (6,600–9,800 ft) and flows north along the eastern continental slope of Africa. Deeper than NADW, Antarctic Bottom Water flows from Enderby Basin to Agulhas Basin across deep channels (<4,000 m (13,000 ft)) in
18096-534: The group. Alternatively, there may be rewards associated with being a leader that fends off intruders; the rank of lionesses in the pride is reflected in these responses. The male or males associated with the pride must defend their relationship with the pride from outside males who may attempt to usurp them. Dominance hierarchies do not appear to exist among individuals of either sex in a pride. Asiatic lion prides differ in group composition. Male Asiatic lions are solitary or associate with up to three males, forming
18252-570: The hunting of large, slower-moving prey like buffalo; and even hunt them on their own. Moderately-sized hunting groups generally have higher success rates than lone females and larger groups. Lions are not particularly known for their stamina. For instance, a lioness's heart comprises only 0.57% of her body weight and a male's is about 0.45% of his body weight, whereas a hyena's heart comprises almost 1% of its body weight. Thus, lions run quickly only in short bursts at about 48–59 km/h (30–37 mph) and need to be close to their prey before starting
18408-593: The islands of the Indian Ocean are textbook cases of evolutionary processes; the dung beetles , day geckos , and lemurs are all examples of adaptive radiation . Many bones (250 bones per square metre) of recently extinct vertebrates have been found in the Mare aux Songes swamp in Mauritius, including bones of the Dodo bird ( Raphus cucullatus ) and Cylindraspis giant tortoise. An analysis of these remains suggests
18564-547: The lion has been recorded up to an elevation of 3,600 m (11,800 ft) and close to the snow line on Mount Kenya . Savannahs with an annual rainfall of 300 to 1,500 mm (12 to 59 in) make up the majority of lion habitat in Africa, estimated at 3,390,821 km (1,309,203 sq mi) at most, but remnant populations are also present in tropical moist forests in West Africa and montane forests in East Africa. The Asiatic lion now survives only in and around Gir National Park in Gujarat, western India. Its habitat
18720-810: The lion population in Kenya's Masai Mara National Reserve declined, the spotted hyena population increased rapidly. Lions tend to dominate cheetahs and leopards, steal their kills and kill their cubs and even adults when given the chance. Cheetahs often lose their kills to lions or other predators. A study in the Serengeti ecosystem revealed that lions killed at least 17 of 125 cheetah cubs born between 1987 and 1990. Cheetahs avoid their competitors by hunting at different times and habitats. Leopards, by contrast, do not appear to be motivated by an avoidance of lions, as they use heavy vegetation regardless of whether lions are present in an area and both cats are active around
18876-486: The lion population to recover. Cub mortality in particular is high. African lions are included in CITES Appendix II . Today, lion populations are stable only in large protected area complexes. IUCN regional offices and many wildlife conservation organisations cooperated to develop a Lion Conservation Strategy for Eastern and Southern Africa in 2006. The strategy envisages to maintain sufficient habitat, ensure
19032-427: The lion ranged throughout Africa and Eurasia, from Southeast Europe to India, but it has been reduced to fragmented populations in sub-Saharan Africa and one population in western India . It has been listed as Vulnerable on the IUCN Red List since 1996 because populations in African countries have declined by about 43% since the early 1990s. Lion populations are untenable outside designated protected areas. Although
19188-507: The lion to the genus Panthera in 1930, when he wrote about Asiatic lions . Ellerman and Morrison-Scott recognized only two lion subspecies in the Palearctic realm , namely the African P. l. leo and the Asiatic P. l. persica . Various authors recognized between seven and 10 African lion subspecies. Others followed the classification proposed by Ellerman and Morrison-Scott, recognizing two subspecies including one in Africa. In
19344-551: The location of the hunt but sometimes drag large prey into cover. They tend to squabble over kills, particularly the males. Cubs suffer most when food is scarce but otherwise all pride members eat their fill, including old and crippled lions, which can live on leftovers. Large kills are shared more widely among pride members. An adult lioness requires an average of about 5 kg (11 lb) of meat per day while males require about 7 kg (15 lb). Lions gorge themselves and eat up to 30 kg (66 lb) in one session. If it
19500-481: The margins of the Indian Ocean. The origin of this diversity is debated; the break-up of Gondwana can explain vicariance older than 100 mya, but the diversity on the younger, smaller islands must have required a Cenozoic dispersal from the rims of the Indian Ocean to the islands. A "reverse colonisation", from islands to continents, apparently occurred more recently; the chameleons , for example, first diversified on Madagascar and then colonised Africa. Several species on
19656-634: The mid-18th century, as opposed to the Western Ocean ( Atlantic ) before the Pacific was surmised. In modern times, the name Afro-Asian Ocean has occasionally been used. The Hindi name for the Ocean is हिंद महासागर ( Hind Mahāsāgar ; lit. transl. Ocean of India ). Conversely, Chinese explorers (e.g., Zheng He during the Ming dynasty ) who traveled to the Indian Ocean during
19812-415: The mid-1970s. CDV is spread by domestic dogs and other carnivores; a 1994 outbreak in Serengeti National Park resulted in many lions developing neurological symptoms such as seizures. During the outbreak, several lions died from pneumonia and encephalitis . Feline immunodeficiency virus and lentivirus also affect captive lions. When resting, lion socialisation occurs through a number of behaviours;
19968-400: The monsoon. Two large gyres , one in the northern hemisphere flowing clockwise and one south of the equator moving anticlockwise (including the Agulhas Current and Agulhas Return Current ), constitute the dominant flow pattern. During the winter monsoon (November–February), however, circulation is reversed north of 30°S and winds are weakened during winter and the transitional periods between
20124-622: The monsoons. The Indian Ocean contains the largest submarine fans of the world, the Bengal Fan and Indus Fan , and the largest areas of slope terraces and rift valleys . The inflow of deep water into the Indian Ocean is 11 Sv , most of which comes from the Circumpolar Deep Water (CDW). The CDW enters the Indian Ocean through the Crozet and Madagascar basins and crosses the Southwest Indian Ridge at 30°S. In
20280-543: The most productive ecosystems of the Indian Ocean ;— coastal areas produce 20 tones of fish per square kilometre. These areas, however, are also being urbanised with populations often exceeding several thousand people per square kilometre and fishing techniques become more effective and often destructive beyond sustainable levels while the increase in sea surface temperature spreads coral bleaching. Mangroves covers 80,984 km (31,268 sq mi) in
20436-589: The northern end while the Laccadive Sea separates the Maldives from the southern tip of India. The Bay of Bengal is off the east coast of India. The Gulf of Mannar and the Palk Strait separate Sri Lanka from India, while Adam's Bridge separates the two. The Andaman Sea is located between the Bay of Bengal and the Andaman Islands. In Indonesia, the so-called Indonesian Seaway is composed of
20592-434: The phylogenetic relationship of the lion; it was thought to form a sister group with the jaguar that diverged 3.46 to 1.22 million years ago , but also with the leopard that diverged 3.1 to 1.95 million years ago to 4.32 to 0.02 million years ago . Hybridisation between lion and snow leopard ancestors possibly continued until about 2.1 million years ago. The lion-leopard clade
20748-660: The population trend is stable there as of 2012. Lions, which can be grouped into the northeastern clade are found in Somalia, Northern Kenya and Ethiopia, with a larger hybridization zone to the Northern lion subspecies ( P. l. leo ) in Ethiopia. Lions, which can be grouped into the southeastern clade are found in Southern Kenya, Western DRC , Tanzania, Mozambique, Malawi, Zambia, southern Namibia and South Africa, with
20904-612: The previous males associated with a pride, the victors often kill any existing young cubs , perhaps because females do not become fertile and receptive until their cubs mature or die. Females often fiercely defend their cubs from a usurping male but are rarely successful unless a group of three or four mothers within a pride join forces against the male. Cubs also die from starvation and abandonment, and predation by leopards, hyenas and wild dogs. Male cubs are excluded from their maternal pride when they reach maturity at around two or three years of age, while some females may leave when they reach
21060-538: The rainforest decreased and thus gave rise to more open habitats, lions moved from West to Central Africa. Lions from North Africa dispersed to southern Europe and Asia between 38,800 and 8,300 years ago. Extinction of lions in southern Europe, North Africa and the Middle East interrupted gene flow between lion populations in Asia and Africa. Genetic evidence revealed numerous mutations in lion samples from East and Southern Africa, which indicates that this group has
21216-457: The range of the now extinct cave lion . In Hungary , the modern lion was present from about 4,500 to 3,200 years Before Present . In Ukraine , the modern lion was present from about 6,400 to 2,000 years Before Present. In Greece, it was common, as reported by Herodotus in 480 BC; it was considered rare by 300 BC and extirpated by AD 100. In Asia the lion once ranged in regions where climatic conditions supported an abundance of prey. It
21372-530: The region include the effects of climate change , piracy, and strategic disputes over island territories. The Indian Ocean has been known by its present name since at least 1515, when the Latin form Oceanus Orientalis Indicus ( lit. ' Indian Eastern Ocean ' ) is attested, named after India, which projects into it. It was earlier known as the Eastern Ocean , a term that was still in use during
21528-488: The removal of the gonads inhibits testosterone production. Rarely, both wild and captive lionesses have manes. Increased testosterone may be the cause of maned lionesses reported in northern Botswana. The white lion is a rare morph with a genetic condition called leucism , which is caused by a double recessive allele . It is not albino; it has normal pigmentation in the eyes and skin. White lions have occasionally been encountered in and around Kruger National Park and
21684-429: The same area. The evolution of sociability in lions was likely driven both by high population density and the clumped resources of savannah habitats. The larger the pride, the more high-quality territory they can defend; "hotspots" being near river confluences , where the cats have better access to water, prey and shelter (via vegetation). The area occupied by a pride is called a "pride area" whereas that occupied by
21840-420: The same size and have an equal chance of survival, and sucklings are not dominated by older cubs. Weaning occurs after six or seven months. Male lions reach maturity at about three years of age and at four to five years are capable of challenging and displacing adult males associated with another pride. They begin to age and weaken at between 10 and 15 years of age at the latest. When one or more new males oust
21996-406: The same time of day. In addition, there is no evidence that lions affect leopard abundance. Leopards take refuge in trees, though lionesses occasionally attempt to climb up and retrieve their kills. Lions similarly dominate African wild dogs , taking their kills and dispatching pups or adult dogs. Population densities of wild dogs are low in areas where lions are more abundant. However, there are
22152-420: The same time. When first introduced to the rest of the pride, lion cubs lack confidence when confronted with adults other than their mother. They soon begin to immerse themselves in the pride life, however, playing among themselves or attempting to initiate play with the adults. Lionesses with cubs of their own are more likely to be tolerant of another lioness's cubs than lionesses without cubs. Male tolerance of
22308-422: The spur are unknown. The tuft is absent at birth and develops at around 5 + 1 ⁄ 2 months of age. It is readily identifiable at the age of seven months. Its skull is very similar to that of the tiger, although the frontal region is usually more depressed and flattened and has a slightly shorter postorbital region and broader nasal openings than those of the tiger. Due to the amount of skull variation in
22464-413: The tiger in size. The size and weight of adult lions vary across its range and habitats. Accounts of a few individuals that were larger than average exist from Africa and India. The male lion's mane is the most recognisable feature of the species. It may have evolved around 320,000–190,000 years ago. It grows downwards and backwards, covering most of the head, neck, shoulders, and chest. The mane
22620-407: The two species, usually only the structure of the lower jaw can be used as a reliable indicator of species. The skeletal muscles of the lion make up 58.8% of its body weight and represent the highest percentage of muscles among mammals. The lion has a high concentration of fast twitch muscle fibers, giving them quick bursts of speed but less stamina. Among felids, the lion is second only to
22776-528: The ventilation of the Indian Ocean thermocline . That continent also drives the Indian Ocean monsoon , the strongest on Earth, which causes large-scale seasonal variations in ocean currents, including the reversal of the Somali Current and Indian Monsoon Current . Because of the Indian Ocean Walker circulation there are no continuous equatorial easterlies. Upwelling occurs near the Horn of Africa and
22932-673: The wild in the 1970s, thus decreasing the white lion gene pool . Nevertheless, 17 births have been recorded in five different prides between 2007 and 2015. White lions were selected for breeding in captivity . They have been bred in South African camps for use as trophies to be killed during canned hunts . In Serengeti National Park , monitoring of lion prides started in 1966. Between 1966 and 1972, two observed lion prides comprised between seven and 10 females each. Females had litters once in 23 months on average. Litters contained two to three cubs. Of 87 cubs born until 1970, only 12 reached
23088-412: The wild. Scientists analysed between 32 and 480 lion samples from up to 22 countries. Results of genetic analyses indicate that the species comprises two main evolutionary groups, one in Southern and East Africa, and the other in the northern and eastern parts of its historical range. These groups genetically diverged between 50,000 and 200,000 years ago. It was assumed that tropical rainforest and
23244-487: The world. Long-term ocean temperature records show a rapid, continuous warming in the Indian Ocean, at about 1.2 °C (34.2 °F) (compared to 0.7 °C (33.3 °F) for the warm pool region) during 1901–2012. Research indicates that human induced greenhouse warming , and changes in the frequency and magnitude of El Niño (or the Indian Ocean Dipole ), events are a trigger to this strong warming in
23400-400: Was a borrowing from Ancient Greek λέων léōn . The Hebrew word לָבִיא lavi may also be related. The generic name Panthera is traceable to the classical Latin word 'panthēra' and the ancient Greek word πάνθηρ 'panther'. Felis leo was the scientific name used by Carl Linnaeus in 1758, who described the lion in his work Systema Naturae . The genus name Panthera
23556-461: Was a melting pot of lion populations after they had become isolated. They possibly migrated through corridors in the Nile Basin during the early Holocene . Among six samples of captive lions from Ethiopia, five clustered with samples from East Africa, but one with samples from the Sahel . For a subsequent phylogeographic study, eight wild lion samples from the Ethiopian Highlands were included in
23712-423: Was an outbreak of tuberculosis in southern park at the time. Skeletal muscles make up 58.8% of the lion's body weight. The largest known lion measured 3.35 m (11.0 ft) in length and weighed 375 kg (825 lb). An exceptionally heavy male lion near Mount Kenya weighed 272 kg (600 lb). The longest wild lion reportedly was a male shot near Mucusso National Park in southern Angola in 1973. In 1936,
23868-465: Was assessed in 2005, and Lion Conservation Units (LCU) mapped. Between 2002 and 2012, educated guesses for size of populations in these LCUs ranged from 33,967 to 32,000 individuals. The LCUs Ruaha−Rungwa, Serengeti−Mara, Tsavo−Mkomazi and Selous in East Africa, as well as Luangwa, Kgalagadi, Okavango−Hwange, Mid−Zambezi, Niassa and Greater Limpopo in Southern Africa are currently considered lion strongholds. These LCUs host more than 500 individuals each, and
24024-593: Was coined by Lorenz Oken in 1816. Between the mid-18th and mid-20th centuries, 26 lion specimens were described and proposed as subspecies, of which 11 were recognised as valid in 2005. They were distinguished mostly by the size and colour of their manes and skins. In the 19th and 20th centuries, several lion type specimens were described and proposed as subspecies , with about a dozen recognised as valid taxa until 2017. Between 2008 and 2016, IUCN Red List assessors used only two subspecific names: P. l. leo for African lion populations, and P. l. persica for
24180-464: Was distributed in the Asian and African Palearctic since at least the early Pliocene . The earliest fossils recognisable as lions were found at Olduvai Gorge in Tanzania and are estimated to be up to 2 million years old. Estimates for the divergence time of the modern and cave lion lineages range from 529,000 to 392,000 years ago based on mutation rate per generation time of the modern lion. There
24336-951: Was present in the Caucasus until the 10th century. It lived in Palestine until the Middle Ages and in Southwest Asia until the late 19th century. By the late 19th century, it had been extirpated in most of Turkey. The last live lion in Iran was sighted in 1942, about 65 km (40 mi) northwest of Dezful , although the corpse of a lioness was found on the banks of the Karun river in Khuzestan province in 1944. It once ranged from Sind and Punjab in Pakistan to Bengal and
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